Cymaenes isus (Godman, 1900)

Zhang, Jing, Cong, Qian, Shen, Jinhui & Grishin, Nick V., 2022, Taxonomic changes suggested by the genomic analysis of Hesperiidae (Lepidoptera), Insecta Mundi 2022 (921), pp. 1-135 : 72

Cymaenes isus (Godman, 1900) , reinstated status

Treated by Evans (1955) as a junior subjective synonym of Cymaenes trebius ( Mabille, 1891) (type locality Colombia: Bogota), Megistias isus Godman, 1900 (type locality Mexico, Guatemala, Nicaragua, Costa Rica, Venezuela and Guyana) may become the senior name for this species after we have shown above that Pamphila trebius Mabille, 1891 is a junior subjective synonym of Cymaenes lumina ( Herrich-Schäffer, 1869) . Cymaenes trebius was misidentified by Evans (1955), because its syntype specimen in the ZMHB (NVG-15034E04) actually (and correctly) keys out to “Sub-sp. odilia” by its characters. i.e. on its hindwing below “grey suffusion along the termen reaches to the discal [pale] band” ( Evans 1955: 133), and specimens from North America that key to Evans’ “Subsp. trebius” are in a different from the P.trebius syntype clade of Cymaenes Scudder, 1872 (type species Cobalus tripunctus Herrich-Schäffer, 1865 ) in the genomic tree ( Fig. 14 ). Indeed, Evans identified all the North American syntypes of M. isus in the BMNH collection as “Sub-sp. trebius” and no other name except M. isus can be applied to this species. Therefore, we reinstate Cymaenes isus (Godman, 1900) as a species-level taxon. Phylogenetic analysis reveals that C. isus is sister to the type species of the genus, C. tripunctus (type locality Cuba) ( Fig. 14 ). The type series of C. isus includes specimens from South America that represent a taxon different from North America specimens. To promote stability of nomenclature and follow Evans (1955) who listed the type locality of C. isus as “ Mexico ”, in addition to the original author of the taxon Godman ( Godman and Salvin 1900b) who illustrated specimens from Mexico, thus giving them more weight in the description, N.V.G. hereby designates one of these illustrated specimens, the male in BMNH collection that is already curated as “the type” with the following labels || Type / H 2222 || Type | H. T. || Venta de Zopilote, | Guerrero, 2800 ft. | Oct. H. H. Smith || ♂ || Sp. figured. || B. C. A. Lep. Rhop. | Megistias | isus, | Godm. || Godman-Salvin | Coll. 1914.–5 || as the lectotype of Megistias isus Godman, 1900 . The type locality of C. isus thus becomes Mexico: Guerrero, 22 km N of Chilpancingo, nr. southern end of the Canon de Zopilote at 2800 ft elevation, approximate GPS 17.77, −99.53, as detailed by Selander and Vaurie (1962), who researched the localities used by Godman and Salvin.

Figures

Figure 14. Genomic tree of Moncina, part 2. See Fig. 1 legend for notations.