Omophron (Phrator) vittulatum Fairmaire, 1894: 651

Anichtchenko, Alexander & Valainis, Uldis, 2024, To the knowledge of the subgenus Phrator Semenov, 1922 of the genus Omophron Latreille, 1802 (Coleoptera: Carabidae), Zootaxa 5447 (1), pp. 77-90 : 88-89

Omophron (Phrator) vittulatum Fairmaire, 1894: 651

Loc. Typ.: “Haut-Soudan”

Omophron (Phrator) alluaudi Dupuis, 1913: 213 syn. n.

Loc. Typ.: “Kondué” [ DR Congo].

Type material examined. Holotype, male: “H. Sóudan / Aubert [handwritten]”, “vittulatum / mih [handwritten]”, “ Ex. Musaeo / L. Fairmaire 1896 [printed]”, “Holo” “Type [red, printed]”, “ Holotype / Omophron / vittulatum / Fairmaire 1894 ” [red, handwritten] ( MNHN). Holotype, female: “ Omophron / alluaudi Dup. / det. P. Dupuis 1913 ”, “Type” [handwritten], “ Omophron alluaudi / Dup. / P. Dupuis det. / 1913”, “ Syntype ” [printed], “ Coll. R.I.Sc.N.B. / Congo belge / Ed. Luja / Kondué / Congo-Belge ” [printed], “ Collection P. Dupuis ”, “ Omophron Alluaudi Dupuis / P. Basilewsky vid. 1960 / (= vittulatus Frm.)” ( ISNB) .

Other material. ANGOLA: 1m: “ Lunda Norte Province, Dundo , 08. - 09.1948, de Baross Machado leg.” ( MRAC) ; 1m, 1f: “ Lunda Norte Province, Riv. Tshlkapa , 50 km S. O. de Dundo, 07.1948, de Baross Machado leg.” ( MRAC). CAMEROON: 1m, 1f: “Edea, Bsse-Sanaga, Vadon” ( MNHN) ; 1 ex: “ Edea , Le Moult leg.” ( ETHZ) ; 1f: “ Edea, Base Senegal , Vadon leg.” ( MNHN) ; 1m: “ Younde ” ( MRAC). CENTRAL AFRICAN REPUBLIC: 1ex: “ Kemo perfecture, Sibut , LeMoult leg.” ( ETHZ). DEMOCRATIC REPUBLIC OF THE CONGO: 18m, 21f: “ Kasai, Djokopunda , 10.08.1921, Schouteden leg.” ( MRAC, ETHZ) ; 1m: “ Kasai, Ilebo , 14- 15.07.1925, Prince Leopold leg.” ( MRAC) ; 3m, 3f: “ Kasai-Occidental Province, BenaMakima , 08.08.1921, Ghesquiere leg.” ( MRAC) ; 30m, 26f: “ Sud-Ubangi Province, Libenge , 04.01.1948, Cremer & Neuman leg.” ( ISNB) ; 1m: “ Sud-Ubangi Province, Libenge, Mawuya , 06.01.1948, Cremer & Neuman leg.” ( ISNB) ; 29m, 33f: “ Sud-Ubangi Province, Libenge, Ubangi riv., 04.01.1948, Crèmer & Neuman leg.” ( ISNB) ; 9m, 32f: “ Sud-Ubangi Province, Libenge, Zambi , 06.01.1948, Cremer & Neuman leg.” ( ISNB, MRAC) ; 1m: “ Mai – Ndombe Province, Inongo, Lake Mai-Ndombe , 11.- 24.06.1925 Prince Leopold leg.” ( ISNB) ; 19m, 23f: “ Sankuru Province, Sankuru river , 09.1924, Ghesquiere leg.” ( MRAC) ; 1m, 2f: “ Orientale Province, Kungulu ; 06.1923, Ghesquiere leg.” ( MRAC) ; 1f: “ Lomami river , 08.1924, Ghesquiere leg.” ( MRAC) ; 1m, 1f: “De Kwamoutha, Port Irangui , 05.1930, Vanderyst leg.” ( MRAC) ; 1m: “ Basongo , 10.09.1930, De Vitte leg.” ( MRAC) ; 1m, 1f: “ Basongo, River Kasai, 06.1929, Overlaet leg.” ( MRAC) ; 1f: “ Basongo , 10.09.1930, Witte leg.” ( MNHN) ; 1f: “ Env. Lac Leipold II, 11- 24.06.1925, Prince Leopold leg.” ( MRAC) ; 9m, 4f: “ Haut-Zaire, Bambesa, Alipago , 02.1935, Leroy leg.” ( MRAC) ; 1m: “ Haut–Zaire, Equater. Bokuma , 1938, Hulstaert leg.” ( MRAC) ; 1m: “ Equater. Bokuma , 07.1952, Lootens leg.” ( MRAC) ; 6m, 4f: “ Tshuapa, Bokuma , 1953, Lootens leg.” ( MRAC) ; 1f: “ Congo Belge, Rives du Congo, Mission du Bourg de Bozas , 1903” ( MNHN) ; 6m, 3f: “ Kimpangu Riv., Fulezi , 850m, 06.07.1949, Leleup leg.” ( MRAC) ; 1f: “ Lomami , 08.1924, Ghesquiere leg.” ( MNHN) ; 1m: “Doutch Zambezi” ( MNHN). CHAD: 1f: “ Fort Archambault, Mission Chari-Tchad , 1904, Decorse leg.” ( MNHN) ; 2m: “ Fort Archambault, Moyen Chari, Bongoul , Mission Chari-Tchad ( Ba Kare ), 04.1904, Decorse leg.” ( MNHN) ; 1m: “ Fort Archambault, Baha Sara, Mission Chari-Tchad , 04.1904, Decorse leg.” ( MNHN). GABON: 1m: “Ogowe” ( MNHN) ; 9m, 5f: “ Ogooue, Lambarene , 1898, Haug leg.” ( MNHN) ; 1m: “ Bas. Ogooue, Lambarene , 1901, Haug leg.” ( MNHN) ; 1m, 1f: “ Ogooue, Lambarene , 1901, Haug leg.” ( MNHN) ; 3m, 1f: “ Congo, Libreville , 1899, Boucher leg.” ( MNHN) ; 2m: “ Lambarene , 1924, Haug” ( ETHZ) ; 1m, 2f: “ Lambarene , 1896, Haug leg.” ( MNHN). NIGERIA: 1m: “ Adamawa State, Yola , Benue riv., 12.-20.04.03, Schultze leg.” ( ETHZ) ; 1m: “ Benue, 16.05.1903 ” ( ZMHB) ; 1m: “ Tarbata State, Sardauna , Sunkuru, 09.1924, Ghesquiere” ( ETHZ) ; 1f: “ Benue b. Lau , 17.04.1905, Glauning leg.” ( ZMHB) ; 1m, 2f: “ Benue, Glauning leg.” ( ZMHB). MALI: 1f: “ Macina, Dioura, OICMA , 20.09.1969, Farrow leg.” ( BMNH) ; 2m, 1f: “ Macina, Tilembeya , Niger Bank , OICMA, 05.06.1969, Farrow leg.” ( BMNH) ; 5m, 6f: “ Kara, Flood plain, OICMA, 29.06.1969, Farrow leg.” ( BMNH) ; 1f: “ Haut Senegal ”, “Comp. Type Basilewsky” ( MRAC) ; 1f: “Haut Senegal” ( MRAC).

Description. Holotype body length 7.54 mm; width 4.40 mm. Habitus ( Figs 23–25 ). Last segment of maxillar palpi elongate, 1.4 times as long as penultimate ( Figs 28–29 ). Sclerite of bursa copulatrix as in fig. 42.

Comments. For a long time this species has remained mysterious. Partly due to the fact that it was described from a single specimen with an unclear label ( Fig. 55 ), and the habitat was considered to be Sudan ( Valainis, 2016). In fact, at the end of the 19th century “Haut-Soudan” was the name of the territory of the French colony, which now for the most part corresponds to the nations of Mali, Burkina Faso and Niger. This place falls within the species’ range of O. alluaudi syn. n. ( Fig. 57 ). In its description Dupuis (1913: 216) indicated that the elytra have 14 striae, but the studied type specimen has 15 striae. The possible synonymy of O. (P.) alluaudi Dupuis, 1913 with O. (P.) vittulatum Fairmaire, 1894 had already been noticed by Basilevsky in 1960, who mentioned it on the label under the holotype of O. alluaudi ( Fig. 56 ), but these data have never been published. We confirm and establish the new synonymy here.

Distribution. Angola, Cameroon, Central African Republic, Democratic Republic of the Congo, Chad, Gabon, Mali, Nigeria ( Fig. 57 ). Previous mentions for Senegal ( Valainis, 2016) belong to “Haut Senegal ” the name of the territory of the French colony which now for the most part corresponds to the nations of Mali, Burkina Faso and Niger.

Figures

FIGURE 57. Distribution of species of the subgenus Phrator.

FIGURES 23–26. Habitus, dorsal view: (23)—O. (P.) alluaudi, holotype, syn. n. = O. (P.) vittulatum; (24)—O. (P.) vittulatum, holotype; (25)—O. (P.) vittulatum, Libenge, DRC; (26)—O. (P.) schoutedeni, Paratype. Scale bar = 5 mm.

FIGURES 27–35. Maxillar palpi and its variability. (27)—O. (P.) barsevskisi; (28)—O. (P.) alluaudi syn. n. = O. (P.) vittulatum; (29)—O. (P.) vittulatum, holotype; (30)—O. (P.) variegatum, Sardinia; (31)—O. (P.) variegatum seurati syn. n. = O. (P.) variegatum; (32)—O. (P.) multiguttatum depressum, Namibia; (33)—O. (P.) multiguttatum s.str., Kenya; (34–35)—holotype and paratype of O. (P.) ethiopiensis syn. n. = O. (P.) multiguttatum s.str. Scale bar = 1 mm.

FIGURES 55–56. Labels of type specimens. (55)—O. (P.) vittulatum, holotype; (56)—O. (P.) alluaudi, Type.