Leuctra nigra (Olivier, 1811)

Vincon, Gilles, Boumans, Louis & Gattolliat, Jean-Luc, 2018, Reinstatement of Leuctrabiellensis Festa, 1942 (Plecoptera, Leuctridae), Alpine Entomology 2, pp. 35-43 : 35-36

Leuctra nigra (Olivier, 1811) Figs 1, 2 a–g

Nemoura nigra Olivier 1811: 186

Leuctra acuminata Bengtsson, 1933 (syn. fide Brinck 1949).

Leuctra nigra Oliv., Mosely 1932: 27, figs 39-40, pl. V/26; Kühtreiber 1934: 76, fig. 55; Despax 1951: 140, fig. 61c-d; Illies 1955: 68, fig. 61; Aubert 1959: 65, figs 159-160; Mendl 1968: 70; Kis 1974: 87, 88, figs 37 a–e; Ravizza and Vinçon 1998: 303, fig. 6 a–f; Teslenko and Zhiltzova 2009, 223, figs 1235-1239; Lubini et al. 2012: 180, fig. 433, 196, fig. 477.

Locus typicus : France, Versailles.

Material examined.

FRANCE: BRITANY: Finistère: Rosnoen, Toulencoat, 22.03.2001, 6♂ (Manach leg). VOSGES MASSIF: Vosges: SW Colmar,> Wasserbourg, Krebsbach, Fecht trib., Ill trib., 700 m, 17.05.1995, 10♂, 13♀; Ballon d’Alsace, Saut-de-la-Truite, 23.08.1996, 1♂, 8♀ (Vin). MASSIF-CENTRAL: Saône-et-Loire: Morvan Massif, Haut-Folon, 800 m, 14.07.1994, 5♂, 7♀ (Vin); Puy-de-Dôme: Forez Mountains,> Vertolaye,> Chansert Pass, Jasserie de la Croix Fossat, 1500 m, 17.07.1994, 3♂, 7♀; St Pierre-la-Bourlhonne, cross road to the Chansert Pass, Dore trib., 17.07.1994, 1♀; W. Clermont-Ferrand, between Pontaumur and La-Forêt, b., 21.04.2009, 1♂, 5♀ (Vin). Loire: Mont Pilat, Graix, 900 m, 17.07.1994, 1♀; Mont Pilat, Crêt de la Perdrix, Gier b., 1400 m, 17.07.1997, 3♀ (Vin). Ardèche: Mont Gerbier-de-Jonc, Loire spring, 16.06.2001, 2♀; Lachamp-Rafael, Bois de Cuze, peat bog, 1350 m, 15.06.2001, 4♂, 6♀ (Vin). Cantal: Plomb-du-Cantal Massif,> Super Lioran, 1300 m, 30.07.2004, 3♀ (Vin). Tarn: NW slope of Pic de Nord, Arnette trib., Thoré trib., 850 m, 1.05.1991, 4♂, 2♀ (Vin). Aude:> Citou, spring, trib. of Argent-double, 900 m, 1.07.1989, 1♀, 1.05.1991, 20♂, 3♀ (Vin). JURA MASSIF: Ain: La Vattay Ski Station, Faucille Pass, 1300 m, 30.05.1991, 1♂, 27.06.1995, 1♀, 13.07.1996, 1♀ (Vin); Valserine, Combe de Mijoux, Le Planet, 1200 m, 19.07.1991, 3♀; Valserine, Joux Verte, 19.07.1991, 2♀ (MZL); Doubs: Jougne, Swiss Border, Les Tavins, La Tavine, Orbe trib., 900 m, 28.06.1995, 1♀ (Vin). ALPS: Haute-Savoie: Montet Pass,> Vallorcine, Eau-Noire trib., 1450 m, 21.06.1991, 2♂, 15♀ (Vin); Savoie: Allevard, Val Pelouse, 1400 m, 7.06.1987, 10♂, 1♀; Saisie Pass, Nant Rouge trib., 1600 m, 22.05.1991, 1♂, 1♀ (Vin). Isère: Chartreuse Massif, <Charmette Pass, Tenaison b., Vence trib., 1200 m, 28.05.1988, 1♂ (Vin); Belledonne Massif, below Oursière Cascade, Doménon trib., brook, 1500 m, 4.06.1995, 1♂, 1♀; <Chamrousse, Fénérieux b., Prémol trib, between Prémol Forest House and Luitel Lake, 1200 m, 10.06.1985, 1♂, 3♀; <Chamrousse, Premol Forest House, Sonnant trib., 1100 m, 10.06.1985, 1♀, 24.05.1998, 1♀, 19.05.2013, 7♂, 5♀, 2.06.2013, 2♂, 1♀ (Vin). SWITZERLAND, JURA MASSIF: NW Tramelan, Gruère Lake, b. right side of the lake, 20.06.2012, 5♂, 11♀ (Vin). GERMANY: Fulda, 1981, 2♂, 1♀ (Zwick leg), SLOVAKIA: Vysoké Mountains, Tatry, Podspady, Javorinka r., 11.07.1967, 5♂, 5♀ (Steinmann & Mur).

Complementary description.

Based on French specimens: Savoie, Allevard, Val Pelouse, 1400 m, 7.06.1987. Male (Fig. 2 a–f): Tergite VI with two small triangular appendices pointing upwards and backwards in side view (Fig. 2b). Tergite VII with non-interrupted anterior margin and wide median bell-shaped membranous area. Tergite VIII with two strong teeth pointing upwards and backwards in side view (Fig. 2a, b, f); teeth sub-rectangular in dorsal view (Fig. 2f). Tergite IX with anterior margin interrupted on nearly half segment width, lateral edges triangular ending in sharp angles near anterior margin. Lateral lobes of paraprocts with a sclerotized hook-shaped expansion turning backwards and extending along distal part of specilla (Fig. 2 c–d). Specilla straight in ventral and side views (Fig. 2 c–d), slightly curved at their tip (Fig. 2c). Sternite IX: ventral vesicle racket-shaped (Fig. 2e). Female (Fig. 2g): subgenital plate with two rounded lobes separated by a small rounded lamella; a triangular sclerite placed between the 2 lobes is visible beneath transparent cuticle.