Mecynostomella hollowayi, Johanson, 2003

Johanson, Kjell Arne, 2003, Revision of the New Caledonian genus Mecynostomella (Trichoptera, Kokiriidae), Zootaxa 270 (1), pp. 1-24 : 15-17

Mecynostomella hollowayi sp.n.

( Figs 6­11 , 36­40 , 53 )

Material examined: Male holotype: Plateau N Col d'Amos, 13.vii. 1971, 440 m (#46) [J.D. Holloway] ( ROM, alcohol). 2 male paratypes: N Mt. Aoupinié, 520 m, Haut Mou, Napoue Areu valley , 30­31.vii.1971 (#62) [J.D.Holloway] ( NRM, alcohol) . 3 male paratypes: same, except ( ROM, alcohol) . 1 male paratype: Province Sud, stream at Koghi Mt. , 22°10.722'S, 166°30.501'E, 462 m, light trap, 6.xii.2001 [Johanson, Pape, Viklund] ( NRM, alcohol) .

Etymology: hollowayi , named after J. D. Holloway who collected the species. To be treated as a noun in genitive case.

Diagnosis: The species has a pale area present between the forewing Cu and A, as in sigma . The two species are separated by the longer and slightly ventrally curving lateral branch, and shorter median branch in hollowayi .

Male.

Colors faded. Antennal flagellum with 42 flagellomeres.

Thorax: Pronotum with stout, dark brownish setae not confined to distinct warts.

Wings ( Figs 6­11 ): Forewing length 8.5­10.2 mm (n=5), hind wing length 7.3­9.3 mm (n=5). Forewing pale brownish, with distinct pale area between A and Cu (a in Fig. 6 ) formed by densely arranged modified sensillae ( Fig. 11 ). Hind wing pale grey. Venation as described for genus.

Genitalia ( Figs 36­40 ): Segment IX, lateral view ( Fig. 36 ), about 2.1x higher than long; anterior part shallowly produced, with rounded, undulating margin; dorsal margin narrowly rounded; in ventral view ( Fig. 38 ), ventral margin with wide hyperbolar, incision. Segment X simple; in dorsal view ( Fig. 37 ), central branch about parallel­sided at distal half; in lateral view ( Fig. 36 ), proximally nearly parallel­sided, tapers distally, dorsal margin slightly bent ventrally at mid­length; lateral margin with few setae on distal half, about 2x longer than high ( Fig. 36 ), apex narrowly rounded in lateral view, slightly incised in dorsal view ( Fig. 37 ). Lateral branch nearly as long as central branch, tubular, slightly curving ventrally along ( Fig. 36 ), slightly converging in dorsal view ( Fig. 37 ); short apical setae present, proximal setae grouped with setae on segment IX. Median branches large, hidden in dorsal view ( Fig. 37 ), with short setae on ventral margin ( Fig. 36 ). Gonocoxite, lateral view ( Fig. 36 ), generally sub­straight, except apex produced dorsad; dorsal margin sigmoid, ventral margin sub­straight ( Fig. 36 ); incised sub­apical part about 0.7x as high as proximal part; sub­straight in ventral view ( Fig. 38 ), oriented posteriad, covered by setae except at proximal part. Phallus simple; phallobasis about three­quarts the height of segment IX ( Fig. 39 ); endotheca produced dorsally from dorsal opening starting midlength on phallus, without spines; in ventral view ( Fig. 40 ), lateral incisions at proximal part located close to posterior margin of anterior opening.

Female unknown.

Figures

FIGURES 6­14. Mecynostomella species, right wings and wing details. 6, Mecynostomella spinosa sp.n. male, right wings and vein terminology; 7, Mecynostomella spinosa sp.n. male, scaloid seta from forewing Cu2, highly magnified; 8, Mecynostomella spinosa sp.n. female, right wings; 9, Mecynostomella brevis sp.n. male, right wings; 10, Mecynostomella hollowayi sp.n. male, right wings, a indicates the position of a pale area in the Cu­ and A­ area of the forewing, and made from minute structures shown in the next figure; 11, details of a single structure forming a pentagon and with a central opening, and which build up a pale area of the forewing as indicated in the previous figure; 12, Mecynostomella fusca Kimmins, 1953, right wings, with area covered by sensillae indicated by a surrounding black line; 13, Mecynostomella flinti sp.n., detail from wing membrane of dark area between forewing Cu and A, showing position of a modified sensilla and sensilla bases (t), dorsal view; 14, same species and sensilla, lateral view.

FIGURES 36­40. Mecynostomella hollowayi sp.n., male holotype. 36, genitalia, lateral view; 37, genitalia, dorsal view; 38, genitalia, ventral view; 39, phallus, lateral view; 40, phallus, ventral view.

FIGURES 53­54. Distribution maps of New Caledonian Mecynostomella. 53, caledonia (circles), hollowayi (rectangles), and sigma (triangle); 54, spinosa (circles), flinti (rectangle), brevis (triangles), and fusca (diamond).