Leptostigma alaemacula, Archibald & Rasnitsyn, 2023

Archibald, S. Bruce & Rasnitsyn, Alexandr P., 2023, Cimbicidae (Hymenoptera, ‘ Symphyta’) in the Paleogene: revision, the new subfamily Cenocimbicinae, and new taxa from the Eocene Okanagan Highlands, Zootaxa 5278 (1), pp. 1-38 : 22-23

Leptostigma alaemacula n. sp.

http://zoobank.org/ urn:lsid:zoobank.org:act:CB061CC9-BB17-4FED-B536-2A35796FCBBA

Fig. 21 .

Holotype. SR 92-18 - 08 ( Fig. 21 ). A forewing, collected by Carma Henry at Republic on September 25, 1992, in the collections of the SR.

Diagnosis. Distinguished from other species of Leptostigma by a combination of forewing dark in cell 1r+2rm. Tentatively diagnostic: R+M shorter than 1-M; 2r-rs joins cell 3rm distinctly near its middle; 2r-m joins M scarcely basal to 2m-cu; anal cells widely separated by fusion of 1A and 2A+3A, without crossvein.

Description. Holotype. Forewing length 14.2 mm, maximum width 4.1 mm, length/width 3.5; membrane lightly infuscate throughout except darker posterior to 1r-rs; pterostigma dark; R+M shorter than 1-M; maximum length of cell 2r shorter than 3r; 2r-rs shorter than Rs between 2r-m, 2r-rs, joining cell 3rm near its middle; 2r-m slightly curved, joins Rs notably basal to 2m-cu joining M, aligned with 2mcu; 1-M almost straight, sub-parallel to 1m-cu; 3r-m distinctly bowed toward base; cell 2mcu with M longer than 1m-cu; 2m-cu curved distinctly toward base; cu-a postfurcal; anal cells separated for distance greater than width of cell 2a, without crossvein between them.

Remarks. Differs from L. brevilatum n. sp., L. longiclava n. sp., L. longipallidum n. sp., L. longitenebricum n. sp. by its forewing dark posterior to 1r-rs and 2r-m joins M close to 2m-cu. Differs from L. proxivena n. sp. and L. fasciatum n. sp. in which the anal cells are not separated by a fusion of 1A with 2A+3A.

Material. Holotype.

Etymology. From the Latin ala, meaning wing, and macula, spot, referring to the forewing spot. The epithet is a noun in apposition.

Locality and age. Republic , Washington, United States of America, Tom Thumb Tuff Member of the Klondike Mountain Formation exposure A0307 ; mid-Ypresian .

Figures

FIGURE 21. Leptostigma alaemaculum holotype SR 92-18-08, forewing: A, photograph; B, drawing. Scale bar 5 mm.