Loomisiella pylei, Shear & Richart & Wong, 2020

Shear, William A., Richart, Casey H. & Wong, Victoria L., 2020, The millipede family Conotylidae in northwestern North America, with a complete bibliography of the family (Diplopoda, Chordeumatida, Heterochordeumatidea, Conotyloidea), Zootaxa 4753 (1), pp. 1-78 : 69

Loomisiella pylei , new species

Figs. 244–246

Types: Male holotype, male and female paratypes from WASHINGTON: Wahakiakum Co., Lower Hendrickson Canyon , 46.3693°N, - 123.6657°W, elev. 30 m, old-growth forest, collected 23 January 2004, by W. Leonard, M. Leonard, C. Richart, R. M. Pyle, and K. Novoselić .

Diagnosis: See under preceding species.

Etymology: The species name honors Robert Michael Pyle, a notable lepidopterist, author, conservation advocate, and as is this species, denizen of the Willapa Hills.

Description: Male: Length, 5.0 mm. Fourteen ocelli in oval-triangular group. Metazonital shoulders poorly developed; segmental setae curved, acute. Legpairs one, two reduced, pairs three, four encrassate, pairs five to seven near same size as postgonopodal legs; femora three, four with basal femoral knobs acute, posteriorly directed.Anterior gonopods (ag, Figs. 244 , 245 ) with large anterior sternum, widely separated, curving lateral to posterior gonopod coxites, apically decurved but without any apical expansion, small lateral branch tightly appressed to gonopod (free in Fig. 245 due to compression on microscope slide). Posterior gonopod telopodites reduced but relatively larger than in L. evergreen ; posterior gonopod coxites (pgc, Figs. 244 , 245 ) short, broad, anteriorly cupped, with broad, lamellate process mesally, basal pseudoflagellum originating posteriorly, two processes distolaterally, more lateral process composed of filaments (seen as separate in Fig. 246 due to compression on microscope slide), mesal process curved, apically fimbriate. Coxae 10, prefemora 11 as usual.

Female 5.2 mm long, nonsexual characters as in male.

Distribution: WASHINGTON: Lewis Co., FS-25 8.5 mi S of Randle, 46.4409°N, - 121.9966°W, elev. 330 m, 21 December 2003, W. Leonard, C. Richart m f; same as previous but 6 December 2003, m. Pacific Co., Cement Creek, SR-401 2.5 mi S of Parpala Road in Naselle, 46.3341°N, - 123.8002°W, elev. 30 m., 15 January 2006, W. Leonard, C. Richart, from the litter of a riparian forest including Alnus rubra , Picea sitchensis , Sambucus cf. caerulea , and Polystichum munitum , m ff; Trap Creek Road 1.7 mi S of SR-6, Trap Creek Basin, 46.5403°N, - 123.6297°W, elev. 60 m., 19 November 2005, W. Leonard, C. Richart, mm ff. Wahkiakum Co., only known from the type locality.

Figures

FIGS. 242–244. Loomisiella spp. Fig. 242. Loomisiella evergreen, anterior gonopods, posterior view. Figs. 243–244. Loomisiella pylei. Fig. 243. Right anterior gonopod, lateral view. Fig. 244. Gonopod complex, anterior view.

FIGS. 245–250. Loomisiella spp. Figs. 245, 246. Loomisiella pylei. Fig. 245. Left anterior gonopod, posterior view. Fig. 246. Left posterior gonopod coxite, posterior view. Figs. 247–250. Loomisiella evergreen. Fig. 247. Left anterior gonopod, posterior view. Fig. 248. Left anterior gonopod, lateral view. Fig. 250. Posterior gonopods, anterior view.