Odites obitsyella ( Viette, 1986 ) Park & Koo & Minet, 2020

Park, Kyu-Tek, Koo, Jun-Mo & Minet, Joël, 2020, Review of the Malagasy lecithocerid species described by Pierre Viette and deposited in MNHN (Paris), with new generic combinations and descriptions of a new subfamily and genus of Momphidae (Lepidoptera: Gelechioidea), Zootaxa 4845 (2), pp. 151-190 : 176-178

publication ID	https://doi.org/ 10.11646/zootaxa.4845.2.1
publication LSID	lsid:zoobank.org:pub:184EB7D9-A6F8-44D4-B7BD-FBCC0B3466E4
DOI	https://doi.org/10.5281/zenodo.4488464
persistent identifier	https://treatment.plazi.org/id/0392796A-FFAE-FFC7-5CE1-9BA4FE00FC9A
treatment provided by	Plazi
scientific name	Odites obitsyella ( Viette, 1986 )
status	comb. nov.

Treatment

Odites obitsyella ( Viette, 1986) , comb. nov. (Oditinae)

( Figs. 19 View FIGURE 19 A–J)

Idiopteryx obitsyella Viette, 1986: 116 . TL: Zombitsy Special Reserve.

Type. 1♀ (holotype), West Madagascar, east of Sakaraha, Zombitsy Special Reserve, matsabory, alt. 640 m, 13 i 1974, P. Viette and A. Peyrieras leg.; genitalia: prep. P. Viette no. 5706 .

Adult ( Figs. 19 View FIGURE 19 A–E). Wingspan, 19 mm. Head orange yellow, with slender, erect scales on vertex ( Fig. 19C View FIGURE 19 ). Antenna black, with whitish apex; its proximal third orange yellow ventrally. Labial palpus extremely large, sickleshaped; first two segments orange yellow, third segment partially black. Thorax orange yellow. Abdomen black dorsally. Legs orange yellow, with blackish/greyish zones on tarsi, notably on fore- and midlegs. Tibial spurs 0-2-4; the medial pair of hindleg from middle of tibia ( Fig. 19D View FIGURE 19 ). Dorsal surface of forewing: black, with orange yellow base; fringe yellow with two black regions: a small one at the apex, a large one at the tornus. Forewing ventral surface: like dorsal surface but with reduced, ill-defined basal yellow spot. Hind wing dark grey on both surfaces; fringe greyish (becoming whitish distally). Forewing venation ( Fig. 19B View FIGURE 19 ): all veins free, separate, except for R 4 and R 5 which are stalked (the latter running to termen, just below apex); R 2 arising approximately midway between R 1 and R 3 (slightly nearer R 3); bases of R 3 and M 1 both moderately remote from R 4+5; M 2 arising closer to M 3 than to M 1; CuA 1 and CuA 2 arched, close to each other proximally; CuA 1 arising near M 3. Hind wing venation ( Fig. 19B View FIGURE 19 ): Rs and M 1 long-stalked; upper edge of discal cell very short, slightly shorter than stem Rs+M 1; discocellular m1- m2 long, strongly oblique; M 3 and CuA 1 short-stalked; M 2 arising near M 3 +CuA 1. Abdominal segments ( Fig. 19J View FIGURE 19 ) without spinose zones; sternum II without anterolateral processes but with short apodemes, distinct venulae, and an arched, transverse ridge near its anterior edge.

Female genitalia ( Figs. 19 View FIGURE 19 F–I): See also the line drawing given by Viette (1986, Fig. 14 View FIGURE 14 ). Ductus bursae narrow, very long, about 3 times the length of the corpus bursae, with an elongate, colliculate, heavily sclerotized plate in posterior 2/5; ductus seminalis narrow, arising approximately from middle of ductus bursae. Corpus bursae rounded, with a pair of large scobinate signa. Ovipositor elongate; apophyses posteriores thinner and much longer than apophyses anteriores (length ratio of ca. 3 to 1).

Distribution. Madagascar (West) ( Viette, 1986).

Remarks. “ Idiopteryx ” obitsyella Viette, 1986 is here transferred to the Oditinae (Peleopodidae) and, tentatively, to Odites Walsingham, 1891 , a speciose genus in Madagascar (and elsewhere). It differs from most Lecithoceridae in its vertex vestiture, mainly composed of slender erect scales (while vertex tends to be smooth-scaled in Lecithoceridae ), and in its metathoracic coxae, which lack the rows of erect piliform scales that are normally present, postero-ventrally, in the Lecithoceridae . Moreover, Odites obitsyella shows, on the dorsal surface of the hind wing, a few piliform scales that arise from the lower edge of the discal cell; as mentioned above (see remarks about Mexytocerus ), we regard this “cubital pecten” as a previously unnoticed autapomorphy of the Oditinae (nevertheless also present, through parallel evolution, in at least core Xyloryctidae and Gelechiidae : Dichomerinae). Typical oditine characters found in O. obitsyella also include: very large, slender, sickle-shaped labial palpi; hind wing with long-stalked Rs and M 1, a very short upper edge of discal cell, and a long, markedly oblique discocellular m1-m2 (compare with the figures given by Lvovsky (1996)); elongate ovipositor (with long apophyses posteriores); on abdominal sternum II, presence of a well indicated, narrow transverse ridge, which lies near the anterior edge of the sternum ( Fig. 19J View FIGURE 19 ). Occurrence of the last trait (also present in Mexytocerus ) requires however further checking in the subfamily Oditinae .