Myriophora longisetarum, Hash, John M. & Brown, Brian V., 2015

Hash, John M. & Brown, Brian V., 2015, Revision of the New World Species of the Millipede-Parasitic Genus Myriophora Brown (Diptera: Phoridae), Zootaxa 4035 (1), pp. 1-79 : 32

Myriophora longisetarum sp. nov.

(Figs 5.4, 12.2, 18.2, 23.10)

Diagnosis. Female. Myriophora longisetarum is similar to M. kungae , but has a more acutely downturned oviscape apex and distinctly longer setae on the sternite (Figs 18.2, 23.10).

Description. Female (Fig. 5.4). Body length 1.33–2.10 mm. Head: Frons dusky yellow, with 2 pairs of supraantennal setae; length of lower supra-antennal setae less than 1/2 the length of upper pair; upper supra-antennals originate even with lower interfrontal setae. First flagellomere yellow-brown, rounded under arista. Palpus yellow, slightly inflated. Labrum normal sized. Labellum normal sized. Thorax: Scutum dusky yellow. Scutellum dusky yellow. Anterior scutellar setae about 3/4 the length of posterior pair. Pleuron yellow; anepisternum with setulae and one large seta. Forefemur yellow. Midfemur yellow. Hind femur yellow with brown patch distally on anterior surface. Wing length 1.23–2.31 mm (Fig. 12.2); costal vein normal; costal length 0.70–1.23 mm; mean costal ratio 0.55. Knob of halter light brown. Abdomen: Tergites 1 and 2 brown; tergite 2 with strong, distinct setae on lateral margin. Tergite 5 brown; posterior setae on tergite 5 short. Tergite 6 rectangular to slightly narrowing posteriorly; posterodorsal setae on segment 6 shorter than posteroventral setae. Venter of abdomen with dense, long setae. Oviscape (Figs 18.2, 23.10) sclerites dark brown; tergite strongly downturned at apex; sternite about 1/5 of the length of tergite and with long setae on lateral margins.

Distribution. Mexico to Colombia.

Host. Unknown.

Etymology. A noun from Latin for "long setae" referring to the long setulae on the ventrolateral side of the oviscape that differentiates this species from the closely related M. kungae .

Holotype. ♀, MEXICO: Tamaulipas: Estación Los Cedros, Gómez Farías, 23.05°N, 99.15°W, 340m, 1.iv.2002, A. Cordoba-Torres, Malaise trap [ LACM ENT 163253].

Paratypes. 7♀, same as holotype, [ LACM ENT 163286, 163283, 163252, 163130, 163264, 163266, 163201].

Other material examined. COLOMBIA: Cundinamarca: PNN Chingaza, Charrascales, 4.52°N, 73.75°W, 2990m, 1♀, 11.i–8.ii.2002, E. Raigoso, CAP-3021, Malaise trap, 2♀, 8–22.ii.2002, E. Raigoso, CAP-3047, Malaise trap; Valle de Cauca: PNN Farallones de Cali, Alto Anchicaya, 3.43°N, 76.8°W, 650m, 1♀, 19.xii– 2.i.2001, S. Sarria, CAP-1539, Malaise trap. COSTA RICA: Alajuela: 2km W Dos Rios, 10.88°N, 85.38°W, 600m, 2♀, viii.1988, San Ramon, 10.22°N, 84.62°W, 900m, 1♀, vii–viii.1995, P. Hanson, Malaise trap; Cartago: 4km NE Cañon, Genesis II, 9.71°N, 83.91°W, 2350m, 1♀, ii.1995, P. Hanson, Malaise trap, 2♀, iv.1995, P. Hanson, Malaise trap, 1♀, ix.1995, P. Hanson, Malaise trap, 1♀, v.1995, P. Hanson, Malaise trap, Tapanti, 9.76°N, 83.78°W, 1200m, 1♀, 17–20.vii.2000, S. Ashe, R. Brooks, FIT#192; Guanacaste: Cerro Pedregal, Volcan Cacao, 10.93°N, 85.48°W, 1000m, 6♀, ii–iv.1989, I. Gauld, D. Janzen, Malaise trap, Estación Cacao, Volcán Cacao, 10.93°N, 85.47°W, 1000m, 1♀, 10–11.vii.2000, R. Brooks, S. Ashe, FIT #97; Heredia: 6km ENE Vara Blanca, 10.18°N, 84.12°W, 2000m, 1♀, 21.iv.2002, ALAS, Malaise trap 20/M/20/100, La Selva Biological Station, 10.43°N, 84.02°W, 40m, 2♀, 1–15.iii.1993, ALAS, Malaise trap M/07/038 ( INBC), 1♀, 1–15.iv.1993, ALAS, Malaise trap M/07/070 ( INBC), 1♀, 1–15.v.1993, ALAS, Malaise trap M/10/104 ( INBC), 1♀, 1.vi.1993, ALAS, Malaise trap M/07/113 ( INBC), 1♀, 10.i.2000, ALAS, Malaise trap M/19/739 ( INBC), 1♀, 14.vi.1996, ALAS, Malaise trap M/12/673 ( INBC), 1♀, 15–17.ii.2002, Brown, Gonzalez, Walker, Malaise trap, successional plots, 1♀, 15.ii–1.iii.1993, ALAS, Malaise trap M/10/025 ( INBC), 1♀, 15.iii–1.iv.1993, ALAS, Malaise trap M/08/055 ( INBC), 1♀, 16–23.v.1988, B.V. Brown, Malaise trap SSO 50, 1♀, 16.ii–2.iii.1993, ALAS, Malaise trap M/07/022 ( INBC), 1♀, 16.viii.1995, ALAS, Malaise trap M/10/431 ( INBC), 1♀, 18.v.1993, ALAS, Malaise trap M/07/101 ( INBC), 1♀, 2.v.1993, ALAS, Malaise trap M/07/086 ( INBC), 1♀, 25.iii.1999, ALAS, Light L/18/607 ( INBC), 1♀, 26.vi–1.vii.1993, B. Brown & D. Feener, Malaise trap #3, 1♀, 27.xii.1999, ALAS, Malaise trap M/19/738 ( INBC), 1♀, 3.iv.2000, ALAS, Malaise trap M/19/745 ( INBC), 1♀, 30.vi.1995, ALAS, Malaise trap M/01/387 ( INBC), 1♀, 4.iv.1994, ALAS, Malaise trap M/09/387 ( INBC), Refugio 2000m, near Varablanca, 10.18°N, 84.11°W, 2000m, 1♀, 10.iii.2002, ALAS, Malaise trap 20/M/17/037, 1♀, 21.ii.2002, ALAS, Malaise trap 20/M/ 20/020, 1♀, 9.iv.2002, ALAS, Malaise trap 20/M/17/077; Puntarenas: 24km W Piedras Blancas [should be Chacarita], 8.7593°N, 83.3861°W, 200m, 3♀, ix–xi.1989, P. Hanson, Malaise trap, 3km SW Rincon, 8.68°N, 83.48°W, 10m, 1♀, ix–xi.1989, P. Hanson, Malaise trap, 1♀, vii–ix.1990, P. Hanson, Malaise trap, 1♀, viii.1991, P. Hanson, Malaise trap, 1♀, xii.1989, P. Hanson, Malaise trap, Cerro Rincon, 8.52°N, 83.47°W, 745m, 1♀, ii.1991, P. Hanson, Malaise trap, Las Cruces, 8.80°N, 82.97°W, 1200m, 1♀, 7–17.xi.2005, G. Kung, A. Kreuter, Malaise trap #1, Monteverde, 10.33°N, 84.79°W, 1700m, 1♀, 1–5.vi.1988, B.V. Brown, Malaise trap, Rios Paraisos, Albergue del Pecori, 9.56°N, 84.12°W, 400m, 1♀, 15–17.ii.2003, B. Brown, E. Zumbado, Malaise trap #2; San José: 6km N San Gerardo, 9.55°N, 83.8°W, 2800m, 2♀, i.1993, P. Hanson, Malaise trap, 1♀, xi.1992, P. Hanson, Malaise trap, 1♀, xii.1992, P. Hanson, Malaise trap, Braulio Carrillo NP, 9.5km E Tunel, 10.12°N, 83.97°W, 1000m, 1♀, x–xii.1989, P. Hanson, Malaise trap, Zona Protectora El Rodeo, 9.91°N, 84.28°W, 1000m, 1♀, 1– 5.viii.2001, B. Brown et al., Malaise trap #2 (upper site), 1♀, 1–5.viii.2001, B. Brown et al., Malaise trap, lower site, Zurquí de Moravia, 10.05°N, 84.02°W, 1600m, 1♀, 17–28.iii.2007, M. Pollet, blue pan traps, CR/2007/306, 1♀, i.1996, P. Hanson, Malaise trap, 1♀, vi.1995, P. Hanson, Malaise trap. GUATEMALA: Sacatepequez: Dunwest Farm, Sumpango, 14.67°N, 90.72°W, 1975m, 1♀, 18.ii–4.iii.2007, M. Hoddle, Malaise trap. MEXICO: Chiapas: Reserva El Triunfo, Sendero Cerro Triunfo, 15.66°N, 92.8°W, 1982m, 1♀, 19–22.vii.1997, J. Woolley, Gonzalez, Malaise trap; Tamaulipas: Estación Los Cedros, Gómez Farías, 23.05°N, 99.15°W, 340m, 1♀, 22.v.2002, S. Triapitsyn, V. Berezovskiy, Malaise trap, 3♀, 23.iv.2002, S. Triapitsyn, V. Berezovskiy, Malaise trap, 1♀, 3.vi.2002, S. Triapitsyn, V. Berezovskiy, Malaise trap, 1♀, 4.iii.2003, S. Triapitsyn, V. Berezovskiy, Malaise trap, 2♀, iii.2002, A. Cordoba-Torres, Malaise trap. NICARAGUA: Matagalpa: Fuente Pura, 13.02°N, 85.92°W, 1500m, 1♀, 6.vii.1994, J. M. Maes, Malaise trap, 1♀, vi.1993, J. M. Maes, Malaise trap. PANAMA: Veraguas: Cerro Tute, 8 km W Santa Fe, 8.50°N, 81.11°W, 1000m, 1♀, 24.vii–8.viii.1999, J. Woolley, Malaise trap.