Manota index, Hippa, Heikki, 2008

Hippa, Heikki, 2008, New species and new records of Manota Williston (Diptera, Mycetophilidae) from the Oriental region, Zootaxa 1723, pp. 1-41 : 21

Manota index sp. n.

( Figs 12 A, B, C)

Male. Colour. Head pale brown, frons and occiput darker brown, mouthparts pale yellowish. Thorax pale yellowish-brown, posterior part of scutum and scutellum darker brown, preepisternum 2 ventrally paler yellowish. Legs pale yellowish-brown, trochanter 3 and femur 3 at base infuscated. Wing unicolorous yellowishbrown; haltere yellowish-brown with dark brown knob. Abdomen brown, as dark as scutellum. All the setosity pale, yellowish or brownish, the thicker setae seeming darker than the finer setae and trichia. Head. Antennal flagellomere 4, Fig. 12 A. Palpomere 3 of maxillary palpus with apicomesial extension, with 4 apically expanded curved sensilla; palpomere 4 with parasegment. Number of postocular setae 12. Thorax. Anepisternum setose, with 41 setae, anterior basalare non-setose, preepisternum 2 setose, with 15 setae, laterotergite setose, with 20 setae, episternum 3 setose, with 12 setae. Wing. Length 1.9 mm. Hypopygium, Figs 12 B and C: Sternite 9 about one-half of the ventral length of gonocoxa, lateral margin sharply delimited, posterior margin with a shallow notch, anterior margin deeply incised, the setae similar to the ventral setae of gonocoxa. Ventral mesial margin of gonocoxa simple. Parastylar lobe large, the mesial margin angular and with several setae. Paraapodemal lobe absent. Dorsal mesial margin of gonocoxa simple. Two juxtagonostylar setae present: they are simple strong setae rather than megasetae and arise from a long basal body which is about one-half of the length of the setae. Gonostylus bilobate, with a narrower lateral and a broader mesial lobe; both lobes with two apical megasetae, otherwise the setosity absent except for a few rather short setae ventrally on the lateral lobe, dorsally across base of mesial lobe and at mesial margin of the latter with those on dorsal side being stronger than those on ventral side. Tegmen narrow, with a long parallel-sided apical part and distinct lateral shoulders. In the hypopygium mount of the single specimen, the median parts, tegmen, gonocoxal apodemes and hypoproct, are probably displaced posteriad. Hypoproct setose only posteriorly where there are a few long setae. Cerci medially united.

Female unknown.

Discussion. M. index is not similar to any described Manota . It is readily distinguished by its characteristic bilobed gonostylus with the two megasetae at the apex of each lobe. There are other species with bilobed gonostylus, e.g. the Oriental M. biloba Hippa and M. flavipes (Enderlein) from the Seychelles, but they lack the megasetae and have sternite 9 laterally fused with the gonocoxa.

Types. Holotype. Male, MALAYSIA, Pahang, Bukit Fraser, 4020 ft, Malaise in forest 14–17.III.1997, H. Hippa, M. Jaschhof, B. Viklund (in SMNH).

Figures

FIGURE 12. Manota index sp. n. (holotype). A. Antennal flagellomere 4, lateral view. B. Hypopygium, dorsal view. C. Hypopygium, ventral view. Scale 0.10 mm.