Europiella knowltoni Knight, 1970: 228

SCHUH, RANDALL T., 2000, Revision of the North American Plant Bug Genus Megalopsallus Knight, with the Description of Eight New Species from the West (Heteroptera: Miridae: Phylinae), American Museum Novitates 3305, pp. 1-70 : 21-36

publication ID	https://doi.org/ 10.1206/0003-0082(2000)305<0001:ROTNAP>2.0.CO;2
persistent identifier	https://treatment.plazi.org/id/4A3FBB30-C673-8F42-C500-DEBD48BCF91A
treatment provided by	Carolina
scientific name	Europiella knowltoni Knight, 1970: 228
status

Treatment

Europiella knowltoni Knight, 1970: 228 View in CoL (n. sp.). Megalopsallus knowltoni: Schuh et al., 1995: 389

(n. comb.).

DIAGNOSIS: Recognized by large size and totally black coloration (fig. 9). Similar in size, general appearance, coloration, and strong sexual dimorphism to Dakota hesperia Uhler, but readily separated by male genitalic structure and hosts, hesperia breeding on Potentilla (Rosaceae) rather than Sarcobatus (Chenopodiaceae) . Within Megalopsallus , most similar in coloration to dark specimens of humeralis , nigrofemoratus , and teretis , but easily distinguished by much larger size.

REDESCRIPTION: Male: Large, total length 4.51–5.20, length apex clypeus–cuneal fracture 2.79–3.31, width across pronotum 1.15– 1.32. COLORATION: Black or nearly so, including eyes (fig. 9); antennal segments 2, 3, and 4 and tibiae lighter, but always at least weakly infuscate; tibial spines black with obscure dark bases. SURFACE AND VESTI TURE: Dorsum very faintly rugose, pronotum and scutellum weakly shining, hemelytra appearing dull; dorsum with long, dark, reclining simple setae intermixed with silvery, woolly setae (fig. 4B, C). STRUCTURE: Hemelytra extremely elongate, nearly parallelsided (fig. 9), apex of abdomen reaching only to cuneal fracture; labium just reaching middle trochanters; claws sharply curving near apex, pulvilli about onehalf the length of claw, free except at base (fig. 4D). MALE GENITALIA: Vesica Sshaped, attenuated apically; gonopore subapical, very weakly sclerotized; gonopore sclerite absent (fig. 14).

Female: Total length 3.62–3.98, length apex clypeus–cuneal fracture 2.75–2.96, width across pronotum 1.25–1.32; elongate ovoid (fig. 9).

HOST: Sarcobatus vermiculatus (Chenopodiaceae) .

DISTRIBUTION: Northern Great Basin.

DISCUSSION: The majority of specimens of M. knowltoni are labeled as having been taken on Sarcobatus vermiculatus .

SPECIMENS EXAMINED: USA. — California: Modoc Co.: 5 mi SE of Fort Bidwell, Surprize Valley Dunes, June 9, 1970, P. Opler, Sarcobatus vermiculatus (Chenopodiaceae) , 213 (UCB). Mono Co.: 8 mi W of Nevada state line on Rt. 359, 6700 ft, July 2, 1983, R. T. Schuh and M. D. Schwartz, Sarcobatus vermiculatus (Chenopodiaceae) , 43, 8♀ (AMNH). Idaho: Oneida Co. : Curlew Reservoir, June 6, 1969, G. F. Knowlton, 1♀ (USNM); 1♀ (USNM). Black Pine, March 31, 1969, G. F. Knowlton, 73 (USNM, USU); Paratypes: 13, 1♀ (CNC). Nevada: Elko Co.: Utah state line on Utah Rt. 30, 4760 ft, June 25, 1983, Schuh and Schwartz, Sarcobatus vermiculatus (Chenopodiaceae) , 13 (AMNH). Nye Co.: 28 mi N of Belmont on Rt. 82, 2013 m, July 13, 1980, R. T. Schuh, Sarcobatus vermiculatus (Chenopodiaceae) , 23 (AMNH). Washoe Co.: 2 mi E of Vya on Rt. 8A, 1750 m, July 2, 1979, Schuh and Massie, Sarcobatus vermiculatus (Chenopodiaceae) , 83, 12♀ (AMNH). Oregon: Harney Co.: Alvord Basin, T37S R33E S26, April 29, 1979, Cobb and Lightfood, Sarcobatus vermiculatus (Chenopodiaceae) , (OSU). Utah: Box Elder Co. : Kelton Pass, May 27, 1969, G. F. Knowlton, 13 (USNM).

Megalopsallus marmoratus Knight

Figures 9 View Fig , 15 View Fig

Megalopsallus marmoratus Knight, 1968: 27 View in CoL (n. sp.).

DIAGNOSIS: Recognized by brownish coloration and marmorate membrane (fig. 9). Most similar in overall coloration to brittoni and femoralis , but those species both lacking marmorate membrane. Marmorate membrane known to occur elsewhere in Megalopsallus only in californicus , that species otherwise being pale green.

REDESCRIPTION: Male: Moderately small, total length 3.00–3.26, length apex clypeus– cuneal fracture 1.99–2.20, width across pronotum 0.94–1.04. COLORATION: Generally tan, much of body and femora with darker brown or reddish spots; eyes brown; entire membrane marmorate (fig. 9); tibial spines pale with reddish or brownish bases. SUR FACE AND VESTITURE: Dorsum smooth, dull, rather sparsely clothed with recumbent, brown, simple setae intermixed with silvery, weakly flattened setae. STRUCTURE: Weakly elongate, nearly parallelsided (fig. 9); labium reaching well onto hind trochanters; claws elongate, curving; pulvilli minute. MALE GENITALIA: Vesica relatively short, Sshaped, with a short decurved apex; gonopore subapical; no gonopore sclerite (fig. 14).

Female: Total length 2.57–3.01, length apex clypeus–cuneal fracture 1.84 –2.16, width across pronotum 0.96–1.13; ovoid (fig. 14); body form elongate ovoid (fig. 9).

HOSTS: Allenrolfea occidentalis , Salicornia sp. (Chenopodiaceae) .

DISTRIBUTION: Arizona, southern California, and Baja California Norte.

SPECIMENS EXAMINED: MEXICO. — Baja California Norte: 1 mi S of Bahia de los Angeles, March 26, 1979, J. D. Pinto, Salicornia sp. (Chenopodiaceae) , 13, 1♀ (UCR). USA. — Arizona: Maricopa Co. : just S of Buckeye, Palo Verde Rd at Rt. 85, 335 m, April 1, 1981, R. T. Schuh and M. D. Schwartz, Allenrolfea occidentalis (Chenopodiaceae) , 163, 113♀ (AMNH, USNM). Yuma Co. : Yuma , April 27, 1939, L. L. Stitt, Paratypes: 23, (USNM). Yuma , May 5, 1942, L. L. Stitt, Paratypes: 33 (CNC, USNM). California: Imperial Co. : Holtville, March 13, 1945, Weed, 13 (USNM). Inyo Co. : Death Valley, Saratoga Springs, March 23, 1957, C. L. Hogue, 13 (LACM). Death Valley, Triangle Spa, April 16, 1943, G. Willett, 13 (LACM). Riverside Co.: Salton Sea, Salt Creek, April 14, 1974, M. Wasbauer, Salicornia sp. (Chenopodiaceae) , 13, 14♀ (CAFA). Salton Sea, Mecca, April 9, 1955, W. R. Richards, picklebush, 63, 6♀ (CNC).

Megalopsallus nicholi (Knight)

Figures 5 View Fig , 9 View Fig , 15 View Fig

Europiella nicholi Knight, 1968: 42 (n. sp.). Megalopsallus nicholi: Schuh et al., 1995: 389 (n.

comb.).

DIAGNOSIS: Recognized by castaneous head, pronotum, and scutellum contrasting with pale corium, clavus, and membrane, and cuneus always at least weakly darkened and reddish (figs. 5, 9). Most easily confused with pale specimens of humeralis (fig. 9), the latter also breeding on Lycium spp. ; distinguished from humeralis by darkened cuneus contrasting with remain der of hemelytra. Genitalia also absolutely distinctive from those of humeralis . Also similar in general appearance to Sarcoba tus feeding rubricornis (fig. 11), but that species always with cuneus unicolorous with remainder of pale hemelytra and head, with pronotum and scutellum often more strongly reddish.

REDESCRIPTION: Male: Moderately small, total length 3.02–3.48, length apex clypeus–cuneal fracture 1.95–2.24, width across pronotum 0.98–1.06. COLORATION: Head, pronotum, and undersurface castaneous or chocolate brown, scutellum partly or entirely, and sometimes femora also, brown; hemelytra and often much of scutellum cream colored, cuneus weakly to strongly reddish brown (fig. 9); eyes dark brown; antennae and tibiae pale; tibial spines dark with faintly darkened bases. SURFACE AND VESTITURE: Dorsum smooth, weakly shining, clothed with pale to brown recumbent simple setae intermixed with golden, shining recumbent setae and some patches of woolly silvery setae. STRUCTURE: Moderately elongate, nearly parallelsided (fig. 9); labium reaching to about posterior margin of middle trochanters; claws nearly straight and sharply curving at apex, pulvilli large and covering nearly entire ventral claw surface. MALE GENITALIA: Vesica elongate, Sshaped, apex in the form of a fingerlike projection adorned with tiny spicules; gonopore subapical, well developed; no gonopore sclerite (fig. 15).

Female: Total length 2.37–2.59, length apex clypeus–cuneal fracture 1.79 –1.95, width across pronotum 0.88–1.02; very broadly ovoid, rotund, membrane short (fig. 9).

HOSTS: Lycium andersonii , L. sp. ( Solanaceae ). Other reported hosts with limited confirmation: Shepherdia argentea (Eleagnaceae) ; Condalia globosa (Rhamnaceae) ; Solanum eleagnifolium . Probable sitting records: Quercus arizonica .

DISTRIBUTION: Arizona and Nevada south to Zacatecas, Mexico.

DISCUSSION: Knight (1968) described this species on the basis of specimens collected on Lycium torreyi . The majority of subsequent records are from Lycium spp. , although the records from Condalia and Shepherdia may indicate that those plants are occasionally used as hosts as well.

SPECIMENS EXAMINED: MEXICO. — Zacatecas: 13 mi SE of Concepcion del Oro, July 9, 1983, Schaffner, Kovarik, Harrison, 153, 18♀ (TAMU). 6 mi S of Concepcion del Oro, July 9, 1983, Kovarik, Harrison, and Schaffner, 463, 62♀ (TAMU). USA. — Arizona: Cochise Co. : July 29, 1927, R. H. Beamer, 1♀ (KU). Gila Co. : 5.5 mi W of Roosevelt Dam on Rt. 188, 2000 ft, May 27, 1983, Schuh, Stonedahl, and Massie, 12♀ (AMNH). Maricopa Co. : 5 mi S of Freeman SE of Gila Bend , 625 m, May 8, 1978, R. T. Schuh, A. F. Guenther, Lycium sp. (Solanaceae) , 13 (AMNH). 1.8 mi W of Roosevelt on Rt. 88, May 19, 1980, J. D. Pinto, Lycium sp. (Solanaceae) , 123, 12♀ (UCR). Pima Co. : Tucson, 2400 ft, April 19, 1926, A. A. Nichol, Lycium sp. (Solanaceae) , Paratypes: 13, 3♀ (USNM, CNC). Rincon Mts, 3300 ft, September 2, 1928, A. A. Nichol, 4♀ (USNM). Organ Pipe Cactus Natl. Mon., Quitobaquito, April 3, 1966, C. W. O’Brien, 13 (UCB). Organ Pipe Cactus Natl. Mon., Alamo Wash, 700 –800 m, R. T. Schuh and M. D. Schwartz, Lycium sp. (Solanaceae) , 203, 13♀ (AMNH). Organ Pipe Cactus Natl. Mon., Ajo Valley, April 10, 1981, D. A. Polhemus, Condalia globosa (Rhamnaceae) , 163, 7♀ (JTP). Baboquivari Mts, Sabino Canyon, April 20, 1982, D. A. and J. T. Polhemus, Quercus arizonica (Fagaceae) , 13 (JTP). Madera Canyon, May 4, 1988, W. A. Jones, Solanum eleagnifolium (Solanaceae) , Nevada: Nye Co.: 2.5 mi NE of Gabbs off Rt. 844, Gabbs Rifle Range, 4800 ft, July 1, 1983, R. T. Schuh and M. D. Schwartz, Lycium andersonii (Solanaceae) , 23, 6♀ (AMNH).

Megalopsallus nigricaput , new species

Figures 6 View Fig , 9 View Fig , 15 View Fig

HOLOTYPE: Male, ARIZONA: Graham County, Stockton Pass, Pinaleno Mts., elev. 52005500 ft., June 1–2, 1983, colls. R. T. Schuh, G. M. Stonedahl, Lycium pallidum Miers (Solanaceae) . Deposited in the American Museum of Natural History.

DIAGNOSIS: Recognized unequivocally by the dark head, first antennal segment, coxae, femora, and thoracic sternum in contrast to otherwise almost white to weakly greenish coloration (figs. 6, 9). Probably most easily confused with rubricornis (fig. 11), but in that species pronotum and scutellum always red to castaneous.

DESCRIPTION: Male: Medium sized, total length 3.35–3.44, length apex clypeus–cuneal fracture 2.23–2.33, width across pronotum 1.02–1.10. COLORATION: Generally white with a faint greenish cast; head includ ing eyes, antennal segment 1, coxae, trochanters, femora, and apex of labium castaneous and strongly contrasting with remain der of body and appendages (fig. 9). SUR FACE AND VESTITURE: Head weakly shining, remainder of dorsum smooth and dull; dorsum clothed with pale recumbent, simple setae intermixed with scattered, silvery, weakly flattened setae. STRUCTURE: Elongate ovoid (fig. 9); labium reaching to middle trochanters; claws long, slender, smoothly curving, pulvilli minute. MALE GENITALIA: Vesica elongate, Sshaped, apex membranous; gonopore subapical, sclerotized; no gonopore sclerite (fig. 15).

Female: Total length 3.11–3.24, length apex clypeus–cuneal fracture 2.19 –2.36, width across pronotum 1.04 –1.06; just slightly more ovoid than male (fig. 9).

ETYMOLOGY: Named for the contrasting dark coloration of the head relative to the remainder of the body; from the Latin niger, black, and caput, head.

HOSTS: Lycium pallidum (Solanaceae) ; Shepherdia argentea (Eleagnaceae) .

DISTRIBUTION: Arizona, Nevada, and Utah.

PARATYPES: USA. — Arizona: Graham Co.: Stockton Pass, Pinaleno Mts, 5200– 5500 ft, June 1, 1983, R. T. Schuh and G. M. Stonedahl, Lycium pallidum (Solanaceae) , 1143, 110♀ (AMNH). Yavapai Co.: 22.7 mi S of Ash Fork on Rt. 89, June 4, 1983, G. M. Stonedahl, Shepherdia argentea (Elaeagnaceae) , 273, 26♀ (AMNH). 5 mi N of Prescott on Rt 89, 1800 m, June 20, 1980, R. T. Schuh, Lycium pallidum (Solanaceae) , 183, 64♀ (AMNH). Nevada: Nye Co.: Atomic Test Site, 4.5 mi S GS500 on Jackass Flats Rd, 3300 ft, June 6, 1983, Schuh, Schwartz, Stonedahl, Lycium pallidum (Solanaceae) , 13 (AMNH). Utah: San Juan Co.: Grand Flat near Collins Canyon, 5600 ft, May 28, 1978, D. A. and J. T. Polhemus, 203, 20♀ (JTP).

Megalopsallus nigrofemoratus (Knight)

Figures 10 View Fig , 15 View Fig

Europiella nigrofemoratus Knight, 1968: 39 View in CoL (n. sp.).

Megalopsallus nigrofemoratus: Schuh et al., 1995: 389 View in CoL (n. comb.).

Europiella grayiae Knight, 1968: 41 View in CoL (n. sp.). NEW SYNONYMY.

Megalopsallus grayiae: Schuh et al., 1995: 389 View in CoL (n. comb.).

Europiella montanae Knight, 1968: 45 (n. sp.). NEW SYNONYMY.

Megalopsallus montanae: Schuh et al., 1995: 389 View in CoL (n. comb.).

DIAGNOSIS: Recognized by frequently dark coloration of most of dorsum and femora, or at least head, pronotum, scutellum, and femora dark and contrasting with pale hemelytra (fig. 10). Most easily confused with humeralis (fig. 10) on basis of size, coloration, and type of sexual dimorphism; readily recognized by structure of male genitalia, vesica in nigrofemoratus being more elongate and slender than that of humeralis , and its association with Atriplex and Grayia , whereas humeralis feeding only on Lycium . Pale specimens separated from the Lycium feeding nicholi by having the cuneus unicolorous with corium.

REDESCRIPTION: Male: Small to medium sized, total length 3.13–3.88, length apex clypeus–cuneal fracture 2.00–2.46, width across pronotum 0.97–1.16. COLORA TION: Dorsum varying from largely deep blackishbrown to having the hemelytra somewhat lighter to much lighter (fig. 10); eyes blackish; antennae usually reddish, sometimes almost entirely pale; femora varying from dark or darkened to pale with a reddish cast and sometimes red spots, tibiae pale, tibial spines dark with dark bases; underside of body dark. SURFACE AND VESTITURE: Dorsum smooth, very weakly shining, clothed with recumbent, dark, simple setae intermixed with woolly, silvery setae. STRUCTURE: Weakly to strongly elongate, parallelsided, specimens in some populations relatively more elongate than those in others; labium reaching posterior margin of middle trochanters; claws relatively short and straight, curving sharply near apex, pulvilli of moderate size, attached to claw only at base of pulvillus. MALE GENITALIA: Vesica elongate, strongly curving, apex membranous, gonopore subapical, gonopore sclerite well developed (fig. 15).

Female: Total length 2.33–2.85, length apex clypeus–cuneal fracture 1.65–2.08, width across pronotum 0.90–1.06; ovoid, often much shorter and more robust than male (fig. 10).

HOSTS: Atriplex canescens , A. confertifolia , A. sp., Grayia spinosa , Sarcobatus baileyi , S. vermiculatus (Chenopodiaceae) . Probable sitting records: Artemisia cana , A. nova , rabbit bush [ Chrysothamnus ] ( Asteraceae ); Condalia globosa , Rhamnus sp. (Rhamnaceae) .

DISTRIBUTION: Interior of western North America from Alberta, Canada, south to Zacatecas, Mexico.

DISCUSSION: Knight (1968) described the species grayiae and montanae on the basis of material from a single locality or area, comprising relatively homogeneous samples with limited host data. After having examined a very large number of specimens from many localities, most with associated host information, I conclude that grayiae and montanae fall within the range of variation of

nigrofemoratus and therefore treat the three nominal species as synonymous.

SPECIMENS EXAMINED: CANADA. — Alberta: Mayberries, July 8, 1952, L. A. Konotopetz, Sarcobatus sp. (Chenopodiaceae) , 83, 10♀ (CNC). Saskatchewan: Wood Mountain, August 8, 1955, A. R. Brooks, 2♀ (CNC). MEXICO. — Baja California Norte: 12 mi E of El Rosario, March 25, 1979, J. D. Pinto, Atriplex sp. (Chenopodiaceae) , 13 (UCR). Nuevo Leon: Santa Ana , Sep

tember 8, 1969, L. A. Kelton, 43, 4♀ (CNC). San Luis Potosi: 19.6 mi S of Huizache, July 25, 1976, Schaffner et al., 43, 4♀ (TAMU). 28.5 mi S of Huizache, July 4, 1985, Jones, Schaffner, 73, 12♀ (TAMU). Sonora: 40 mi W of Moctezuma, April 27, 1981, D. A. and J. T. Polhemus, Rhamnus sp. (Rhamnaceae) , 13, 10♀ (JTP). Zacatecas: 13 mi SW of Concepcion del Oro, July 9, 1983, Kovarik, Harrison, and Schaffner, 173, 14♀ (TAMU). Concepcion del Oro, July 8, 1983, Kovarik, Harrison, Schaffner, 23 (TAMU). 30 mi SW of Concepcion del Oro, July 9, 1983, Kovarik, Harrison, and Schaffner, 353, 21♀ (TAMU). 6 mi S of Concepcion del Oro, July 9, 1983, Kovarik, Harrison, Schaffner, 13, 3♀ (TAMU). USA. — Arizona: Eager, Apache Natl. Forest, 7500 ft, July 12, 1968, L. A. Kelton, ( Asteraceae ), 3♀ (CNC). Coconino Co.: 1 mi E of Tuba City on Rt. 163, 5000 ft, June 16, 1983, R. T. Schuh and M. D. Schwartz, Sarcobatus vermiculatus (Chenopodiaceae) , 13, 6♀ (AMNH). Marble Canyon Monument, T7E R39N, 3603 ft, April 18, 1982, M. D. Schwartz, Atriplex confertifolia (Chenopodiaceae) , 203, 6♀ (AMNH). Red Lake on Rt. 160, June 26, 1980, K. & R. Schmidt, Sarcobatus vermiculatus (Chenopodiaceae) , 3♀ (AMNH). Rt. 64 just SE of Grand Canyon Natl. Park, June 26, 1980, K.&R. Schmidt, Atriplex canescens (Chenopodiaceae) , 1♀ (AMNH). Maricopa Co. : Palo Verde Rd at Rt. 85 just S of Buckeye, 335 m, April 1, 1981, R. T. Schuh and M. D. Schwartz, Atriplex sp. (Chenopodiaceae) , 23 (AMNH). Pima Co. : 10 mi S of Robles jct., April 20, 1982, D. A. and J. T. Polhemus, Atriplex sp. (Chenopodiaceae) , 13 (JTP). Tucson, April 16, 1939, B. P. Bliven, 23 (CAS). Santa Catalina Mts, Finger Rock Canyon Trail , 3000– 3500 ft, April 5, 1981, M. D. Schwartz, Artemisia cana (Asteraceae) , 13, 2♀ (AMNH). Organ Pipe Cactus Natl. Mon., Ajo Valley, April 10, 1981, D. A. Polhemus, Condalia globosa (Rhamnaceae) , 63, 5♀ (JTP). 7 mi SE of Continental on rd to Madeira Canyon, 3700 ft, September 28, 1988, M. D. Schwartz, Atriplex canescens (Chenopodiaceae) , 2♀ (AMNH). Yavapai Co.: Caslte Wash north of Phoenix, March 21, 1980, J. T. Polhemus, 13, 3♀ (JTP). California: Inyo Co. : Mono Lake, Tioga Lodge, June 22, 1929, R. L. Usinger, 2♀ (CAS). Saline Valley, May 25, 1975 – June 5, 1976, D. Guiliani, 33 (LACM). Darwin, May 12, 1969, P. A. Opler, 1♀ (UCB). 9 mi NE of Big Pine, 6300 ft, June 9, 1966, W. Gagne, 13 (UCB). 8 mi NE of Independence, Mazourka Canyon, May 11, 1969, J. T. Doyen, 53, 18♀ (UCB). 7 mi NE of Panamint Springs, May 16, 1969, P. Rude and J. Doyen, 123, 2♀ (UCB). 2 mi E of Westgard Pass Summit, White Mts, 2125 m, July 2, 1980, R. T. Schuh, Atriplex sp. (Chenopodiaceae) , 63, 11♀ (AMNH). Surprise Canyon, Panamint Mts, April 24, 1957, J. Powell, 13 (UCB). San Bernardino Co.: 1.5 mi W of Kramer Junction, April 24, 1980, M. D. Schwartz, Atriplex confertifolia (Chenopodiaceae) , 73, 13♀ (AMNH). 3 mi S of Kramer Junction, April 15, 1965, D. Veirs, 53, 3♀ (UCB). 3 mi W of Lucerne Valley, May 5, 1975, J. D. Pinto, 53, 1♀ (UCR). 6 mi SE of Kramer Junction, April 8, 1966, L. and C. W. O’Brien, 153, 13♀ (UCB). Goldstone Lake, April 26, 1953, R.O Schuster, 2♀ (UCB). Colorado: Eagle Co.: N of Eagle, June 24, 1979, J. T. Polhemus, 2♀ (JTP). Mesa Co. : Rabbit Valley, June 7, 1981, J. T. Polhemus, 13 (JTP). E of Palisade, Plateau Creek, June 6, 1981, J. T. Polhemus, 43 (JTP). Mineral Co.: Creede, June 21, 1990, J. T. and D. A. Polhemus, 63, 10♀ (JTP). Moffat Co.: 5 mi S of Baggs, Wyoming, 6350 ft, August 14, 1986, R. T. Schuh, Sarcobatus vermiculatus (Chenopodiaceae) , 153, 20♀ (AMNH). Gates of Lodore, June 28, 1979, D. A. Polhemus, 23, 3♀ (JTP). Montezuma Co. : 4 mi E of Cortez, August 16, 1973, J. C. Schaffner, 8♀ (TAMU). Cortez, July 19, 1968, L. A. Kelton, Salicornia sp. (Chenopodiaceae) , 63, 14♀ (CNC). Mesa Verde Natl. Park , June 10, 1968, J. E. Slansky, 2♀ (UCD). Montrose Co.: 6 mi E of Montrose, August 13, 1987, T.J. Henry, Atriplex canescens (Chenopodiaceae) , 53, 10♀ (USNM). Pueblo Co.: Pueblo, June 15, 1900, 43 (USNM). Rio Blanco Co. : W Evacuation Creek, 4 mi SE of state line on Rt. 45, 6400 ft, July 9, 1981, M. D. Schwartz, Sarcobatus vermiculatus (Chenopodiaceae) , 33, 1♀ (AMNH). Montana: Carbon Co.: Bear Creek between Red Lodge and Belfry, 5000 ft, August 12, 1986, M. D. Schwartz and G. M. Stonedahl, Sarcobatus vermiculatus (Chenopodiaceae) , 213, 45♀ (AMNH). Gallatin Co.: Bozeman, July 17, 1926, W. Downes, 1♀ (USNM). Yellowstone Co.: Billings, August 4, 1927, H. H. Knight, 3♀ (USNM). Nevada: Clark Co.: 1 mi E of Searchlight, 1095 m, May 17, 1978, R. T. Schuh, 33, 5♀ (AMNH). Elko Co.: 2.2 mi NE of Cobre on state Rt. 233, 5900 ft, June 25, 1983, R. T. Schuh and M. D. Schwartz, Atriplex confertifolia (Chenopodiaceae) , 6♀ (AMNH). 2.2 mi NE of Cobre on state Rt. 233, 5900 ft, June 25, 1983, R. T. Schuh and M. D. Schwartz, Grayia spinosa (Chenopodiaceae) , 183, 19♀ (AMNH). 23.7 mi S of jct. Rt. 229 on Rt. 93, July 19, 1980, G. M. Stonedahl, Atriplex sp. (Chenopodiaceae) , 4♀ (AMNH). Utah state line on Utah Rt. 30, 4760 ft, June 25, 1983, R. T. Schuh and M. D. Schwartz, Grayia spinosa (Chenopodiaceae) , 223, 25♀ (AMNH). Eureka Co.: Garden Summit on Rt. 278, 23.5 mi N of Rt. 50, 6500 ft, June 27, 1983, R. T. Schuh and M. D. Schwartz, Grayia spinosa (Chenopodiaceae) , 73, 18♀ (AMNH). Humboldt Co.: 27 mi W of Denio, June 23, 1971, P. Oman, 13 (OSU). Sheldon Natl. Antelope Range. 0.5 mi S of jct. 8A and 343A, 1950 m, July 2, 1979, R. T. Schuh and B. M. Massie, Atriplex sp. (Chenopodiaceae) , 133, 26♀ (AMNH). Lander Co.: 1.5 mi S of Rt 50 on Rt 376, T18N R45E, 6000 ft, June 28, 1983, R. T. Schuh and M. D. Schwartz, Grayia spinosa (Chenopodiaceae) , 183, 60♀ (AMNH). Lincoln Co.: 5 mi NE of jct. Rts 38 & 93, 2500 ft, May 19, 1982, M. D. Schwartz, Grayia spinosa (Chenopodiaceae) , 133, 27♀ (AMNH). Lyon Co.: Weeks, May 29, 1967, C. W. O’Brien, Sarcobatus vermiculatus (Chenopodiaceae) , 23 (UCB). Mineral Co.: 27 mi SW of Hawthorne on Rt 359, 1 mi NE of Anchorite Summit, 7400 ft, July 2, 1983, R. T. Schuh and M. D. Schwartz, Grayia spinosa (Chenopodiaceae) , 183, 14♀ (AMNH). Nye Co.: Mercury, 18 M, June 11, 1965, H. Knight and J. Merino, Grayia spinosa (Chenopodiaceae) , 173, 13♀ (USNM). Test Site, 6.8 mi SE Mercury Hwy on Orange Blossom Rd, 4000 ft, June 8, 1983, Schuh, Schwartz, and Stonedahl, 13 (AMNH). Mercury, 40M, June 14, 1965, H. Knight and J. Merino, Grayia spinosa (Chenopodiaceae) , 73, 8♀ (USNM). Berlin Ichthyosaur State Mon. on Rt. 844, 6350 ft, July 1, 1983, R. T. Schuh and M. D. Schwartz, Sarcobatus baileyi (Chenopodiaceae) , 1♀ (AMNH). Atomic Test Site, Mercury Hwy at Angle Rd, 3800 ft, June 8, 1983, Schuh, Schwartz, and Stonedahl, Grayia spinosa (Chenopodiaceae) , 63, 5♀ (AMNH). Atomic Test Site, Mercury Hwy at Angle Rd, 3800 ft, June 8, 1983, Schuh, Schwartz, and Stonedahl, Atriplex canescens (Chenopodiaceae) , 63, 2♀ (AMNH). Atomic Test Site, Jackass Flats Road, 3300 ft, June 6, 1983, Schuh, Schwartz, and Stonedahl, 23 (AMNH). 5.5 mi S of Belmont on Rt. 82, 2031 m, July 13, 1980, R. T. Schuh and G. M. Stonedahl, Atriplex sp. (Chenopodiaceae) , 43, 13♀ (AMNH). 35 mi N of Tonapah, Coyote Hole Spring/Sevier Reservoir, T8 R42E S11 & 23, 6000 ft, June 30, 1983, R. T. Schuh and M. D. Schwartz, Sarcobatus vermiculatus var. baileyi (Chenopodiaceae) , 5♀ (AMNH). Northumberland Canyon Rd, Toquima Mts, 6400 ft, June 28, 1983, R. T. Schuh and M. D. Schwartz, Grayia spinosa (Chenopodiaceae) , 23, 2♀ (AMNH). Washoe Co.: Verdi, July 9, 1967, W. Gagne, 1♀ (UCB). New Mexico: Taos Co.: Ojo Caliente, June 6, 1982, D. A. and J. T. Polhemus, Atriplex sp. (Chenopodiaceae) , 13, 5♀ (JTP). Oregon: Harney Co.: Alvord Basin, T37S R33E S26, April 26, 1979, Cobb and Lightfoot, Atriplex spinosa (Chenopodiaceae) , 13, 1♀ (OSU). T37S R33E, Lightfoot and Cobb, Atriplex spinosa (Chenopodiaceae) , 43, 4♀ (OSU). Utah: Duchesne Co.: 23.7 mi S of Myton, Well’s Draw, T10S R15E, 6000 ft, M. D. Schwartz, 1♀ (AMNH). Garfield Co.: Capitol Reef Natl. Park, Grand Wash, Cohab Canyon Trail, 5350–6640 ft, June 21, 1983, R. T. Schuh and M. D. Schwartz, Sarcobatus vermiculatus (Chenopodiaceae) , 563, 33♀ (AMNH). Grand Co.: East of Moab, Utah, Colorado River Bridge, May 26, 1979, J. T. and D. A. Polhemus, 103, 14♀ (JTP). 11 mi SE of Rt 163 toward Dead Horse Point on Rt. 313, 5200 ft, June 11, 1982, M. D. Schwartz, Grayia spinosa (Chenopodiaceae) , 113, 12♀ (AMNH). Millard Co.: Longridge Reservoir Cutoff on Rt. 50/6, 4500 ft, May 19, 1982, M. D. Schwartz, Atriplex confertifolia (Chenopodiaceae) , 203, 14♀ (AMNH). San Juan Co.: 25 mi N of Monticello on Rt. 191, 5700 ft, July 18, 1986, R. T. Schuh, Sarcobatus vermiculatus (Chenopodiaceae) , 343, 63♀ (AMNH). Sevier Co.: 32 mi N of Hiway 24 on Hiway 72, 7560 ft, July 17, 1980, G. M. Stonedahl, Atriplex sp. (Chenopodiaceae) , 1♀ (AMNH). Uintah Co.: 5–10 mi SW of Bonanza, T10S R24E, 5000–5600 ft, June 6, 1981 – June 8, 1982, M. D. Schwartz, Sarcobatus vermiculatus (Chenopodiaceae) , 823, 92♀ (AMNH). Wyoming: Big Horn Co.: 1 mi W of Shell Creek Falls Scenic Overlook on Rt. 14, 5600 ft, August 13, 1986, Schuh, Schwartz, and