Loomisiella evergreen, Shear & Richart & Wong, 2020

Shear, William A., Richart, Casey H. & Wong, Victoria L., 2020, The millipede family Conotylidae in northwestern North America, with a complete bibliography of the family (Diplopoda, Chordeumatida, Heterochordeumatidea, Conotyloidea), Zootaxa 4753 (1), pp. 1-78 : 66-68

Loomisiella evergreen , new species

Figs. 240–243 , 247–250

Types: Male holotype and male and female paratypes from WASHINGTON: Thurston Co., The Evergreen State College , Olympia , beside main parking lot, 47.0791°N, 122.9776°W, elev. 60 m, collected 14 February 2003, by W. Leonard, C. Richart .

Diagnosis: Distinct from L. pylei , n. sp. in the more broadly flared tips of the anterior gonopods and in lacking a mesal plate-like process of the posterior gonopod coxites. The posterior gonopod telopodites of L. pylei are reduced, as in this species, but comparatively larger.

Etymology: The species epithet, a noun in apposition, is from the type locality on the campus of The Evergreen State College, Olympia, Washington.

Description: Male paratype: Length, 4.8 mm. Thirteen ocelli in oval to triangular group. Metazonital shoulders poorly developed, segmental setae curved, acute. Legpairs one and two reduced, pairs three to five encrassate, pairs six and seven about as postgonopodal legs; femora three and four with acute basal knobs. Anterior gonopods (ag, Figs. 240 , 242, 243 , 247, 248 ) widely separated at base, embracing posterior gonopod coxites, narrowed, with right-angled or less bend basal to flaring tip; small processes mesal at base. Posterior gonopod telopodites strongly reduced, femur about one-third length of prefemur, sternum broad, well-sclerotized anteriorly. Posterior gonopod coxites (pgc, Figs. 240, 241 , 249, 250 ) short, broad, anteriorly cupped, with two curved processes distolaterally, two smaller processes arising from anterior face of body of coxite. Coxae 10 and eleventh prefemora typical of family.

Female 5.0 mm long, nonsexual characters as in male.

Distribution: WASHINGTON: Grays Harbor Co., FS-2153 5.3 mi N of Kelley Road, Canyon River basin, 47.3035° N, - 123.5081°W, elev. 160 m, 17 January 2003, by W. Leonard, mm ff; same as previous but 12 December 2004; Middle Satsop Road 2.7 mi N of Monte Elma Road, 47.0332°N, - 123.5258°W, elev. 20 m., 17 January 2003, W. Leonard et al., riparian forest, mm ff; Canyon River, 6 mi W, 1.25 mi N of Matlock, ca. 47.2546°N, - 123.5346°W, in maple and fern litter, 17 January 2003, W. Leonard et al., mm f. Thurston Co., Capitol State Forest, Potosi Creek, 2 mi S of Rock Candy Mountain, ca. 46.9809°N, - 123.109°W, ca. 290 m., estimated coordinates from Potosi Creek 1 horiz mi S of Rock Candy Mountain Summit, 10 December 2002, W. Leonard et al., mm ff.

Figures

FIGS. 240, 241. Loomisiella evergreen. Fig. 240. Gonopod complex, posterior view. Fig. 241. Posterior gonopods, anterior view.

FIGS. 242–244. Loomisiella spp. Fig. 242. Loomisiella evergreen, anterior gonopods, posterior view. Figs. 243–244. Loomisiella pylei. Fig. 243. Right anterior gonopod, lateral view. Fig. 244. Gonopod complex, anterior view.

FIGS. 245–250. Loomisiella spp. Figs. 245, 246. Loomisiella pylei. Fig. 245. Left anterior gonopod, posterior view. Fig. 246. Left posterior gonopod coxite, posterior view. Figs. 247–250. Loomisiella evergreen. Fig. 247. Left anterior gonopod, posterior view. Fig. 248. Left anterior gonopod, lateral view. Fig. 250. Posterior gonopods, anterior view.