Novichneumoninae Li et al., 2017

Kopylov, Dmitry S., Zhang, Qi & Zhang, Hai-Chun, 2021, The Darwin wasps (Hymenoptera: Ichneumonidae) of Burmese amber, Palaeoentomology 4 (6), pp. 592-603 : 593-594

Subfamily Novichneumoninae Li et al., 2017

Type genus. Novichneumon Li et al., 2017 .

Included genera. 6 genera with 8 species. Novichneumon Li et al., 2017 , Caloichneumon Li et al., 2017 , Heteropimpla Li et al., 2019 , Rasnichneumon gen. nov., Heteroichneumon gen. nov. and Rogichneumon gen. nov.

Diagnosis (based on Li et al. (2017), with improvements). Mandibles bidentate. Antennae filiform, with 9–20 flagellomeres. Notauli converging, sometimes weak or absent; mesonotum smooth to rough, never with coarse sculpture. Legs slender, with simple pretarsal claws. Fore wing with 1-Rs base strongly displaced basad (except Caloichneumon ); 1-Rs+M reduced or only preserved as a stub (ramulus); areolet with 2-Rs as long as, or longer than 2+3-M, meeting at obtuse or, rarely, right angle, rm usually absent (except Heteropimpla ); radial cell (3r) ending far proximal of the fore wing apex; 2m-cu with one bulla, usually straight, rarely slightly arched; 1-Cu very long, 0.2–0.5× as long as 2-Cu. Hind wing with 1-Rs short or absent, rarely (in Heteropimpla ) as long as r-m, but never longer; 2-Cu usually absent, when present (in Heteropimpla and Heteroichneumon ), nervellus broken at or below middle. Free endings of longitudinal veins in fore and hind wings usually reaching or almost reaching wing margin. Propodeum areolated. The first metasomal segment stout, with glymmae. Ovipositor medium-sized (0.3–0.5x as long as fore wing length).

Key to the genera and species of Novichneumoninae

- Fore wing with r-m absent. Hind wing usually without 2-Cu (excepting Heteroichneumon ) .......................................... 3

2. Fore wing with 1-Rs&1-M twice as long as r-rs, 2-Rs+M as long as 1m-cu, 2m-cu reclined converging to 3-Cu; hind wing with 1-Rs 0.5× as long as r-m...................................... ..................................................... H. megista Li et al., 2019

- Fore wing with 1-Rs&1-M equals r-rs, 2-Rs+M significantly longer than 1m-cu, 2m-cu proclined, parallel to 3-Cu; hind wing with 1-Rs equals r-m........... H. pulverulenta sp. nov.

3. Antennae with ≥19 flagellomeres..................................... 4

- Antennae with ≤ 12 flagellomeres.................................... 6

4. Fore wing with 2-Rs twice as long as 2+3-M. Notauli weak. Genus Novichneumon Li et al., 2017 .................................... ....................................................... N. longus Li et al., 2017

- Fore wing with 2-Rs nearly as long as 2+3-M. Notauli strong ................................................................................ 5

5. Notauli joining before scutellum. Hind wing with 1-Rs 0.5× as long as r-m, nervellus broken below middle, 2-Cu present. Genus Heteroichneumon gen. nov.......................... ........................................................... H. rasnitsyni sp. nov.

- Notauli converging, but not joining. Hind wing with 1-Rs vanishingly short, nervellus not broken, 2-Cu absent. Genus Rogichneumon gen. nov.............. R. braconidicus sp. nov.

6. Antennae with 9 flagellomeres. Fore wing with 2-Rs twice as long as 2+3-M and cell 1mcu as long as 2a. Genus Caloichneumon Li et al., 2017 . C. perrarus Li et al., 2017

- Antennae with 12 flagellomeres. Fore wing with 2-Rs as long as or shorter than 2+3-M and cell 1mcu much longer than 2a. Genus Rasnichneumon gen. nov......................... 7

7. Fore wing with r-rs issuing before pterostigma midlength, twice as long as distance between r-rs and m-cu; 2m-cu reclined; hind wing with 2-Rs and 2-M reaching wing margin................................................ R. alexandri sp. nov.

- Fore wing with r-rs issuing after pterostigma midlength, as long as distance between r-rs and m-cu; 2m-cu proclined; hind wing with 2-Rs as short stub, 2-M lost....................... ............................................................... R. gracilis sp. nov.