Beutelius sagitta ( Neboiss, 1989 ) Escalona & Lawrence & Ślipiński, 2020

Escalona, Hermes E., Lawrence, John F. & Ślipiński, Adam, 2020, The extant species of the genus Omma Newman and description of Beutelius gen. nov. (Coleoptera: Archostemata: Ommatidae: Ommatinae), Zootaxa 4728 (4), pp. 547-574 : 558-562

Beutelius sagitta ( Neboiss, 1989) , comb. nov.

( Figs 10–11 , 22–25 , 38 , 46, 48, 50 , 56 , 68 , 73, 75 , 82–83 , 95 )

Omma sagitta Neboiss 1989: 110 .

Diagnosis. Antennae ( Fig. 10 ) moniliform, shorter than head and prothorax combined. Prothorax slightly transverse, about 0.8 times as long as wide (PL/PW). Prothorax with notopleural suture complete and sternopleural suture absent. Elytral lateral margins with white ribbed scales around the middle; epipleura with about 4 rows of ribbed scales. Outer surface of protibia in male with deep and large setose excavation at basal third. Parameres with translucent apical portion extended forward beyond apex of penis ( Fig. 95 ).

Description. Length: 9 mm (♂, ♀). Body 3.1 times as long as wide (BL/EW), dorsal surface clothed with ribbed dark brown scales, whited ribbed scales present on frons and scattered behind eye, at base of prothorax and forming patterns on elytra, ventral side with mixed dark brown setae and white ribbed setae. Head subquadrate, median length of visible portion about 1.0 times (HL/HW) as long as greatest width. Temples present. Frons slightly concave. Labrum about as long as wide, with apex truncate and bordered with pointed tubercles. Gulamentum deeply impressed to accommodate labial and maxillary palps. Mentum 1.4 times as long as wide, median area prominent with surface flat. Antennae relatively short ( Fig. 10 ) moniliform, shorter than head and prothorax combined; antennomere length/width ratios (♂), 1: 1.1, 2: 0.7, 3: 1.8, 4: 0.9, 5: 0.8, 6: 0.7, 7: 0.7, 8: 0.7, 9: 0.7, 10: 0.6, 11: 0.8.

Prothorax slightly transverse, about 0.8 times as long as wide (PL/PW); prothorax with notopleural suture complete and sternopleural suture absent; prosternal process extended between procoxae. Elytra about 2.1 times as long as wide (EL/EW) and 4.0 times as long as pronotum (EL/PL), lateral margins with white ribbed scales around middle; epipleura broad, with about 4 rows of ribbed scales, white ribbed scales concentrated in basal third, internal edge lined with row of white setae. Outer surface of protibia in male with deep and large setose excavation at basal third. Protarsus (♂) 0.7 times as long as protibia (TIL/TAL). Mesotarsus 0.8 times as long as wide mesotibia (TIL/TAL). Metatarsus 1.4 times as long as long as metatibia (TIL/TAL). Aedeagus with apex of penis emarginate, parameres with translucent apical portion extended forward beyond apex of penis ( Fig. 95 ).

Remarks. This species was previously known only from the holotype (♀), its type locality is about 200 km east of Perth in WA. The new records extend considerably (about 2000 km) the distribution of the species.

Material examined. SOUTH AUSTRALIA: New Records: Eyre Pen. Nr. Sinclair Cap Salt Lake, 33.12°S 137.05°E, in yellow bucket, 18.xii.1999, S. Baker (1 ♂, SAMA-25-038605). Uno Stn. 4.3km NW Harris Bluff, 32°40’27”S 136°42’54”E, 28.xii.2009 – 2.iii.2010, in red bucket, Mallee, P. Hudson (1 ♀, SAMA-25-038606). WESTERN AUSTRALIA: Kellerberrin, W Crawshaw, Brit Mus 1927-139, Omma sagitta sp. nov. det. by Neboiss 1988, Omma mastersi det. by Lea, (1 ♀, holotype, NHML).

Figures

FIGURES 10–13. Habitus. 10–11, Beutelius sagitta: 10, dorsal (♀ 9 mm, SAMA); 11, same, ventral; 12–13, B. rutherfordi: 12, dorsal (♂ 6 mm, SAMA); 13, same, ventral.

FIGURES 22–26. Beutelius sagitta (♀): 22–25. Head: 22, dorsal, 23, ventral, 24, latero–ventral, 25, frontal; 26, B. mastersi (♀) habitus, lateral.

FIGURES 33–38. Beutelius rutherfordi (♂): 33–37. 33, head, dorsal, 34, head, ventral, 35, head, lateral, 36, head, dorsal (paratype ♂), 37, antenna. 38, B. sagitta, antenna.

FIGURES 45–50. Beutelius mastersi: 45, 47, 49. 45, pronotum, 47, prothorax, ventral, 49, prothorax, lateral; B. sagitta: 46, 48, 50. 46, pronotum, 48, prothorax, ventral, 50, prothorax, lateral.

FIGURES 56–63. Beutelius sagitta (♂): 56, epipleuron, ventral view; B. rutherfordi (♂): 57, epipleuron, ventral view; O. stanleyi (♀) elytron: 58–60, 63. 58, ventral view with epipleuron, 59, dorsal, 60, internal, 63 dorsal; B. mastersi (♀) elytron: 61–62. 61, dorsal, 62, ventral view with epipleuron.

FIGURES 67–70. Beutelius mastersi (♂): 67, wing (veinal nomenclature as in Fig. 64); B. sagitta (♂): 68, wing; Omma stanleyi: 69–70. 69, elytron base, 70, elytron, close detail of sutural flange.

FIGURES 71–75. Omma stanleyi and Beutelius spp. abdominal ventrites and tergites: 71, B. rutherfordi (♂, holotype); 72, B. mastersi (♀) ventrites; 73, B. sagitta (♂) ventrites; 74, O. stanleyi (♀) ventrites; 75, B. sagitta (♂), tergites.

FIGURES 76–84. Omma stanleyi (♀): 76–78. 76, foreleg, 77, midleg, 78, hind leg. Forelegs (♂): 79–84. 79, Beutelius rutherfordi; 80–81, O. stanleyi, 80, anterior view, 81, lateral view; 82–83, B. sagitta, 82, anterior view, 83, posterior view; 84, B. mastersi, anterior view.

FIGURES 85–95. Beutelius spp., mentum: 85–86. 85, B. mastersi, 86, B. rutherfordi (paratype ♂); 87, B. mastersi, mesonotum; 88, B. mastersi, pterothorax, ventral; 89, B. rutherfordi, pterothorax, ventral; aedeagus: 90–95. 90, Omma stanleyi; 91–92, B. mastersi; 93, B. reidi sp. nov.; 94, B. rutherfordi; 95, B. sagitta.