Mimophytum alienoides (G.L.Nesom) Holstein & Weigend, 2016

Holstein, Norbert, Chacón, Juliana, Otero, Ana, Jiménez-Mejías, Pedro & Weigend, Maximilian, 2016, Towards a monophyletic Omphalodes-or an expansion of North American Mimophytum, Phytotaxa 288 (2), pp. 131-144 : 139

1. Mimophytum alienoides (G.L.Nesom) Holstein & Weigend comb. nov.

Basionym: Omphalodes alienoides G.L.Nesom, Phytoneuron 2013 -64 (2013: 10, figs 1, 2, 3, 5).

Holotype:— USA. Texas: Val Verde Co., N-facing ledge of Pecos River below the park overlooking the mouth of the river, corolla light blue at the edge and white in the center, 3 April 1965, D. S. Correll & H. B. Correll 30816 ( LL00035424 , isotypes GH00294204 , SRSC00001251 !).

Type:— MEXICO. Coahuila: Canyon E side of Sierra de las Margaritas ca 13 km N of Las Margaritas, 1100– 1400 m, limestone and gypsum slopes, also limestone gravel arroyo, matorral desertico inerme, also some mesquital, Larrea , Viguiera stenoloba , Parthenium incanum , Agave lechuguilla , Hechtia , 23 March 1973, M. C. Johnston et al. 10357 A (paratype TEX). Type:— MEXICO. Coahuila: ca. 1 km W of Las Delicias at and near spring, top of alluvial fan on side of mountain, 1000–1300 m, calcareous gravel somewhat gypsiferous fan and limestone mountainsides, matorral desertico inerme, also some mesquital, Larrea tridentata , Flourensia cernua , Parthenium incanum , Viguiera stenoloba , 24 March 1973, M. C. Johnston et al. 10390 (paratypes MO-451150, TEX). Type:— MEXICO. Coahuila: Cuatro Cienegas Basin, tip of Sierra San Marcos, ridge, campsite, 3 April 1969, E. Lehto et al. 6180 (paratypes ASU 0020739, TEX). Type:— MEXICO. Coahuila: 18 km SW of Cuatrocienegas, 27 March 1992, Neff 92-3-27-1 (paratype TEX). Type:— MEXICO. Coahuila: Cuatro Cienegas Basin, Sierra San Marcos, NE-facing slope, near tip of mountain, bajada and mountain slopes, desert shrub, 19 March 1973, D. J. Pinkava 10369 (paratypes ASU 0020729!, DES 00015101, TEX). Type:— MEXICO. Coahuila: lomerias cercanos a la Poza de la Becerra en Cuatro Cienegas, veg. de Agave lechuguilla , Agave greggii , Larrea , Viguiera stenoloba , 7 Apri1984, Rodriguez y Carranza 1348 (paratypes ANSM fide J. Villarreal, TEX). Type:— MEXICO. Coahuila: Cañon de Jora, along road, 5.3 miles E of Est. Los Belloc and 3.8 mi W of Est. Socorro turnoff, major desert arroyo in low limestone canyon area with Chilopsis , Celtis pallida , Acacia berlandieri , Lycium berlandieri , Acacia neovernicosa , Viguiera stenoloba , Porophyllum scoparium , 1050 m, 19 March 1977, Wendt et al. 1961 (paratype TEX). Type:— USA. Texas: Brewster Co.: On N-facing slopes of Bullis Range, in the vicinity of Bullis Gap, on the Bullis Gap Ranch, infrequent annual herb among rock crevices, 17 May 1977, M. Butterwick & E. Lott 3601 (paratype TEX 00401507). Type:— USA. Texas: Boquillas Canyon (ca. 10 miles below W entrance), 11 February 1965, D. S. Correll & W. O. Douglas 30715 (paratype LL 00035422). Type:— USA. Texas: Reagan Canyon, ca. 1 mile above mouth, 500–600 m, USGS, Reagan Canyon Quad, steep rocky (limestone) slopes, abundant, 4 April 1947, R. McVaugh 7808 (paratypes DES 00033915, P 03512913!, TEX 00401508). Type:— USA. Texas: Burro Bluff, bluff on the Rio Grande (across from Tule Canyon, Coahuila), high limestone bluff, talus slopes, limestone gravel, associates Aloysia , Acacia , Rhus , Karwinskia , Agave , Hechtia , Euphorbia antisiph ., Opuntia , 21 March–6 April 1973, D. H. Riskind 1009 (paratype LL 00401509). Type:— USA. Texas: Santa Helena Canyon ledges, 19 March 1941, B. H. Warnock 21579 (paratype TEX 00401506). Presidio Co.: N facing cliffs, 15 miles SE of Redford, on limestone cliffs, 14 May 1959, D. S. Correll & I. M. Johnston 21915 (paratype LL 00401505). Type:— USA. Texas: ca. 15 mi SE of Redford, in shade of bluffs, in canyon, 21 April 1961, D. S. Correll & R. C. Rollins 23704 (paratype LL 00035421). Type:— USA. Texas: Terrell Co.: N side of current channel of Independence Creek, S side of Independence Creek Rd. from a point 3.6–3.7 road miles E of jct St. Rte. 349, Oasis Ranch Preserve, Oasis Ranch Quadrangle, common in tightly packed riverwashed gravel, sand and silt, in shade of thicket on floodplain, 1960 ft., associates include Juglans microcarpa , Morus sp. , Celtis reticulata , 4 April 2001, W. R. Carr 19471 (paratype TEX 00199780). Type:— USA. Texas: in short N-draining canyon on S side of Independence Creek, ca. 3.5–2.6 air miles E to ESE of jct Independence Creek Rd. and St. Rte. 349, Oasis Ranch Preserve ( TNC), Oasis Ranch Quadrangle, common among Quercus pungens var. vaseyana , Juniperus pinchotii , Diospyros texana and other tall shrubs on rocky slopes and bottom of steep-sided canyon cut into plateau of Edwards Limestone, 2000–2200 ft., in spring 2001, this species was abundant in stream terrace woodlands and locally frequent to common in mesic canyon woodlands, 5 April 2001, W. R. Carr & J. Maresh 19513 (paratype TEX 00199781). Type:— USA. Texas: Sanderson, common and abundant on rocky hillsides, 15 March 1919, H. C. Hanson 381 (paratypes TEX 00401510, US 01080292!). Type:— USA. Texas: Morton Canyon, Rio Grande ca. 25 mi S of Dryden, rocky limestone bluffs at dam site, 1 April 1947, R. McVaugh 7751 (paratypes P 03512911!, TEX 00035423). Type:— USA. Texas: Val Verde Co.: Pump Canyon, Langtry, on ledges, 15 April 1961, D. S. Correll & R. C. Rollins 23575 (paratype LL 00401500). Type:— USA. Texas: W side of Pecos River below Pandale, at roadcrossing 26 miles N of Langtry, sandy bank, scattered, but quite abundant, 30 March 1947, R. McVaugh 7712 (paratypes P 03512909!, TEX 00401502). Type:— USA. Texas: canyon of Pecos River 1 mile above mouth, at crossing of route US 90, steep rocky limestone slopes, abundant, 29 March 1947, R. McVaugh 7699 (paratypes P 03512910!, TEX 00401504). Type:— USA. Texas: W wall of Pecos River Canyon, in small draw between limestone bluffs, 23 March 1941, R. Rose-Innes & B. H. Warnock 603 (paratype TEX 00401501). Type:— USA. Texas: at mouth of Pecos River, frequent atop limestone bluffs, 29 March 1947, B. H. Warnock & R. McVaugh 47200 (paratype TEX 00401503). Type:— USA. Texas: Mile Canyon, Langtry, limestone soil, 1300 ft, frequent annual, 10 April 1949, B. H. Warnock et al. 224 (paratype LL 00401499).