Lispe albimaculata Stein

Pont, Adrian C., 2019, Studies on the Australian Muscidae (Diptera). VIII. The genus Lispe Latreille, 1797, Zootaxa 4557 (1), pp. 1-232 : 24-30

Lispe albimaculata Stein

( Figs 28–41 )

Lispa albimaculata Stein, 1910: 561 .

Lispa albimaculata .— Stein, 1919: 141.

Xenolispa albimaculata .— Malloch, 1923: 610; Lee, Crust & Sabrosky, 1956: 306.

Lispe albimaculata .— Séguy, 1937: 182; Pont, 1989: 692.

Type-material. Stein described this species from 3♂ from Brisbane and 1♀ from Botany Bay, from material in HNHM. The 3♂, collected in 1900 by L. Biró, were studied by me in HNHM in 1986 (Pont & Werner, 2006: 18; Pont, 2013: 53), having survived the destruction of the HNHM Calyptrate Diptera collections in 1956; the Botany Bay ♀ has not been found ( Pont, 2013). The three ♂ syntypes are slightly mouldy, and two have the head missing. The ♂ with the head is herewith designated as lectotype, and the other 2♂ are paralectotypes. Each of the three has two printed labels: “ Australia / Biró” and “ Queensland / Brisbane”. In addition, the lectotype has Stein’s handwritten label on pale green paper “ Lispa / albimaculata / sp.n.”. Apart from the mould, it is in good condition. All three specimens belong to Lispe albimaculata as here defined.

Diagnosis. This is the only Australian species with a dark band in apical part of wing. The ♂ may also be easily recognised by the large boss-like tubercle on sternite 3 ( Fig. 33 ).

Material Examined. AUSTRALIA: Western Australia: 12 km NE by N of Broome, 17.52S 122.18E, 25.viii.1976, I.F.B. Common, 1♂ ( ANIC); 3 km ENE of Cape Borda, West Kimberley , 27.iv.1977, D.H. Colless, 1♀ ( ANIC) ; Carson Escarpment , 14.49S 126.49E, 9–15.viii.1975, I.F.B. Common & M.S. Upton, 12♂ 7♀ ( ANIC & BMNH)) ; N.W. Coastal Highway , 7 miles NE of Karratha, on Eucalyptus sp., 16.ii.1973, E.M. Exley, 1♂ ( QMBA) ; 3 km S of Coulomb Point, West Kimberley , at light, 20.iv.1977, D.H. Colless, 2♂ 2♀ ( ANIC) ; Drysdale River , 15. 02S 126.55E, 3–8.viii.1975, I.F.B. Common & M.S. Upton, 2♂ ( ANIC) ; Kimberley Res. Stn , light trap, x.1961, K.T. Richards, 1♀ ( ANIC) ; Logues Springs , 102 km SE by E of Broome, 18.25S 123.05E, 18.viii.1976, I.F.B. Common, 1♂ ( ANIC) ; 8 km WSW of Martin 's Well, West Kimberley , Malaise trap, 29.iv.1977, D.H. Colless, 1♂ ( ANIC) ; Millstream , 22 & 24.x.1970, D.H. Colless, 2♂ 1♀ ( ANIC) ; Millstream , at light, 12.iv.1971, D.H. Colless, 1♀ ( ANIC) ; 10 km NW by N of Mining Camp, stop 15, Mitchell Plateau, 14.45S 125.47E, Kimberley District , 17.v.1983, D.C.F. Rentz & J. Balderson, 1♂ ( ANIC) ; Murchison River , 27. 49S 114.41E, 28.iii.1971, E.F. Riek, 1♂ 1♀, and Upton & Mitchell, 1♀ ( ANIC) . Northern Territory: 8 km S of Alice Springs , on Wahlenbergia , 29.x.1974, E.M. Exley & R.I. Storey, 1♀ ( QMBA) ; 12 km SW of Alroy Downs , at light, 10.iv.1976, D.H. Colless, 2♂ 2♀ ( ANIC) ; Alice Springs , 14.ii.1966, J.A. Grant, 1♀ ( BMNH) ; 32 km WNW of Alice Springs , 8.x.1978, D.H. Colless, 2♂ ( ANIC & BMNH) ; 30 km NW by W of Alice Springs , at light, 7.x.1978, D.H. Colless, 1♂ ( ANIC) ; Amadeus Basin , 3.vii & 16.ix.1962, P. Ranford, 2♂ ( ANIC) ; Ayers Rock , 18.viii.1972, C.W.Sabrosky, 1♀ ( USNM) ; Batten Creek, 31 km WSW of Borroloola, Malaise trap, 16.iv.1976, D.H. Colless, 1♂ ( ANIC) ; Batten Point , 30 km NE by E of Borroloola, at light, 18.iv.1976, D.H. Colless, 1♀ ( ANIC) ; Birraduk Creek , 17 km WSW of Nimbuwah Rock, 5.vi.1973, D.H. Colless, 1♂ ( ANIC) ; Border Waterhole, 15 km W of Musselbrook Resource Centre, Lawn Hill Nat. Park , 18.3644S, 137.5930E, 200 m, 2.v.1995, G. Daniels & M.A. Schneider, 1♂ 5♀ ( QMBA) ; 12 km NNE of Borroloola , 19.iv.1976, D.H. Colless, 1♂ ( ANIC) ; 22 km WSW of Borroloola , at light, 16.iv.1976, D.H. Colless, 1♂ 1♀ ( ANIC) ; same locality, Malaise trap, 17.iv.1976, D.H. Colless, 1♂ ( ANIC) ; Caranbirini W/H, 33 km SW of Borroloola, at light, 21.iv.1976, D.H. Colless, 1♀ ( ANIC) ; Chilla Well , 21.31S, 130.59E, 5.xi.1968, A.O. Nicholls, 22♂ 2♀ ( ANIC) , & 1♂ 1♀ ( BMNH) ; Cooper Creek , 19 km E by S of Mt Borradaile, 5.vi.1973, D.H. Colless, 1♂ ( ANIC) ; 34 miles NW of Dorisvale H.S., 14.13S, 130.55E, M. Mendum, 2♀ ( ANIC) ; 46 miles E of Frewana, Barkly Tableland , 17.viii.1960, E. Exley, 3♂ 4♀ ( QMBA) & 1♂ 1♀ ( OUMNH) ; Horn Islet, Pellew Group , 8–14.ii.1968, B. Cantrell, 1♂ ( QMBA) ; Hugh River , 28 miles WSW of Alice Springs, 10.ii.1966, Britton, Upton & McInnes, 1♂ ( ANIC) ; Hull River, Mannanana Range, Petermann Ranges , 18.v.1983, G.A. Holloway, 1♀ ( AMS) ; 14 miles W of Humpty Doo , 10 & 14.vi.1964, K.R. Norris, 3♂ 7♀ ( ANIC, BMNH) ; Katherine , 7.vi.1964, K.R. Norris, 2♂ 4♀ ( ANIC, BMNH) ; Koongarra , 15 km E of Mt Cahill, at light, 6–9.iii.1973, D.H. Colless, 2♂ 3♀ ( ANIC, BMNH) ; Koongarra area , 15–22 km E of Mt Cahill, 9.iii.1973, D.H. Colless, 1♂ ( ANIC) ; 8 km N of Kulgera , 21.ix.1978, M.S. Upton & R.A. Barrett, 1♀ ( ANIC) ; Maggie Springs at Uluru National Park , 4.iv.1980, G.F. Hevel & J.A. Fortin, 3♂ 4♀ ( USNM) ; McArthur River , 14 km S by W of Cape Crawford, at light, 11–12.iv.1976, D.H. Colless, 2♂ 1♀ ( ANIC) ; McArthur River , 48 km SW by S of Borroloola, at light, 13–14.iv.1976, D.H. Colless, 5♂ 5♀ ( ANIC, BMNH) ; Nourlangie Creek , 8 km N of Mt Cahill, at light, 16.vi.1973, D.H. Colless, 1♀ ( ANIC) ; Roe Creek , 12 km SW by W of Alice Springs, at light, 9.x.1978, D.H. Colless, 1♀ ( ANIC) ; Standley Chasm , NW of Jay Creek, 4.x.1972, Z. Liepa, 2♀ ( ANIC) ; Surprise Creek, 45 km SW by S of Borroloola, Malaise trap, 15.iv.1976, D.H. Colless, 2♂ ( ANIC) ; Todd River , 9 km N by E of Alice Springs, 28.ix.1978, M.S. Upton & R.A. Barrett, 1♂ ( ANIC) ; same locality & date, 23.38S 133.53E, J.C. Cardale, 1♂ 1♀ ( ANIC) ; Todd River , 9 km N by E of Alice Springs, at light, 10.x.1978, D.H. Colless, 2♂ ( ANIC) ; 12.8 km N–NE of Victoria River Downs , 8.vi.1973, L.P. Kelsey, 1♀ ( ANIC) ; Wauchope , 6.iv.1980, G.F. Hevel & J.A. Fortin, 1♂ 1♀ ( USNM) . Queensland: no further data, 1♀ ( BMNH) ; Acacia Ridge, Brisbane , 19.vii.1963, E.C. Dahms, 1♂ 1♀ ( QMBA) ; Ayr , 30.ix.1960, R. Hughes, 1♂ ( ANIC) ; Amphitheatre waterhole area, 27 km N of Musselbrook Resource Centre, Lawn Hill Nat. Park , 18.21.08S, 138.09.43E, 200 m, m. v. lamp , 3.v.1995, G. Daniels & M.A. Schneider, 1♂ 2♀ ( QMBA) ; Ayr, Swans Lagoon , on Leersia hexandra , 11.iii.1986, J.den Hollander, 1♂ ( NMWC) ; Babinda, J.F. Illingworth, 1♂ ( USNM) ; Barcoo River, near Blackall , 15.vi.1976, J.F. Donaldson, 4♂ 4♀ ( QDPC, BMNH) ; Belyando River, approx. halfway between Charters Towers and Clermont , 6.vii.1975, M.S. & B.J. Moulds, 4♀ ( AMS, BMNH) ; Big Mitchell Creek, Mareeba–Molloy Road , 4.v.1967, D.H. Colless, 1♂ 1♀ ( ANIC) ; Bilyana , 25.7 km SW of Tully, 15.vii.1971, Z. Liepa, 1♂ ( ANIC) ; Bluff Range, near Biggenden , 7–19.viii.1971, H. Frauca, 3♀ ( ANIC) ; Boggomoss, Taroom District , 25.31S, 150.03E, 13.xi.1996, C.J. Burwell & S. Evans, 1♀ ( QMBA) ; 25 km NW by N of Boulia , at light, 8.iv.1976, D.H. Colless, 2♂ 1♀ ( ANIC) ; Bribie I., Malaise trap, 2-11.ix.1983, E. Hamacek, 3♀ ( QDPC) ; Bribie Island ( QDPC) Fisheries site, heathland–Acacia regrowth, 27.03S, 153.11E, Malaise traps, 5–12.ix.1997, S. Winterton, N. Power, D. White, 1♂ 4♀ ( QMBA) ; Brisbane , 1900, L. Biró, ♂ lectotype & 2♂ paralectotypes ( HNHM) ; Brisbane , 2.ii.1954, 1♂ ( QMBA) ; Brisbane , 4.viii.1955, F.A. Perkins, 4♂ 4♀ ( QMBA) ; Brisbane , 17.viii.1955, F.A. Perkins, 1♀ ( QMBA) ; Brisbane , 23.viii.1955, F.A. Perkins, 1♂ ( QMBA) ; Brisbane , 17.xi.1962, T. Brooks, 1♂ ( QMBA) ; Brisbane , 25.viii.1967, J. Hargreaves, 1♂ ( QMBA) ; Brisbane, DPI Indooroopilly site, Malaise trap, viii.1977, J. Grimshaw, 1♀ ( QDPC) ; same locality, 15–22.xi.1979, 1♂ ( QDPC) ; Brisbane, Sanford , 29 & 30.vii.1972, B.Hocking, 2♂ ( QMBA & OUMNH) ; Lake Broadwater via Dalby , 9.xii.1987, m. v. light, D.K. Yeates , 1♂ ( QMBA) ; Bundaberg , 8.iii.1972, H. Frauca, 1♀ ( ANIC) ; 2 miles N of Bundaberg, Tea-tree swamp, 26.vi.1971, H. Frauca, 2♂ 1♀ ( ANIC) ; Burnett River, Bundaberg , 24–28.iii.1972, H. Frauca, 1♂ ( ANIC) ; Burpengary , no date, T.L. Bancroft, 2♂ ( BMNH) ; Caboohure , 7.v.1914, S. Johnson, 1♂ ( QNBA) ; Cairns , on window, J.F. Illingworth, 1♀ ( USNM) ; Caloundra , 17.viii.1934, F.A. Perkins, 1♂ ( QMBA) ; Caloundra , 27.viii.1963, H.C. Shepherd, 1♀ ( QMBA) ; Canungra , 27.iii.1937, N.T.M. Yeates, 1♂ ( QMBA) ; Carnarvon Nat. Park, Mt. Moffatt section, 25.03.49 S, 148.01.57E, Malaise trap, 27–28.xi.1997, J. Skevington, C. Lambkin, 3♂ ( QMBA) ; Carnarvon Nat. Park, Mt. Moffatt section, swamp edge near landing ground, 25.02.26 S, 147.56.13E, 2 m, Malaise trap, 27.xi.1997, J. Skevington, C. Lambkin, S. Evans, 1♂ ( QMBA) ; Charleville , 7.iv.1976, D.H. Colless, 1♂ ( ANIC) ; 15 km S of Charleville , 6.iv.1976, D.H. Colless, 1♂ ( ANIC) ; Childers , 5.ix.1973, H. Frauca, 1♀ ( ANIC) ; Coolum , 29.vii.1972, E.C. Dahms, 1♂ ( QMBA) ; Coppermine Creek , 67.5 km N of Marlborough, 5.vii.1971, Z. Liepa, 1♂ 2♀ ( ANIC) ; Dawson River, near Duaringa , 8.v.1970, Z. Liepa, 2♂ ( ANIC) ; Desailly Creek , 10 km W by N of Mt Carbine, 16.30S, 144.55E, at light, 19.v.1981, D.H. Colless, 1♂ ( ANIC) ; Dalby , 23.x.1958, S. Sekhon, 2♂ 1♀ ( QMBA) ; Eidsvold , 11.iv.1924, Bancroft, 1♂ ( AMS) ; Einasleigh River Crossing, E of Georgetown, 21.vi.1976, J.F. Donaldson, 1♂ ( QDPC) ; 26 km W Fairview , 15.35S, 144.04E, m. v. lamp , 24.iv.1989, G. & A. Daniels, 1♂ ( QMBA) ; Fig Tree Creek , 39 km S of Ingham, 14.iv.1980, G.F. Hevel & J.A. Fortin, 1♂ ( USNM) ; 7–14 miles W of Herberton, via Watsonville , 1.v.1967, D.H. Colless, 2♂ 1♀ ( ANIC, BMNH) ; Hughenden , 1♂ ( USNM) ; DPI Indooroopilly site, Malaise trap, 6–12.xii.1984, 4♂ ( QDPC, BMNH) ; Inglewood , 1.ix.1925, 1♂ ( QMBA) ; Karara , 3.v.1954, O.R. Byrne, 1♂ ( QMBA) ; Jondaryan , 8.vi.1963, C. Speed, 1♂ ( QMBA) ; Lake Broadwater, near Dalby , 27.21S, 151.06E, m. v. lamp , 2.v.1987, G. & A. Daniels, 1♂ ( QMBA) ; 45 km W of Lynd River , wild rice,?D-Vac, 24.iv.1983, 1♂ ( QDPC) ; Mackay , 12.v.1970, Z. Liepa, 1♂ ( ANIC) ; Mackay , in grass on edge of sugar cane, D-Vac, 8.xi.1975, I.D. Galloway, 1♀ ( QDPC) ; Marlborough Creek , 2 km N of Marlborough, 29.i.1975, B.K. Cantrell, 4♂ 3♀ ( QDPC) ; McIlwraith Range, NE of Coen, Cape York Peninsula , 29.vi–5.vii.1976, J.F. Donaldson, 1♀ ( QDPC) ; Mooloolaba , 24.x.1965, C.F. Ashby, 1♂ ( ANIC) ; 3 km SW by S of Mt Baird , 15.10S, 145.07E, light & Malaise traps, 3–4.v.1981, D.H. Colless, 1♂ 3♀ ( ANIC) ; Mt Irving, 14.viii.1979, R.H. Broadley, 1♂ ( QDPC) ; 66 km NW of Mt Isa , at light, 9.iv.1976, D.H. Colless, 2♂ ( ANIC) ; Tamborine , 10.x.1979, B.K. Cantrell & J.F. Donaldson, 1♂ ( QDPC) ; Mungallala , 15.vi.1976, J.F. Donaldson, 1♂ ( QDPC) ; near monument, Musselbrook Resource Centre, Lawn Hill Nat. Park , 18.35.52S, 138.07.23E, 160 m, m. v. lamp , 19.iv.1995, G. Daniels & M.A. Schneider, 1♀ ( QMBA) ; Murrays Creek, 1km W of Musselbrook Resource Centre, Lawn Hill Nat. Park , 18.35.50S, 138.07.18E, 160 m, 20.iv.1995, G. Daniels & M.A. Schneider, 1♀ ( QMBA) ; Murrays Spring, 7 km W of Musselbrook Resource Centre, Lawn Hill Nat. Park , 18.35.15S, 138.04.28E, 200 m, 21.iv–14.v.1995, G. Daniels & M.A. Schneider, 13♂ 12♀ ( QMBA) & 1♂ 1♀ ( BMNH) ; Musselbrook Creek, 11 km ESE of Musselbrook Resource Centre, Lawn Hill Nat. Park , 18.36.45S, 138.07.46E, 140 m, 8.v.1995, G. Daniels & M. A. Schneider, 1♂ 2♀ ( QMBA) ; Noosa area , 3.v.1999, J. & A. Skevington, S.A. Marshall, 1♂ ( QMBA) ; 1 mile E of Paluma , 2800 ft, m. v. lamp , 17.i.1970, G.A. Holloway, 1♂ ( AMS) ; Ranges Bridge, Condamine River , 25.iv.1963, B.V. Timms, 1♂ ( QMBA) ; Ridgepole Waterhole, 24 km ESE of Musselbrook Resource Centre, Lawn Hill Nat. Park , 18.40.15S, 138.22.15E, 180 m, 23.iv–14.v.1995, G. Daniels & M.A. Schneider, 26♂ 20♀ ( QMBA) & 1♂ 1♀ ( BMNH) ; Samsonvale Cemetery , 8.5 km SSE Dayboro, 27.16S, 152.52E, 3.ix.1996, S. Evans, 1♀ ( QMBA) ; Stanthorpe, Mount Marlay , 28.40S 151.57E, 10–11.x.1998, J. & A. Skevington, 1♂ ( QMBA) ; 15 miles N Tambo , 2.v.1963, E.C. Dahms, 1♀ ( QMBA) ; Tibrogargan Creek , 4.ix.1956, F.A. Perkins, 1♂ ( QMBA) ; Tinaroo Falls Dam, open savannah, 27.iv.1967, D.H. Colless, 1♂ ( ANIC) ; Wenlock H., Cape York Peninsula, 25.vi.1976, J.F. Donaldson, 1♀ ( QDPC) ; 205 km W Windorah, Malaise trap in dry creek bed, 4.ix.1997, S. Winterton, J. & A. Skevington, 1♂ ( QMBA) ; Woodford , 26.x.1958, S. Sekhon, 1♂ ( QMBA) . New South Wales: Beecroft Peninsula , S of Currarong, 20.xii.1974, G. Daniels, 1♀ ( QMBA) ; 13 miles SE of Braidwood , 14.x.1955, I.F.B. Common, 1♂ ( ANIC) ; Brunswick Hds , viii.1963, D.E. Havenstein, 7♂ 6♀ ( ANIC, BMNH) ; Careel Bay, Avalon, Casuarina swamp, 9.vi.1962, D.K. McAlpine, 1♂ 2♀ ( AMS) ; Coff 's Harbour , 25.vi.1976, Z. Liepa, 1♂ ( ANIC) ; Como West, near Sydney , 5–8.xii.1972, L.S. Willan, 1♂ ( AMS) ; same locality, m. v. light , 6–9.xi.1972, 1♀ ( AMS) ; Goonoo State Forest , 5 miles S of Mendooran, 24.iii.1971, D.K. McAlpine, 8♂ 3♀ ( AMS, BMNH) ; Goonoo State Forest , 31.54S, 148.57E, Malaise trap, 12–14.i.1994, M.E. Irwin & D.K. Yeates, 1♀ ( QMBA) ; Kurnell , 28.ix.1954, D.K. McAlpine, 1♂ ( AMS) ; Mendooran , 8.xii.1973, G. Daniels, 1♂ ( AMS) ; 5 miles S of Mendooran , m. v. lamp , 23.iv.1973, G. Daniels, 1♂ 1♀ ( AMS) ; Moonan Forest, Gloucester Tops , 19.xi.1976, K.R. Norris, 1♀ ( ANIC) ; Tarro, Hunter River , 18.x.1922, 1♂ ( USNM) ; Tubrabucca, Upper Hunter , 19.x.1956, D.K. McAlpine, 1♀ ( AMS) ; Warrumbungle Nat. Park , 22.iv.1973 & 26.ix.1976, A. & G. Daniels, 2♂ ( AMS) ; West Head , 4.xi.1972, E.A. Fonseca, 2♂ ( BMNH) . Australian Capital Territory: Black Mt , light trap, 15.x.1955 & 15.xii.1964, I.F.B. Common, 1♂ 1♀ ( ANIC) . South Australia: 71 km S by E Cooper Pedy , at light, 22.xi.1977, J.A.L. Watson, 1♂ ( ANIC) ; Dutchmans Stern Conservation Park , Malaise trap, 32.17.49S, 137.58.59E, 6–7.x.1997, S. Winterton, J. & A. Skevington, C. Lambkin, 2♂ ( QMBA) ; Flinders Ranges, Parachilna Creek , 1.ix.1977, A.N. Clements, 1♀ ( BMNH) ; Flinders Ranges Nat. Park, Dingley Dell camping ground, Oraparinna Creek , Malaise trap, 32.21.17S, 138.42.16E, 8–9.x.1997, J. & A. Skevington, S. Winterton, C. Lambkin, 2♂ ( QMBA) ; Gammon Ranges Nat. Park , Malaise trap, 13.x.1997, J. Skevington, S. Winterton, C. Lambkin, 1♂ 1♀ ( QMBA) .

Description ♂ ♀. Head ( Figs 28–29 ). Ground-colour black. Frons at broadest point 0.28 (♂) or 0.33 (♀), at lunula 0.45 (♂) or 0.41 (♀), of maximum head-width (frontal view). Fronto-orbital plate mostly shining, with a small brown pruinose area around upper frontal seta followed by a short silvery area before lunula; parafacial and face mainly yellowish pruinose, facial ridges tinged with brown; gena and lower 2/3 of occiput grey dusted, upper 1/3 of occiput undusted, shining, not differentiated from fronto-orbital plates. Frontal triangle quite broad, shining black. Parafacial narrow, at middle 1/4 (♂) to 1/3 (♀) width of postpedicel; setulae short, fine, in 1 row. Antenna black, except tip of pedicel and base of postpedicel which are orange, pedicel often more extensively orange; postpedicel 2.5 (♂) to 2.8 (♀) times as long as broad, falling short of mouth-margin by about 1/3 its length. Arista plumose, the hairing at widest point about 1.5 times width of postpedicel. Vibrissa strong, vibrissal area otherwise bare. Gena narrow, about 0.1 of vertical eye-length; genal setae weak. Palpus yellow, swollen at tip ( Fig. 29 ).

Thorax. Ground-colour black. Scutum mainly shining from most angles but dull in dorsal view; in posterior view with a pair of brownish-grey vittae inside the dc lines, running from neck to scutellum and separated medially by the shining ground-colour (only narrowly separated in ♀); scutellum shining black, without dust; inner half of postpronotal lobe and all pleura (including notopleuron) light grey dusted, outer half of the lobe shining like scutum. Anterior spiracle brown. Acr setulae in 2 rows at suture. Dc 0+1, with 1–2 additional post dc groundsetulae slightly longer than the others ( Fig. 30 ). 1 pprn. 0+0 ia. 1+1 sa, the prst seta very weak. 1 proepisternal seta, usually with an adjacent setula. Proepimeral seta absent, this area with 2-4 tiny setulae. Anepisternum with 1 short seta in upper anterior corner; posterior row with 1 strong and 2 weak setae. 3 kepst, lower one closer to posterior than to anterior one, only the posterior one strong ( Fig. 31 ). Meron bare below spiracle and above hind coxa. Scutellum with disc sparsely setulose, bare on sides; ventral surface bare.

Legs. Fore coxa yellow, mid and hind coxae mainly black; trochanters yellow; femora black, extreme tips (knees) yellow, mid femur yellow on apical 1/4 to 1/5; ♂ tibiae yellow, fore tibia partly infuscated; ♀ fore tibia mainly dark, mid and hind tibiae yellow but partly infuscated; basal 1–2 tarsomeres yellow in ♂, weakly so in ♀, otherwise tarsomeres black. Fore coxa bare behind. Fore femur without av setae; pv surface with 3–4 short setae in apical half and 1 fine seta at base; pd row complete, short. Fore tibia without submedian setae; with short d and pv apical setae; in ♀ tibia slightly expanded, and white dusted along pd to p surfaces. Fore tarsomeres without modifications. Mid femur without av or pv setae; 1 a at basal third; without preapical ad; 1 p preapical, preceded by a p seta at apical 1/4. Mid tibia with 0 ad and 1 p; with moderate a, v and pv apicals, and a short d. Mid tarsosmeres without modifications. Hind coxa without a seta on posterior apical margin. Hind femur in ♂ with a row of short mainly spinose pv setae, none more than half femoral depth; ♂ without av setae, but the setulae in apical third becoming rather spine-like, similar to those on pv surface but usually shorter; ventral surfaces completely bare in ♀; ad row complete; 0 pd preapical. Hind tibia with 1 short ad, without av or pd; d preapical seta short, equal to tibial depth, the ad absent; 1 av but no pv apical. Hind tarsomeres not modified.

Wing. Narrow. Not darkened at base, but with a dark band beyond crossvein dm-cu, followed by a milky-white tip. Tegula and basicosta yellow. Crossvein r-m placed well apicad of the point where vein R1 enters costa; dm-cu straight, longer than apical section of vein CuA1. Vein M running straight to wing-margin. Calypters white to creamy, with creamy margins. Haltere yellow.

Abdomen. Ground-colour black. Syntergite 1+2 and tergites 3–5 not modified in shape ( Fig. 34 ); sternite 2 very narrow; ♂ sternite 3 with a large boss-like tubercle ( Fig. 33 ), and sternite 5 with a median excavation on hindmargin. Tergites subshining black with very thin dark brown dust; ♂ with white dusted spots as in Figs 32–33 (tergite 4 entirely dark); these spots all smaller in ♀, that on syntergite 1+2 restricted to dorsal surface, that on tergite 3 forming a band that extends to ventral edge, that on tergite 5 small and oval. ♂ epandrium subshining. ♀ ovipositor with upper part of tergite 7 white dusted, visible tergal areas otherwise dark like rest of abdomen. Sternites 1–3 grey dusted, sternite 4 subshining black. In ♂ syntergite 1+2 and tergites 3–5 each with 2 pairs of weak lateral marginal setae, those on tergite 5 forming a row, without discals; shorter and less erect in ♀. ♂ sternites 2–4 with sparse setulae.

Ƌ terminalia. Epandrium attached to tergite 5, the intermediate tergite 6 and syntergosternite 8 lost; with 2 spiracles on lower anterior edge ( Fig. 35 ). Sternite 6 vestigial but clearly developed, symmetrical, withdrawn beneath sternite 5. Surstylus well-developed, distinct and discrete from epandrium ( Fig. 35 ). Cercal plate moderately divided ( Fig. 36 ). Hypandrium attached to surstylus and, posteriorly, to wall of epandrium close to inner dorsal tip of cercal plate. Phallic complex ( Figs 37–38 ): praegonite absent; postgonite weakly sclerotised, short, curved; phallapodeme enlarged, bifid at mid-length and the branches fusing with hypandrium; phallus short, juxta eversible, spinulose and containing about 7 large thorn-bearing platelets.

♀ ovipositor ( Figs 39–41 ). Tergite 6 complete dorsally; tergite 7 complete dorsally and large, on lateral surface with an extra, large, superimposed fold or lobe on each side, in dry specimens this appearing shining and contrasting with the dusted tergite; tergite 8 divided dorsally, enlarged below. Sternites 6 and 7 large, broad, plate- like; sternite 8 consisting of 2 small plates and a third, median, plate. Epiproct V-shaped, with 1 pair of setae. Hypoproct not large, spinose. Cercus weakly spinose. 3 spermathecae.

Measurements. Wing-length, 3.0–4.0 mm (♂), 2.5–4.0 mm (♀). Body-length, 3.5–4.5mm (♂), 3.5–5.0 mm (♀).

Biology. Unknown. Adults appear to be euryoecious, having been collected in a wide range of habitats: rivers, lakes, swamps, inland and coastal, and even the most arid (Uluru).

Distribution. Known only from Australia (WA, NT, Q, NSW, ACT, SA).

Figures

FIGURES 28–33. Lispe albimaculata Stein. 28. ♂ head, lateral view. 29. ♀ palpus. 30. Outline of ♂ scutum in lateral view, showing the dorsocentral setae. 31. ♂ katepisternum. 32. ♂ abdomen, dorsal view. 33. ♂ abdomen, lateral view (arrow shows the boss-like tubercle on sternite 3). (28–31, NT, McArthur River; 32–33, Q, 2 miles N of Bundaberg.)

FIGURES 34–38. Lispe albimaculata Stein, ♂ terminalia. 34. Sternites 1–5. 35. Hypopygium, lateral view. 36. Cercal plate, dorsal view. 37. Phallic complex, lateral view. 38. Phallus, ventral view. (WA, Carson Escarpment.)

FIGURES 39–41. Lispe albimaculata Stein, ♀ ovipositor. 39. Dorsal view. 40. Ventral view. 41. Lateral view. (NSW, Brunswick Heads.)