Lithostege coassata, Hubner, Hubner

Sh, Hossein Rajaei, Stüning, Dieter & Viidalepp, Jaan, 2011, A review of the species of Lithostege Hübner, [1825] 1816 (Lepidoptera: Geometridae, Larentiinae), occurring in Iran and adjacent countries, with description of two new species from Iran and Pakistan, Zootaxa 3105, pp. 1-46 : 18-19

L. coassata (Hübner)

( Figs. 22 , 48 , 71 ; Map 5)

Ortholitha coassata Hübner, [1825] : 338 [Replacement name for Geometra duplicata Hübner, [1817] ). Type locality: Europe. Geometra duplicata Hübner, [1817] : pl. 95, fig. 491. Types not traced. Type locality: Europe.

Minoa assinata Freyer, 1830: 123 , pl. 132, fig. 2. Type locality: Odessa region, Ukraine.

Eubolia coassaria Boisduval, 1840: 202 . [Emendation]

Lithostege coassata: Prout, 1914: 173 , pl. 6e (as duplicata ); Prout, 1938: 240; Viidalepp, 1996: 47; Parsons et al., 1999: 545.

Material examined. 29 3, 103 Ƥ: 19 3, 95 Ƥ: N-Iran, Masandaran, Gonbad Qabus, O-m Zone, 28.4.1969, Einh. Sammler leg., preps 3 1041/2010 and 1042, Ƥ 1043/2010 H. R.; 1 3: Turkestan Oriental, Fort-Naryne, Prov. Semirechgensee, S. Akulin, 1920–1911 [m], gen. prep. 1040/2010 H. R.; 2 3: NW-Iran, 15 km SÖ. Maku, 1050 m, 3.6.1975, H.G. Amsel leg., preps 1068/2010 and 1069/2010 H. R.; all in SMNK. 1 Ƥ: same data, 6.6.1975, [leg.] Abai; 1 3: [ Iran, Golestan prov.] Gorgan, 14.4.1970, gen. prep. 1070/2010 H. R.; 1 3: NW-Iran, 40 km westl. Marand, 1100 m, 6.6.1975, H. G. Amsel leg., gen. prep. 1071/2010 H. R.; 1 Ƥ: [ Iran, Golestan prov.] Gorgan, Naharkhoran, 500 m, 5.6.1982, [leg.] Hashemi; 1 Ƥ: [ Iran, Golestan prov.] Gorgan, 26.4.1970; 1 3: [ Iran, Golestan prov.] Gonbad Ghabous, 11.5.[19]69, gen. prep. 1084/2010 H. R.; 1 3, 1 Ƥ: [ Iran] Azarbaidjan, Moghan, 2.5.1968, [leg.] Arghand; all in HMIM. 3/Ƥ: Turkmenistan: Kopet-Dagh Mts.: Kara-Kala, Ai-Dere, Tshuli, Yablonovka, Askhabat, Keledjar, Keshi, Kyzyl-Arvat; Kara-Bogaz; Repetek; Morgunovskii, Badkhyz, in ZIAT. 1 3, 2 Ƥ: „Turkmeenia” [ Turkmenistan] 1977, Sumbar riv., Ai-Dere, 24–25.04, J.V.; 1 3: the same locality but 10.05.1974, P. Ivinskis leg.; 1 Ƥ: “Turkmeenia” [ Turkmenistan], Morgunovka, 0 3.05.1974, P. Ivinskis leg.; 1 Ƥ: the same locality but 26.03.1987, E. Veromann leg.; all in IZBE. Preparations of genitalia: 6 3, 2 Ƥ.

Description & Diagnosis. Wingspan 26–32 mm. Wings brownish-grey, with darker brown pattern elements. Similar to L. griseata griseata in size and wing-shape, but distinguishable by the presence of transverse bands or lines demarcating the median area of forewing and an oblique line reaching the apex of forewing. Also an indistinct antemedian line present. Underside concolorous, almost without pattern. Also similar to L. luigi , but the latter is smaller, the transverse lines are indistinct, less oblique and the submarginal line does not reach the apex ( Fig. 22 ). Male genitalia ( Fig. 48 ) similar to farinata , but the apical projection from costa shorter and pointed, curved ventrad; harpe rather elongate, dorsal arm shorter, stouter, with longer spines apically: Uncus longer, distally trucate, saccus shorter; aedeagus shorter, less narrow. Female genitalia ( Fig. 71 ) with a narrow, tubular ductus bursae; corpus bursae globular, completely spined internally; a membranous anterior diverticulum is present.

Bionomics. Univoltine, common on irrigated and cultivated areas in arid zones. Specimens studied are collected end of March-beginning of June, at altitudes of 500–1100 m.

Additional data on distribution based on literature sources. Tsvetajev (1972): Repetek; Falkovitsh (1986): Uzbekistan; Mirzayans and Kalali (1970): Gonbad-e Qabus (Golestan prov.); Gorgan (Golestan prov.) ( Lederer, 1871); Kopet-Dagh (Khorasan-e Shomali) ( Viidalepp, 1988).

Distribution. Widely distributed from southern Ukraine and southern European Russia to SW Siberia, Turkmenistan and N Iran (Map 5).

Figures

FIGURES 22 – 26. Adult moths of Lithostege Hübner (left column, a: upperside, right column, b: underside). 22. L. coassata (3, Gonbad, N Iran); 23. L. ancyrana (3, Aladag, Turkey); 24. L. narynensis (3, Samarkand, Kasakhstan); 25. L. griseata griseata (3, Agri-Dagh, Armenia); 26. L. griseata gigantea (3, Karaj, N Iran). Scale bar: 10 mm.

FIGURES 48 – 51. Lithostege Hübner: Male genitalia. 48. L. coassata (gen. prep. 1041 H. R.; Gonbad, N Iran), a: genital capsule, b: aedeagus, lateral view; 49. L. ancyrana (gen. prep. 1013 H. R.; Gürün, Turkey), a: genital capsule, b: aedeagus, lateral view; 50. L. narynensis (gen. prep. 1055 H. R.; Narine, Turkestan, type locality), a: genital capsule, b: aedeagus, lateral view; 51. L. griseata griseata (gen. prep. 1048 H. R.; Agri-Dagh, Armenian), a: genital capsule, b: aedeagus, lateral view. Scale bar: 1 mm.

FIGURES 66 – 71. Lithostege Hübner: Female genitalia & details of abdominal integument. 66. L. samandooki sp. nov. (gen. prep. 1076 H. R.; type locality, S Iran); 67. L. senata (gen. prep. 8611 H. R.; Uzboi, Turkmenistan); 68. L. fissurata inanis (gen. prep. 1012 H. R.; Baluchistan, E Iran), a: genital organ, b: 7 th tergite, dorsal view; 69. L. parva (gen. prep. 1615 H. R.; Wachsch- Tal, Tajiikistan), a: genital organ, b: terminal segments of abdomen, lateral view; 70. L. turkmenica (gen. prep. 7649 J. V.; Uzbekistan, Karakum desert); 71. L. coassata (gen. prep. 1043 H. R., Gonbad, N Iran). Scale bar: 1 mm.