Miridiba borneensis ( Moser, 1918 ) Gao & Coca-Abia, 2021

Gao, Chuan-bu & Coca-Abia, María Milagro, 2021, Revision of the genus Miridiba Reitter, 1902 (Coleoptera, Scarabaeidae, Melolonthinae): genital morphotypes and new taxonomic data, European Journal of Taxonomy 749, pp. 1-94 : 68-70

Miridiba borneensis ( Moser, 1918) comb. nov.

Figs 15 , 87

Holotrichia borneensis Moser, 1918: 317 (type loc.: Borneo, Malaysia).

Pentelia borneensis – Matsumoto 2005: 479, figs 2, 6, 12, 17, 22, 26, 30, 33, 37, 40 (combination).

Diagnosis

External morphology of adult ( Fig. 87 ). Body size 19.0 mm. Dorsal surface glabrous. Clypeus almost as long as frons, with conspicuous punctures, rounded laterally, anterior edge strongly emarginate at middle. Frons punctate; carina developed, blunt. Anterior pronotal margin glabrous, regularly flanged; posterior margin glabrous, with a row of punctures near each posterior angle; lateral margins serrated and pubescent; anterior angles almost right and elevated, posterior angles obtuse and rounded. Prosternal process trapezoidal, sunk at middle. Scutellum irregularly punctate and glabrous. Elytral surface punctate, glabrous, costae weakly defined. Foretibia with dorsal carina moderately developed. Metatibia with transverse carina complete and inner margin of dorsal surface with few small spines. Pygidium with strong punctures irregularly distributed; surface glabrous, at most with a tiny seta on each puncture hardly visible, posterior margin with long pubescence on apex. Ventrite 1 not visible. Ventrites 2, 3 and 4 sparsely punctate, without conspicuous pubescence. Ventrite 5 with strong punctures and long pubescence irregularly distributed at posterior half. Ventrite 6 concave moderately bulging at anterior margin; punctures irregularly distributed and long pubescence near posterior margin. Male genitalia: parameres ( Fig. 15 ) with dorsal branches wrinkled, separated from each other and convergent at distal end. Ventral branches elevated at middle and apices strongly curved downward. Parameres and phallobase joined by a membrane oval-shaped. Endophallus and female not studied.

Material examined

Lectotype (here designated)

MALAYSIA • ♂; Borneo , Sarawak; Oct. 1911; Kling Kang leg.; “ Miridiba borneensis Chuan-bu Gao det. 2018”; MFNB.

Other material

MALAYSIA • 1 ♂; Borneo ; MFNB .

Remarks

Moser (1918) described Holotrichia borneensis from Borneo ( Sarawak) without specifying the number of specimens studied. Matsumoto (2005) transferred this species to the genus Pentelia ( Brenske 1891) . The first author of present paper has studied a syntype male of H. borneensis (MFNB) , which is designated herein as lectotype of M. borneensis , just in case new syntype specimens are found in the future.

Distribution

Malaysia (Borneo).

Figures

Figs 15–17. Morphotype VI “Borneensis”. 15. Parameres of M. borneensis (Moser, 1918), lectotype (BMNH) in dorsal (A) and lateral (B) views. 16. Endophallus and temones of M. coxalis (Arrow, 1944), lectotype (BMNH) in dorsal view. 17. Female genitalia of M. coxalis, paralectotype (BMNH) in dorsal (A) and ventral (B) views. Scale bars = 1.0 mm.

Figs 87–89. 87. Miridiba borneensis (Moser, 1918), lectotype, ♂ (MFNB). Habitus in dorsal view. 88–89. M. coxalis (Arrow, 1944), lectotype, ♂ (BMNH). 88. Habitus in dorsal view. 89. Parameres in dorsal (A), ventral (B) and lateral (C) views. Scale bars: 87–88 = 5.0 mm; 89 = 1.0 mm.