Monotes magnificus Gilg var. gigantophyllus (P.A.Duvign.) Meerts, 2017

Meerts, Pierre, Rougelot, Quentin & Sosef, Marc, 2017, Revision of the genus Monotes (Dipterocarpaceae) in D. R. Congo, with implications for Angola and its distinction from Marquesia, Phytotaxa 308 (2), pp. 151-205 : 192

11b. Monotes magnificus Gilg var. gigantophyllus (P.A.Duvign.) Meerts , comb. et stat. nov.

Basionym: Monotes gigantophyllus Duvigneaud (1949: 56) ; Lebrun & Stork (1991: 144); Meerts (2016: 223).

Type: —D.R. CONGO. Upper Katanga, in sylvis siccis prope Kasenga, 19 August 1948, Duvigneaud 1395M (holo-: BRLU! [barcode BRLU0000090020698], iso-: BRLU (3 sheets)).

Diagnosis: — Differs from the type variety in being a tortuose low shrub up to 3 m high, with much larger leaves 25–45 × 17–30 cm, bigger fruits 20–35 mm in diameter and fruit wings 8–11 cm long.

Illustration: — Fig. 5J .

Distribution in D.R. Congo: —Upper Katanga ( Fig. 16E ). Endemic of D.R. Congo.

Habitat and ecology: —Wooded savannah, open miombo woodland, dambos, grassy savannah, occasionnally on slightly mineralised soil.

Other representative specimens examined:—D.R. CONGO. Upper Katanga: Kasenga, forêt claire, 20 August 1949, Duvigneaud 1400 (BRLU!); Kakontwe, dembo à Brachystegia stipulata en contrebas des collines rocheuses, 15 July 1956, Duvigneaud & Timperman 2031Mo (BRLU!); Mukumbi, dembo incendié à Cryptosepalum , Loudetia simplex , 3 August 1956, Duvigneaud & Timperman 2196 (BRLU!); Kisenge-Kamata, haute savane sur épaisse terre brune manganésifère, 16 August 1956, Duvigneaud & Timperman 2324 (BRLU!); Kisenga-Kamata, forêt sur sol léger à Brachystegia longifolia , 18 August 1956, Duvigneaud & Timperman 2364 (BRLU!); Mitwaba, 26 km au sud du poste, steppe sec arbustif à Uapaca robynsii sur 70 cm de limon ocre sur grenaille latéritique, Duvigneaud & Timperman 2731 (BRLU!); Route Sogefor, steppe à Vellozia sur colline minéralisée, 19 June 1957, Duvigneaud 3588 (BRLU!); Tilwizembe, verger de pente enrochée en contrebas du dembo d’empoisonnement, 15 December 1959, Duvigneaud 4597 (BRLU!); Shinkolobwe, dembo arbustif dans une plaine latéritique, 23 December 1959, Duvigneaud 4752M (BRLU!); Village Kilembi, près de Katofio, bord de route, 1230 m, Lukuesa 242 (LSHI!); Tilwizembe, forêt claire, 23 September 1956, Plancke 133/1792 (BRLU!); Route Katofio-Msipashi, contreforts des Kundelungu, forêt claire à Brachystegia , 9 September 1954, Schmitz 4653 (BR!); Kilembi, près de Katofio, bord de route près du village, 5 September 1957, Symoens 242 (BRLU!).

Discussion:— M. gigantophyllus and M. magnificus differ in no qualitative trait, and the variation in leaf size is virtually continuous. Therefore, species rank does not seem justified and the varietal level was deemed the mostappropriate.

Figures

FIGURE 5 A–H. SEM photographs of leaf blade surfaces for 12 members of Monotes. A. M. adenophyllus var. adenophyllus, lower surface (Delevoy 524, type of M. delevoyi). B. M. autennei, lower surface (Duvigneaud 3267 (type of M. autennei)). C. M. autennei, upper surface (Duvigneaud 3267). D. M. hypoleucus var. angolensis, lower surface (Duvigneaud 950M2, type of M. oxyphyllinus). E. M. hypoleucus var. angolensis, upper surface (Duvigneaud 950M2). F.M. hypoleucus var. caloneurus, lower surface (Schmitz 1332 (type of M. schmitzii)). G. M. hypoleucus var. caloneurus, upper surface (Schmitz 1332 (type of M. schmitzii)). H. M. hypoleucus var. hypoleucus, lower surface (Welwitsch 1036 (type of M. hypoleucus)).

FIGURE 16. Distribution maps.A.Monotes duvigneaudii var.duvigneaudii: Θ; var.concolor:z; B. M.rubriglans subsp.upembensis var. griseocoriaceus: ·; C. M. rubriglans subsp. upembensis var.upembensis:; D. M. doryphorus: „; E. M.magnificus var. gigantophyllus: z; M. hirtii: ·; F. M. dasyanthus var. heterotrichus:.