Macrophysoderes cirripilosa, Hwang & Weirauch, 2017

Hwang, Wei Song & Weirauch, Christiane, 2017, Uncovering hidden diversity: phylogeny and taxonomy of Physoderinae (Reduviidae, Heteroptera), with emphasis on Physoderes Westwood in the Oriental and Australasian regions, European Journal of Taxonomy 341, pp. 1-118 : 41-46

Macrophysoderes cirripilosa sp. nov.

urn:lsid:zoobank.org:act:9B61EDE2-4732-4BCB-92B7-8C200A34A7BE

Figs 8–12 ; Appendix

Diagnosis

This species is recognized among species of Macrophysoderes gen. nov. by the short conical head that is covered with semi-erect, curved setae, with a distinct concentration of setae on the postocular lobe between the ocelli, the length of the scape reaching the apex of the clypeus, the very large eye that attains the ventral margin of the head in lateral view, the median pronotal depression not contiguous with the transverse sulcus and the short scutellar process. It most closely resembles M. finisterre gen. et sp. nov. and M. elongata gen. et sp. nov. It is differentiated from the former by the overall coloration, the shape of the antero-lateral pronotal projections, the wider anterior pronotal lobe and the shape of the abdominal apical margin. It is differentiated from M. elongata gen. et sp. nov. by the size, the shape of the scutellar process, the relative length of head to body and the less swollen fore femur.

Etymology

The name combines the Latin noun “cirrus”, meaning curl or hair-tuft, with the Latin adjective “pilosus”, meaning hairy, to describe the abundant, curved setae on the head, especially between the ocelli.

Material examined

Holotype

PAPUA NEW GUINEA: ♂, Western Province , Fly River, Kiunga, 6.11944° S, 141.29194° E, 8–10 Aug. 1957, W.W. Brandt leg. ( UCR_ENT 00073448 ) ( BPBM).

Paratypes

PAPUA NEW GUINEA: 4 ♂♂, same collection data as for holotype ( UCR _ENT 00073446, UCR _ENT 00073447, UCR_ENT 00073453, UCR _ENT 00073454) ( BPBM).

Other material

INDONESIA: Irian Jaya: 1 ♀, Cyclops Mts, Ifar, 2.6° S, 140.61° E, 300 m, 23–25 Jul. 1962, J. Sedlacek leg. ( UCR _ENT 00073499) ( BPBM); 1 ♀, Cyclops Mts, 2.8775° S, 140.70333° E, 1067 m, Mar. 1936, L.E. Cheesman leg. ( UCR _ENT 00069271) ( BMNH); 1 ♂, Cyclops Mts, Mt Lina, 2.43055° S, 140.45333° E, 1067 m, Mar. 1936, L.E. Cheesman leg. ( UCR _ENT 00069263) ( BMNH); 1 ♂, Guega, W of Swart Valley, 3.6° S, 138.41666° E, 1200 m, 15 Nov. 1958, J.L. Gressitt leg. ( UCR _ENT 00073468) ( BPBM); 1 ♀, Ifar, Cyclops Mts, 2.83166° S, 140.60555° E, 450 m, 8 Sep. 1962, J. Sedlacek leg. ( UCR _ ENT 00073500) ( BPBM); 1 ♂, Yapen, 1.79472° S, 136.30361° E, 600 m, 4 Jan. 2007, S. Bily leg. ( UCR _ENT 00073850) ( NMPC). – Papua: 1 ♂, Paniai Division Co., Wisselmeren Enarotadi, 4.205° S, 136.59305° E, 1900 m, 16 Jul. 1962, J. Sedlacek leg. ( UCR _ENT 00073466) ( BPBM); 1 ♀, Araboebivak, 2.61527° S, 140.62805° E, 6 Oct. 1939, collector unknown ( UCR _ENT 00024031) ( RMNH); 1 ♀, Araucaria Camp, 5.65916° S, 139.13583° E, 800 m, Mar. 1939, L.J. Toxopeus leg. ( UCR _ENT 00024011) ( RMNH); 1 ♀, Bernhard Camp, 6.21305° S, 141.54805° E, 50 m, 5 Oct. 1938, J. Olthof leg. ( UCR _ENT 00014057) ( RMNH); 2 ♀♀, same collection data as preceding but Aug. 1938 ( UCR _ENT 00024007, UCR _ENT 00024008) ( RMNH); 2 ♀♀, same collection data as preceding but Jul.–Nov. 1938 ( UCR _ENT 00024009, UCR _ENT 00024010) ( RMNH); 1 ♂, Hollandia (Jayapura), 2.61527° S, 140.62805° E, 500 m, Jul. 1938, L.J. Toxopeus leg. ( UCR _ENT 00024005) ( RMNH); 1 ♂, Nabire, S Geelwink Bay, 3.36667° S, 135.48333° E, 2–9 Jul. 1962, J.L. Gressitt leg. ( UCR _ENT 00073469) ( BPBM); 1 ♂, Sibil, Star Mountain Range, 5.2375° S, 141.14861° E, 1260 m, 18 Apr. 1959, collector unknown( UCR _ENT 00024028)( RMNH); 1♂,same collection data as preceding but 21Apr.1959 ( UCR _ ENT 00024030)( RMNH); 1♂, same collection data as preceding but 25Apr. 1959 ( UCR _ENT 00024029) ( RMNH); 1 ♂, same collection data as preceding but 25 May 1959 ( UCR _ENT 00024006) ( RMNH); 1 ♀, Wamena, 4.10055° S, 138.90472° E, 700 m, 10–25 Feb. 1960, T.C. Maa leg. ( UCR _ENT 00073485) ( BPBM), 3 ♂♂, same collection data as preceding ( UCR _ENT 00073530– UCR _ENT 00073532) ( BPBM).

PAPUA NEW GUINEA: Bougainville Province: 1 ♂, Mt Nomo, S of Mt Bougainville, 6.05361° S, 155.19055° E, 213 m, Feb. 1936, L.E. Cheesman leg. ( UCR _ENT 00069295) ( BMNH). – Central Province: 1 ♂, Daradae Plain, 80 km N of Port Moresby, 8.73665° S, 146.8088° E, 580 m, 6 Sep 1959, T.C. Maa leg. ( UCR _ENT 00073480) ( BPBM); 1 ♂, Iriri, near Kerema, 8.98416° S, 146.98944° E, 7 May 1959, C.D. Michener leg. ( UCR _ENT 00073484) ( BPBM); 1 ♀, Murua River near Kerema, 8.08361° S, 145.91138° E, 6 May 1959, C.D. Michener leg. ( UCR _ENT 00073492) ( BPBM); 1 ♂, Otomata Plantation, 1 mi. E of Moresby, 9.48166° S, 147.10361° E, 2 Nov. 1960, J.L. Gressitt leg. ( UCR _ENT 00073481) ( BPBM); 1 ♂, Owen Stanley Range, Goilala, Bome, 9.57666° S, 146.37194° E, 1950 m, 8–15 Mar. 1958, W.W. Brandt leg. ( UCR _ENT 00073483) ( BPBM); 1 ♀, Owen Stanley Range, Goilala, Loloipa, 9.23583° S, 147.98444° E, 1–15 Feb. 1958, W.W. Brandt leg. ( UCR _ENT 00073498) ( BPBM). – East New Britain Province: 1 ♀, near Rabaul, 4.175° S, 152.24805° E, Feb. 1929, collector unknown ( UCR _ENT 00046667) ( CAS). – East Sepik Province: 1 ♀, Amboin Patrol Post, Karawari Lodge, 4.40444° S, 142.98555° E, Feb. 1983, A.C. Messer leg. ( UCR _ENT 00031394) ( USNM); 1 ♀, Mar.–Apr. 1983, A.C. Messer leg. ( UCR _ENT 00031395) ( USNM); 1 ♀, May River, 4.40694° S, 141.83944° E, 6 Jun. 1963, R. Straatman leg. ( UCR _ENT 00073503) ( BPBM). – Madang Province: 1 ♂, Erima, Astrolabe Bay, 5.4225° S, 145.73361° E, 1897, Biro leg. ( UCR _ENT 00069770) ( HNHM); 1 ♀, same collection data as preceding ( UCR _ENT 00069777) ( HNHM); 1 ♂, same collection data as preceding but 1896 ( UCR _ENT 00069767) ( HNHM); 5 ♀♀, same collection data as preceding ( UCR _ ENT 00069772– UCR _ENT 00069776) ( HNHM); 1 ♂, Mondo, 5.41666° S, 144.76138° E, 1524 m, Feb. 1934, L.E. Cheesman leg. ( UCR _ENT 00069261) ( BMNH); 1 ♂, Stephansort, Astrolabe Bay, 5.43638° S, 145.74138° E, 1897, Biro leg. ( UCR _ENT 00069771) ( HNHM). – Morobe Province: 2 ♂♂, Huon Penninsula Co., Finschhafen, 6.55527° S, 147.17361° E, 20 Apr. 1944, E.S. Ross leg. ( UCR _ENT 00046650, UCR _ENT 00046651) ( CAS); 1 ♀, same collection data as preceding ( UCR _ENT 00046654) ( CAS); 1 ♀, same locality as preceding, May 1944, F. Skinner leg. ( UCR _ENT 00046655) ( CAS); 1 ♂, same locality as preceding, 7 May 1944, E.S. Ross leg. ( UCR _ENT 00046652) ( CAS); 1 ♂, same locality as preceding, May 1944, E.S. Ross leg. ( UCR _ENT 00046653) ( CAS); 1 ♀, 15 Nov. 1969, J.E. Tobler leg. ( UCR _ENT 00046673) ( CAS); 1 ♀, Huon Gulf, Morobe District, 6.55861° S, 147.50805° E, 22 May–19 Jun. 1937, J.L. Froggatt leg. ( UCR _ENT 00069312) ( BMNH); 1 ♂, Sattelberg, Huon-Golf, 6.485° S, 147.75861° E, 1899, Biro leg. ( UCR _ENT 00069788) ( HNHM); 1 ♂, same collection data as preceding but 1898 ( UCR _ENT 00069787) ( HNHM); 1 ♂, Bulolo, 7.20472° S, 146.63166° E, 900 m, 27 Mar. 1968, P. Colman leg. ( UCR _ENT 00073467) ( BPBM); 1 ♀, same collection data as preceding ( UCR _ENT 00073501) ( BPBM); 1 ♀, Bumayong, 6.63444° S, 147.0025° E, Jul. 1957, R.W. Paine leg. ( UCR _ENT 00069302) ( BMNH); 1 ♀, Garaina, 7.88333° S, 147.13333° E, 800 m, 4 Jan. 1968, J. and M. Sedlacek leg. ( UCR _ENT 00073496) ( BPBM); 3 ♂♂, same collection data as preceding but 15 Jan. 1968 ( UCR _ENT 00073474– UCR _ENT 00073476) ( BPBM); 1 ♂, same collection data as preceding but 13–15 Jan. 1968 ( UCR _ENT 00073477) ( BPBM); 1 ♂, Gewak, Salawaket Range (Saruwaged Range), 6.21694° S, 146.75° E, 1530 m, 6 Sep. 1956, E.J. Ford Jr leg. ( UCR _ENT 00073464) ( BPBM); 1 ♂, NE Wau, 7.33805° S, 146.71555° E, 1270 m, 7 May 1962, J. Sedlacek leg. ( UCR _ENT 00073612) ( BPBM); 1 ♂, same collection data as preceding but 2–4 Jan. 1963 ( UCR _ENT 00073607) ( BPBM); 1 ♀, NE Wau, 7.33166° S, 146.71805° E, 1010 m, 18 Dec. 1968, J. Sedlacek leg. ( UCR _ENT 00073620) ( BPBM); 1 ♂, NE Wau, 7.32138° S, 146.71583° E, 1050 m, 5 Jan. 1963, G. Monteith leg. ( UCR _ENT 00073611) ( BPBM); 2 ♂♂, same collection data as preceding but 7 Jan. 1963, J. Sedlacek leg. ( UCR _ENT 00073609, UCR _ENT 00073610) ( BPBM); 1 ♀, same collection data as preceding ( UCR _ENT 00073613) ( BPBM); 1 ♂, NE Wau, Hospital Creek, 7.33805° S, 146.71555° E, 1300 m, 5 Dec. 1965, J. Sedlacek leg. ( UCR _ENT 00073606) ( BPBM); 1 ♂, NE Wau, Mt Missim, 7.1167° S, 146.9167° E, 950–1300 m, Mar. 1965, J. and M. Sedlacek leg. ( UCR _ ENT 00073478) ( BPBM); 2 ♂♂, Waing, ca 18 mi. (?) of Lae, 6.72444° S, 146.96805° E, 13–14 Apr. 1965, Balogh and Szent-Ivany leg. ( UCR _ENT 00069768, UCR _ENT 00069769); 1 ♀, same collection data as preceding ( UCR _ENT 00069778) ( HNHM); 1 ♀, Wau, 7.33333° S, 146.71667° E, Jan.–Mar. 1982, R.T. Bell leg. ( UCR _ENT 00073618) ( BPBM); 2 ♀♀, same collection data as preceding but Feb– May 1982 ( UCR _ENT 00073616, UCR _ENT 00073617) ( BPBM); 1 ♂, Wau, 7.32138° S, 146.71555° E, 1097 m, 11 Sep. 1971, W. Gagne leg. ( UCR _ENT 00046657) ( CAS); 1 ♀, Wau, 7.3333° S, 146.71667° E, 1200 m, 27 Jul. 1961, J. and J.H. Sedlacek leg. ( UCR _ENT 00073614) ( BPBM); 1 ♂, same collection data as preceding but 23 Dec. 1961, G. Monteith leg. ( UCR _ENT 00073603) ( BPBM); 1 ♂, same collection data as preceding but 31 Aug. 1961, J. Sedlacek leg. ( UCR _ ENT 00073602) ( BPBM); 1 ♂, same collection data as preceding but 18–25 Jun. 1962 ( UCR _ENT 00073599) ( BPBM); 1 ♂, same collection data as preceding but 14–24 Mar. 1964 ( UCR _ENT 00073601) ( BPBM); 1 ♂, same collection data as preceding but 17–19 Sep. 1964 ( UCR _ENT 00073600) ( BPBM); 1 ♂, 8–14 Dec. 1976, G.F. Hevel and R.E. Dietz IV leg. ( UCR _ENT 00031396) ( USNM); 1 ♂, Wau, 7.3333° S, 146.71667° E, 1300 m, 24 Nov. 1963, J.L. Gressitt leg. ( UCR _ENT 00073605) ( BPBM); 1 ♂, Wau, 7.3333° S, 146.71667° E, 1100 m, 4 Oct. 1962, J. Sedlacek leg. ( UCR _ENT 00073604) ( BPBM); 1 ♂, Wau, Hospital Creek, 7.33805° S, 146.71555° E, 1250 m, 10 Apr. 196X [exact year unknown], J. and M. Sedlacek leg. ( UCR _ENT 00073608) ( BPBM); 1 ♀, Wau Ecological Institute, 7.33805° S, 146.71555° E, 4 Dec. 1988, R. Holyuski leg. ( UCR _ENT 00069789) ( HNHM). – Northern Province: 1 ♂, SE Popondetta, 8.76666° S, 148.23333° E, 25 m, May 1966, P. Shanahan leg. ( UCR _ ENT 00073465) ( BPBM). – Oro Province: 1 ♀, Kokoda, 8.86083° S, 147.73722° E, 350 m, 21 Mar. 1956, J.L. Gressitt leg. ( UCR _ENT 00073497) ( BPBM); 2 ♀♀, Kokoda, 8.87777° S, 147.7375° E, 366 m, Apr. 1933, L.E. Cheesman leg. ( UCR _ENT 00069264, UCR _ENT 00069265) ( BMNH); 2 ♀♀, Kokoda, 8.87722° S, 147.7375° E, 366 m, Jun. 1933, L.E. Cheesman leg. ( UCR _ENT 00069268, UCR _ ENT 00069269) ( BMNH); 3 ♂♂, same collection data as preceding but Sep. 1933 ( UCR _ENT 00069258- UCR _ENT 00069260) ( BMNH); 2 ♀♀, same collection data as preceding ( UCR _ENT 00069266, UCR _ENT 00069267) ( BMNH); 3 ♂♂, same collection data as preceding but Apr. 1933 ( UCR _ENT 00069255– UCR _ENT 00069257) ( BMNH); 1 ♂, Kokoda-Pitoki, 8.8775° S, 147.73722° E, 400 m, 24 Mar. 1956, J.L. Gressitt leg. ( UCR _ENT 00073461) ( BPBM); 2 ♂♂, same collection data as preceding but 23 Mar. 1956 ( UCR _ENT 00073459, UCR _ENT 00073460) ( BPBM); 1 ♀, same collection data as preceding ( UCR _ENT 00073491) ( BPBM); 1 ♂, NE Kokoda, 8.86861° S, 147.74777° E, 400 m, 17–18 Nov. 1965, J. and M. Sedlacek leg. ( UCR _ENT 00073462) ( BPBM); 1 ♂, same collection data as preceding but 19 Nov. 1965 ( UCR _ENT 00073463) ( BPBM). – Sandaun (West Sepik) Province: 1 ♂, Waris, S of Hollandia, 3.26667° S, 141.05° E, 450–500 m, 16–23 Aug. 1959, T.C. Maa leg. ( UCR _ENT 00073470) ( BPBM). – Simbu Province: 1 ♀, Karimui, 6.49611° S, 144.82277° E, 1080 m, 13 Jul. 1963, J. Sedlacek leg. ( UCR _ENT 00073504) ( BPBM). – West New Britain Province: 1 ♀, Silanga, Nakanai Mts, 5.5525° S, 150.87° E, 150 m, 31 Jul. 1956, E.J. Ford Jr leg. ( UCR _ENT 00073507) ( BPBM). – Western Highlands Province: 1 ♂, Wum, Upper Jimmi Valley, 5.92972° S, 144.26805° E, 840 m, 18 Jul. 1955, J.L. Gressitt leg. ( UCR _ENT 00073482) ( BPBM). – Western Province: 1 ♂, Eliptamin Valley, 5.17638° S, 141.54361° E, 1200–1350 m, 16–30 Aug. 1959, W.W. Brandt leg. ( UCR _ENT 00073479) ( BPBM); 1 ♀, same collection data as preceding but 1–15 Sep. 1959 ( UCR _ENT 00073502) ( BPBM); 1 ♂, Fly River, Kiunga, 6.11944° S, 141.29194° E, 11–14 Jul. 1957, W.W. Brandt leg. ( UCR _ENT 00073458) ( BPBM); 2 ♀♀, same collection data as preceding ( UCR _ ENT 00073493, UCR _ENT 00073494) ( BPBM); 1 ♂, same collection data as preceding but 23–25 Jul. 1957 ( UCR _ENT 00073457) ( BPBM); 1 ♀, same collection data as preceding ( UCR _ENT 00073495) ( BPBM); 1 ♂, same collection data as preceding but 5–7 Aug. 1957 ( UCR _ENT 00073456) ( BPBM); 5 ♂♂, same collection data as preceding but 8–10 Aug. 1957 ( UCR _ENT 00073449– UCR _ENT 00073452, UCR _ENT 00073455); 5 ♀♀, same collection data as preceding ( UCR _ENT 00073486– UCR _ENT 00073490) ( BPBM); 2 ♂♂, Star Mts, Sibil Valley, 5.04823° S, 140.97958° E, 1245 m, 18 Oct.–8 Nov. 1961, S. and L. Quate leg. ( UCR _ENT 00073472, UCR _ENT 00073473) ( BPBM); 1 ♀, Koitaki, 8.92527° S, 147.73861° E, 457 m, Nov.–Dec. 1928, collector unknown ( UCR _ENT 00046656) ( CAS); 1 ♂, Maffin Bay, Dutch New Guinea, 2.09222° S, 139.01472° E, Sep. 1944, E.S. Ross leg. ( UCR _ENT 00046658) ( CAS); 1 ♂, Peria Creek, Kwagira River, 9.63111° S, 149.38555° E, 50 m, 14 Aug.–6 Sep. 1953, G.M. Tate leg. ( UCR _ENT 00068948) ( AMNH); 2 ♀♀, Kalalo, 6.76° S, 147.91027° E, 750 m, 20–30 Aug. 1966, G.A. Samuelson leg. ( UCR _ENT 00073508, UCR _ENT 00073509) ( BPBM); 1 ♂, Mafulu, 8.45388° S, 146.7425° E, 1219 m, Jan. 1934, L.E. Cheesman leg. ( UCR _ENT 00069262) ( BMNH). – 1 ♂, unknown locality, no date provided, Saunders leg. ( UCR _ENT 00069270) ( BMNH).

Description

Male

BODY LENGTH. Medium size, total length 8.7 mm, SD ± 0.66 (Appendix).

COLORATION ( Figs 10–11 ). Straw-colored and brown. Head brown. Scape and pedicel of antenna strawcolored, sometimes with brown suffusion, basiflagellomere brown, distiflagellomere straw-colored with brown base. First segment of labium brown with straw-colored apex, second segment basally and apically brown, medially straw-colored, third segment brown. Anterior lobe of pronotum brown with straw-colored markings, posterior lobe straw-colored to light brown, parascutellar lobe with lighter margin, scutellum basally dark brown, apically including process straw-colored, pleuron mixture of straw-color to dark brown, with straw-colored margin of acetabula, sternum dark brown. Corium of hemelytron light brown to brown, membrane brown. Femora straw-colored with basal and apical brown annulations, tibiae straw-colored with basal, medial and apical brown annulations, tarsi and claws strawcolored. Abdomen dorsally dull yellow, ventrally brown medially, straw-colored laterally, with dark brown sub-lateral markings, anterior half of connexivum dark brown, posterior half straw-colored, exposed part of pygophore dark brown.

VESTITURE. Densely setose. Head with widespread curved, setigerous tubercles, ventral surface of postocular lobe with two rows of small, setigerous tubercles, with pair of long, straight setae on postocular lobe posterior to ocelli. Anterior lobe of thorax with tuberculated, short, curved setae on lateral margins and along dorsal ridges, posterior lobe with short, curved setae on humeral angle and sparsely distributed along dorsal surface. Corium of hemelytron with short, curved setae. Legs with two rows of spines and setigerous tubercles, tibiae with regular rows of tuberculated, stout, sharp setae. Connexival margin of abdomen with a few clubbed setae on each segment.

HEAD. Short conical; maxillary plate rounded apically; scape reaching apex of clypeus; eye distinctly projecting in dorsal view, about 1/5 length of head, attaining ventral margin of head in lateral view; height of anteocular lobe shorter than postocular lobe.

THORAX. Antero-lateral paired projections acute, diverging; surface of anterior lobe with low ridges; median pronotal depression not contiguous with transverse sulcus; paramedian carina strongly defined; posterior lobe medially weakly rugose; anterior pronotal lobe shorter than posterior lobe, slightly narrower than posterior lobe, anterior lobe lower than posterior lobe in lateral view; parascutellar lobe bell-shaped and skewed towards median; scutellar process short with rounded apex.

HEMELYTRON. Not attaining tip of abdomen.

LEGS. Same as genus description.

ABDOMEN. Elongate ovoid, with rounded terminal margin; connexival margin slightly undulating.

GENITALIA. Anterior margin of mediosternite 8 sharply emarginated, with medial apodeme; transverse bridge of pygophore narrow, margin of anterior opening rounded; cup-like sclerite apically rounded and rim ventrally sclerotized; arms of basal plate converging; ductifer with sclerotized rounded ring; endosomal struts conical, with subacute apex, basally divided into two plates; dorsal phallothecal sclerite subacute, with lateral, broad, plate-like prolongations, short triangular.

Female

Similar in size and shape to males except with rounder or wider abdomen.

Ecology

One specimen was recorded as having been found in logs and another as caught in a light trap.

Distribution

Found throughout the island of New Guinea, on the nearby island New Britain, and on Bougainville Island (Solomon Archipelago).

Remarks

This species is described based on the examination of 147 specimens. It is the most commonly collected physoderine from the island of New Guinea. There are slight differences among specimens collected from different regions, most apparently variation of size and color. These differences include overlapping variation and are therefore not indicative of multiple species. Genitalic dissections of specimens from different regions showed no differences. Based on the phylogenetic analysis above, the species is placed in the newly created Macrophysoderes gen. nov. Two other new species, M. grandis gen. et sp. nov. and M. elongata gen. et sp. nov., share several characters with M. cirripilosa gen. et sp. nov., including the setose head, the very large eyes, the shape of the parascutellar lobe and the anterior pronotal lobe being almost as wide as the posterior lobe.

Figures

Fig. 8. Dorsal view of the male pygophore in species of Breviphysoderes gen. nov., Macrophysoderes gen. nov., Paraphysoderes Villiers, 1962 and Physoderes Westwood, 1845. For an explanation of the colored arrows, see keys on pages 37 and 66.

Fig. 9. Dorsal view of the phallus in species of Breviphysoderes gen. nov., Macrophysoderes gen. nov., Paraphysoderes Villiers, 1962 and Physoderes Westwood, 1845 (partial). Scale bars = 0.20 mm.

Fig. 10. Dorsal habitus images for species of Macrophysoderes gen. nov. and Nanophysoderes gen. nov. For an explanation of the colored arrows, see key on page 37. Scale bars = 2 mm.

Fig. 11. Ventral habitus images for species of Macrophysoderes gen. nov. and Nanophysoderes gen. nov. Scale bars = 2 mm.

Fig. 12. Distribution map for species of Macrophysoderes gen. nov.