Lathrobium mocsarskii, Bernhauer, 1913

Herman, Lee H., 2001, Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the End of the Second Millennium. I. Introduction, History, Biographical Sketches, and Omaliine Group, Bulletin of the American Museum of Natural History 2001 (265), pp. 1-650 : 1-650

mocsarskii Bernhauer, 1913 , see: Eusphalerum .

moczarskii Scheerpeltz, 1961c: 77 , 85 ( Lathrimaeum ; subgenus Lathrimaeum ; Type locality:

Süd-Spanien auf der Sierra Alfacar).

- DISTRIBUTION: Spain.

montanum Erichson, 1840 , see: Eusphalerum .

monticola Cameron, 1924b: 169 ( Lathrimaeum ; Type locality: Chakrata and Simla Districts,

7000 to 9000 feet).

— Cameron, 1930: 154 ( Lathrimaeum ; characters; India).

— Scheerpeltz, 1933: 1056 ( Lathrimaeum ; catalog).

— Scheerpeltz, 1961c: 78 ( Lathrimaeum ; subgenus Lathrimaeum ; characters; India).

- DISTRIBUTION: India.

mucronatum Stephens, 1834 , see: Eusphalerum .

nepalense Scheerpeltz, 1976 , see: Eusphalerum .

notabile Cameron, 1941 , see: Deinopteroloma .

nigerrimum Casey, 1894 , see: Eusphalerum .

nigricorne Stephens, 1834 , see: Proteinus .

nigriventre Stephens, 1834 , see: Eusphalerum .

nigriventre Motschulsky, 1860 , see: Eusphalerum .

nigropiceum Casey, 1894: 417 ( Lathrimaeum ; Type locality: California: Santa Cruz Co.).

— Bernhauer and Schubert, 1910: 65 ( Lathrimaeum ; catalog).

— Moore, 1966: 55 ( Anthobium ).

- DISTRIBUTION: USA.

nigrum Erichson, 1840 , see: Eusphalerum .

nigrum Cameron, 1924b: 170 ( Lathrimaeum ; [Note: Preoccupied as a secondary homonym];

Type locality: Simla Hills; Gahan, 7000 feet).

— Cameron, 1930: 152 ( Lathrimaeum ; characters; India).

— Scheerpeltz, 1933: 1056 ( Lathrimaeum ; catalog).

— Scheerpeltz, 1961c: 75 ( Lathrimaeum ; subgenus Lathrimaeum ; characters; Himalayan region).

- DISTRIBUTION: India.

nitidicolle Baudi, 1857 , see: Eusphalerum .

nitidifrons Luze, 1910 , see: Eusphalerum .

nitidum Stephens, 1834 , see: Proteinus .

nudum Luze, 1910 , see: Eusphalerum .

nuristanicum Scheerpeltz, 1961 , see: Eusphalerum .

obliquum Mulsant and Rey, 1861 , see: Eusphalerum .

oblitum Fairmaire and Laboulbène, 1856 , see: Eusphalerum .

oblongum Lacordaire, 1835 , see: Xylodromus .

obscurellum Luze, 1911 [Note: Nomen nudum], see: Eusphalerum .

obscurum Brisout, 1866 , see: Eusphalerum .

obsoletum Erichson, 1840 , see: Eusphalerum .

obtusicolle Fauvel, 1876 , see: Eusphalerum .

occiduum Casey, 1894: 420 ( Deliphrum ; Type locality: California: Siskiyou Co.).

— Hatch, 1957: 67 ( Deliphrum ; cited as occidum; characters; British Columbia ; Washington;

Oregon).

— Moore, 1966: 55 ( Anthobium ).

- DISTRIBUTION: Canada, USA.

octavii Fauvel, 1871 , see: Eusphalerum .

orientale Bernhauer, 1912 , see: Eusphalerum .

palligerum Kiesenwetter, 1847 , see: Eusphalerum .

parallelum Sharp, 1889 , see: Eusphalerum .

parnassicum Bernhauer, 1910 , see: Eusphalerum .

parvulum Scheerpeltz, 1976 , see: Eusphalerum .

pectorale Luze, 1910 , see: Eusphalerum .

petzi Bernhauer, 1910 , see: Eusphalerum .

petzianum Bernhauer, 1929 , see: Eusphalerum .

pfefferi Roubal, 1941 , see: Eusphalerum .

piceum Stephens, 1834 , see: Omalium .

picipenne Stephens, 1834 , see: Eusphalerum .

pictum Fauvel, 1878 , see: Deinopteroloma .

pilosellum Luze, 1910 , see: Eusphalerum .

pollens Sharp, 1889 , see: Eusphalerum .

pothos Mannerheim, 1843 , see: Eusphalerum .

pourtoyi Coiffait, 1978f: 273 ( Lathrimaeum ; Type locality: Hautes-Pyrénées, Lac Bleu ).

- DISTRIBUTION: France.

procerum Baudi, 1857 , see: Eusphalerum .

prolongatum Rottenberg, 1873 , see: Deliphrosoma .

pruinosum Fauvel, 1871 , see: Eusphalerum .

pseudaucupariae E. Strand, 1917 , see: Eusphalerum .

pulcherrimum Bernhauer, 1902 , see: Eusphalerum .

pumilio Rosenhauer, 1856 , see: Eusphalerum .

punctatum Casey, 1894 , see: Eusphalerum .

puncticeps Luze, 1910 , see: Eusphalerum .

puncticolle Gredler, 1863 , see: Eusphalerum .

punctulatum Mulsant and Rey, 1880 , see: Eusphalerum .

ragusae Bernhauer, 1923 , see: Eusphalerum .

rectanguloides Smetana, 1967 , see: Eusphalerum .

rectangulum Baudi, 1870 , see: Eusphalerum .

rectangulum Fauvel, 1871 , see: Eusphalerum .

reflexicolle Casey, 1894: 417 ( Lathrimaeum ; Type locality: British Columbia : Stickeen River

Cañon).

— Bernhauer and Schubert, 1910: 65 ( Lathrimaeum ; catalog).

— Hatch, 1957: 65 ( Anthobium ; characters; British Columbia ; Washington; Oregon; Idaho).

— Moore, 1966: 55 ( Anthobium ).

- DISTRIBUTION: Canada, USA.

reflexum Reitter, 1891b: 195 ( Lathrimaeum ; Type locality: bei Taschkent).

— Luze, 1905a: 68 ( Lathrimaeum ; subgenus Prionothrax ; characters; Turkestan; Buchara).

— Scheerpeltz, 1961c: 85 ( Lathrimaeum ; subgenus Prionothorax ; characters; Turkestan;

Buchara).

— Coiffait, 1970d: 144 ( Lathrimaeum ; subgenus Prionothorax ; Uzbekistan).

— Tikhomirova, 1973a: 139 ( Lathrimaeum ; subgenus Lathrimaeum ; checklist; Central Asia).

- DISTRIBUTION: Uzbekistan.

reitteri Bernhauer, 1935 , see: Eusphalerum .

retowskii Bernhauer, 1914 , see: Eusphalerum .

rhododendri Baudi, 1848 , see: Eusphalerum .

rhododendri Motschulsky, 1857 , see: Eusphalerum .

roubali Luze, 1911: 293 ( Lathrimaeum ; Type locality: Kaukasus occid., Krasnaja Poljana).

— Scheerpeltz, 1933: 1056 ( Lathrimaeum ; catalog).

— Scheerpeltz, 1961c: 81 ( Lathrimaeum ; subgenus Lathrimaeum ; characters; Caucasus).

- DISTRIBUTION: Russia.

ruffoi Scheerpeltz, 1956 , see: Eusphalerum .

ruficolle Stephens, 1834 , see: atrocephalum Gyllenhal, 1827 .

ruficorne Stephens, 1834: 338 ( Anthobium ; [Note: Nomen dubium]; Type locality: Not cited,

but presumably Britain).

— Stephens, 1839: 425 ( Anthobium ; characters; England).

- DISTRIBUTION: England.

rufipenne Gerhardt, 1910 [Note: Nomen nudum], see: Eusphalerum .

rufiventre Cameron, 1930 , see: Eusphalerum .

rufoscutellatum Eppelsheim, 1881 , see: Eusphalerum .

rufotestaceum Motschulsky, 1860 , see: Eusphalernum.

rufulum Luze, 1910 [Note: Nomen nudum], see: Eusphalerum .

rugosum Champion, 1925: 103 ( Lathrimaeum ; Type locality: Upper Gumti Valley , W. Almora

Division, alt. 6000 ft.).

— Cameron, 1930: 154 ( Lathrimaeum ; characters; India).

— Scheerpeltz, 1933: 1056 ( Lathrimaeum ; catalog).

— Scheerpeltz, 1961c: 85 ( Lathrimaeum ; subgenus Lathrimaeum ; characters; India).

— Coiffait, 1977: 244 ( Lathrimaeum ; Nepal).

- DISTRIBUTION: India, Nepal.

rugulosum Mäklin, 1853 , see: Eusphalerum .

sahlbergi Luze, 1910 , see: Eusphalerum .

sardoum Scheerpeltz, 1961, see: atrocephalum Gyllenhal, 1827 .

sareptanum Eppelsheim, 1878 , see: Eusphalerum .

satanas Luze, 1911 [Note: Nomen nudum], see: Eusphalerum .

schatzmayri C. Koch, 1938 , see: Eusphalerum .

scribae Schaufuss, 1862 , see: Eusphalerum .

scutellare Erichson, 1840 , see: Eusphalerum .

sibiricum Luze, 1910 , see: Eusphalerum .

signatum Märkel, 1857 , see: Eusphalerum .

sikkimi Fauvel, 1904 , see: Eusphalerum .

silesiacum Letzner, 1868 , see: Eusphalerum .

sinuatocolle Lokay, 1919 , see: Eusphalerum .

sinuosum Hatch, 1957: 66 ( Anthobium ; Type locality: B. C. [ British Columbia ]: Creston. Also cited from Washington, Oregon, Idaho). sociale Cameron, 1941: 59 ( Lathrimaeum ; subgenus Prionothorax ; Type locality: Kashmir:

Gulmarg, alt. 8000– 10,000 feet).

- DISTRIBUTION: India.

solitare Sharp, 1874 , see: Eusphalerum .

sorbicola Y. Kangas, 1941 , see: Eusphalerum .

sordidulum Kraatz, 1857 , see: Eusphalerum .

sordidum Erichson, 1840: 871 ( Lathrimaeum ; Type locality: America septentrionalis ).

— LeConte, 1850: 221 ( Lathrimaeum ; Lake Superior).

— Provancher, 1877: 260 ( Lathrimaeum ; characters; Quebec).

— Fauvel, 1878d: 235 [= 1878e: 71] ( Lathrimaeum ; characters; North America ).

— Bernhauer and Schubert, 1910: 65 ( Lathrimaeum ; catalog).

- DISTRIBUTION: Canada, USA.

sparsum Fauvel, 1876 , see: Eusphalerum .

spretum Casey, 1894: 418 ( Lathrimaeum ; Type locality: California: Siskiyou Co.).

— Bernhauer and Schubert, 1910: 65 ( Lathrimaeum ; catalog).

— Moore, 1966: 55 ( Anthobium ).

- DISTRIBUTION: USA.

starcki Reitter, 1890 , see: Eusphalerum .

stramineum Kraatz, 1857 , see: Eusphalerum .

stussineri Bernhauer, 1909 , see: Eusphalerum .

subangulatum Casey, 1894 , see: Eusphalerum .

subcostatum Mäklin, 1852 , see: Deinopteroloma .

subjectum Mulsant and Rey, 1880 , see: Eusphalerum .

subsulcatum Stephens, 1834 , see: Proteinus .

sulculum Stephens, 1834 , see: Acrolocha .

tempestivum Erichson, 1840 , see: Eusphalerum .

tenebaumi Bernhauer, 1932, see: Eusphalerum .

tenue Eppelsheim, 1881: 507 ( Lathrimaeum ; Type locality: Suramgebirge).

— Luze, 1905a: 60 ( Lathrimaeum ; characters; Caucasus).

— Bernhauer and Schubert, 1910: 65 ( Lathrimaeum ; catalog).

— Scheerpeltz, 1961c: 80 ( Lathrimaeum ; subgenus Lathrimaeum ; characters; Caucasus).

— Bordoni, 1986c: 43 ( Lathrimaeum ; characters).

— Tikhomirova, 1973a: 139 ( Lathrimaeum ; subgenus Lathrimaeum ; checklist; Caucasus).

- DISTRIBUTION: Georgia.

tibiale Casey, 1894 , see: Eusphalerum .

tibiale Luze, 1910 , see: Eusphalerum .

transcaucasicum Bernhauer, 1902 , see: Eusphalerum .

triviale Erichson, 1839 , see: Eusphalerum .

umbellatarum Kiesenwetter, 1850 , see: Eusphalerum .

umbricolor Roubal, 1915 , see: Eusphalerum .

uncipes C. Koch, 1938 , see: Eusphalerum .

unicolor Marsham, 1802: 127 ( Silpha ; Type locality: Britain).

— Stephens, 1834: 343 ( Anthobium ; characters; England).

— Stephens, 1839: 426 ( Lathrimaeum ; characters; England).

— Stephens, 1839: 427 ( Olophrum ; [Note: Misidentified as assimile according to Waterhouse,

1858: 31]; characters; England).

— Hardy, 1851: 58 ( Lathrimaeum ; synonym of atrocephalum ).

— Fauvel, 1876a: 52 [= 1876: 237] ( Lathrimaeum ; Spain).

— Fowler, 1888: 406 ( Lathrimaeum ; characters; Britain).

— Ganglbauer, 1895: 722 ( Lathrimaeum ; characters; middle Europe; Portugal).

— Luze, 1905a: 61 ( Lathrimaeum ; characters; central Europe; Spain; Portugal).

— Reitter, 1909: 188 ( Lathrimaeum ; characters; Germany).

— Bernhauer and Schubert, 1910: 65 ( Lathrimaeum ; catalog).

— Johansen, 1914: 596 ( Lathrimaeum ; characters; Denmark).

— Portevin, 1929: 441 ( Lathrimaeum ; characters; France).

— Scheerpeltz, 1933: 1056 ( Lathrimaeum ; catalog).

— Palm, 1948: 101 ( Lathrimaeum ; characters; Sweden; Denmark; Norway).

— Scheerpeltz, 1961c: 81 ( Lathrimaeum ; subgenus Lathrimaeum ; characters; Europe an Mediterranean region).

— Horion, 1963: 106 ( Lathrimaeum ; Britain; Denmark; Norway; Sweden; Holland; Belgium; France; Spain; Portugal; Romania?; Slovakia?; Germany; Austria).

— Lohse, 1964: 52 ( Lathrimaeum ; characters; central Europe).

— Kasule, 1968: 134 ( Lathrimaeum ; larval characters).

— Kasule, 1968a: 73 ( Lathrimaeum ; life history).

— Steel, 1970a: 18 ( Anthobium ; biological notes).

— Pope, 1977: 22 ( Anthobium ; Britain).

— Coiffait, 1978f: 275 ( Lathrimaeum ; characters; France).

— Burakowski, Mroczkowski, and Stefańska, 1979: 57 ( Anthobium ; Poland).

— Hammond, 1980: 136 ( Anthobium ; Ireland).

— Tóth, 1982: 80 ( Lathrimaeum ; characters).

— Lucht, 1987: 88 ( Lathrimaeum ; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

— Dettner and Reissenweber, 1991 ( Lathrimaeum ; defensive secretions: chemistry and systematics).

— Welch, 1993: 227 ( Anthobium ; ovariole number and ovary structure).

— Boháč, 1993: 41 ( Anthobium ; Slovakia?).

— Owen, 1999b: 61 ( Anthobium ; captured underground at roots of old trees; Britain).

— Owen, 2000: 251 ( Anthobium ; pitfall trap collections from underground at roots of trees; Britain).

— Assing, 2001: 75 ( Anthobium ; collected from xerothermous site; Germany).

- DISTRIBUTION: Europe.

assimile Stephens, 1834: 342 ( Anthobium ; Type locality: London ; Suffolk).

— G. Waterhouse, 1858: 31 ( Lathrimaeum ; synonym of unicolor ).

— Bernhauer and Schubert, 1910: 65 ( Lathrimaeum ; synonym of unicolor ).

luteum Erichson, 1840: 869 ( Lathrimaeum ; Type locality: Lusitania).

— Fairmaire and Laboulbène, 1856: 636 ( Lathrimaeum ; characters; France).

— Redtenbacher, 1857: 249 ( Olophrum ; characters; Germany).

— Kraatz, 1857d: 945 ( Lathrimaeum ; characters; Germany).

— G. Waterhouse, 1858: 31 ( Lathrimaeum ; synonym of unicolor ).

— Thomson, 1867: 315 ( Lathrimaeum ; characters; Scandinavia).

— Thomson, 1867a: 48 ( Lathrimaeum ; characters; Scandinavia).

— Fauvel, 1871a: 119 [= 1872: 119] ( Lathrimaeum ; synonym of unicolor ).

— Redtenbacher, 1874: 269 ( Lathrimaeum ; characters; Germany).

— Seidlitz, 1875: 239 ( Lathrimaeum ; characters; Baltic region).

— Mulsant and Rey, 1880: 123 ( Lathrimaeum ; characters; France).

— Fowler, 1888: 406 ( Lathrimaeum ; synonym of unicolor ).

— Ganglbauer, 1895: 722 ( Lathrimaeum ; synonym of unicolor ).

— Bernhauer and Schubert, 1910: 65 ( Lathrimaeum ; synonym of unicolor ).

— Scheerpeltz, 1933: 1056 ( Lathrimaeum ; synonym of unicolor ).

diabolicum Luze, 1905 a: 61 ( Lathrimaeum ; variety of unicolor ; Type locality: Not cited).

— Bernhauer and Schubert, 1910: 65 ( Lathrimaeum ; variety of unicolor ).

— Scheerpeltz, 1961c: 81 ( Lathrimaeum ; subgenus Lathrimaeum ; aberration of unicolor ; characurgelense Fagel, 1968: 122 ( Lathrimaeum ; Type locality: Espagne: Catalogne, Seo de Urgel,

contreforts de la Sierra del Cadi, 1000 m).

- DISTRIBUTION: Spain.

ustulatum Fairmaire and Laboulbène, 1856 , see: Eusphalerum .

viertli Ganglbauer, 1895 , see: Eusphalerum .

zolotarevi Reitter, 1909 , see: Eusphalerum .

Anthophagus

[36 species (7 doubtful); Palaearctic and Nearctic Regions]

Anthophagus Gravenhorst, 1802: 120 (species included: caraboides , abbreviatus , testaceus , armiger , obscurus , plagiatus , dichrous , alpinus ; [Note: Without action by the Commission the name of this genus must be Lesteva and the generic name for the species currently in Lesteva must be Tevales . See Herman, 2001: 7]). TYPE SPECIES: Staphylinus alpinus Fabricius , fixed by subsequent designation by Thomson, 1859: 48.

— Latreille, 1802: 129 (cited as antophagus; synonym of Lesteva ).

— Latreille, 1804: 286, 366 (cited as synonym of Lesteva ).

— Latreille, 1806: 296 (synonym of Lesteva ).

— Gravenhorst, 1806: 220 (characters).

— Gyllenhal, 1810: 191 (characters).

— Leach, 1815: 92 (characters).

— Billberg, 1820: 15 (characters; list of some species).

— Dejean, 1821: 25 (cited as Antophagus ; list of species in collection).

— C. Sahlberg, 1827: 273 (characters).

— Zetterstedt, 1828: 44 (characters).

— Latreille, 1829: 439 (synonym of Lesteva ).

— Dejean, 1833: 68 (list of species).

— Gistel, 1834: 9 (list of species).

— Say, 1834: 463 (characters).

— Zetterstedt, 1838: 48 (characters).

— Erichson, 1839a: 614 (characters).

— Heer, 1839: 194 (characters).

— Erichson, 1840: 847 (characters).

— Laporte, 1840: 192 (characters).

— Kiesenwetter, 1845: 21 (notes).

— Redtenbacher, 1849: 745 (characters).

— Schaum, 1852: 29 (list of species; Europe).

— Lacordaire, 1854: 135 (characters; notes; list of species).

— Fairmaire and Laboulbène, 1856: 625 (characters).

— Gistel, 1856: 267, 389 (list of species).

— Jacquelin du Val, 1857: 69 (characters).

— Redtenbacher, 1857: 241 (characters; key to Austrian species).

— Kraatz, 1857d: 912 (characters).

— Thomson, 1858: 37 (characters).

— G. Waterhouse, 1858: 30 (catalog of British species).

— Thomson, 1859: 48 (characters; type species: alpinus ).

— Schaum, 1859: 31 (catalog; European species).

— LeConte, 1861a: 70 (characters).

— Thomson, 1861: 178 (characters).

— Gredler, 1863: 119 (list of species from Tirol).

— LeConte, 1863: 26 (list of North American species).

— Wencker and Silbermann, 1866: 35 (list of species; collecting notes; France).

— Gemminger and Harold, 1868: 657 (catalog).

— Jacquelin du Val, 1868: 79 (catalog; European species).

— Fauvel, 1871a: 137 [= 1872: 111] (characters; key to species).

— Redtenbacher, 1874: 263 (characters; key to Austrian species).

— Fauvel, 1875a:VII [= 1875b: 209] (catalog).

— Provancher, 1877: 259 (characters).

— Mulsant and Rey, 1880: 8 (characters; key to French species).

— Heyden, 1880: 81 (list of species of Siberian region).

— Duvivier, 1883: 193 (catalog).

— Fowler, 1888: 398 (notes; key to British species).

— Seidlitz, 1891: 348 (key to some European species).

— Seidlitz, 1891a: 371 (key to some European species).

— Heyden, Reitter, and Weise, 1891: 117 (list of species of Europe and Caucasus).

— Ganglbauer, 1895: 704 (characters; key to middle European species).

— Luze, 1902d: 505 (characters; key to Palaearctic species).

— Heyden, Reitter, and Weise, 1906: 136 (list of species of Europe and Caucasus).

— Sainte-Claire Deville, 1907: 50 (characters; key to and annotated list of species of Seine Basin, France).

— Reitter, 1909: 182 (characters; key to German species).

— Bernhauer and Schubert, 1910: 78 (world catalog; 29 species).

— Johansen, 1914: 585 (characters).

— Winkler, 1925: 335 (catalog for Palaearctic region).

— Porta, 1926: 31 (characters; key to species of Italy).

— Portevin, 1929: 433 (characters; key to French species).

— Roubal, 1930: 317 (catalog; Slovakia).

— Joy, 1932: 98 (key to British species).

— Scheerpeltz, 1933: 1067 (world catalog supplement).

— C. Koch, 1933a (discussion; classification).

— C. Koch, 1934a (revision; key to species; redescription of species; discussion; distribution).

— Scheerpeltz, 1937b: 199 (list of Bulgarian species).

— Kloet and Hincks, 1945: 161 (list of British species).

— Palm, 1948: 123 (characters; key to species of Sweden).

— Tottenham, 1949: 358, 408 (type species: alpinus Paykull ; list of British species).

— Blackwelder, 1952: 56 (type species).

— Tottenham, 1954: 34 (characters; key to British species).

— Smetana, 1959a: 196 (list of species collected in Albania).

— Horion, 1963: 138 (list of central European species).

— Székessy, 1963: 22 (characters).

— Lohse, 1964: 63 (characters; key to central European species).

— Moore, 1966: 48, 51 (characters; notes).

— Pototskaia, 1967: 20 (larval characters).

— Scheerpeltz, 1968a: 17 (catalog of Austrian species).

— Steel, 1970a: 21 (larval characters; biological notes).

— Tikhomirova, 1973a: 141 (checklist of species of USSR).

— Moore and Legner, 1975: 183 (catalog; one unnamed species from California).

— Pope, 1977: 23 (list of British species).

— Topp, 1978: 309 (larval characters).

— Burakowski, Mroczkowski, and Stefańska, 1979: 71 (catalog; Poland).

— Moore and Legner, 1979: 178 (characters; notes).

— Muona, 1979: 18 (list of Scandinavian species).

— Uhlig, Vogel, and Sieber, 1980: 240 (some species collected in Germany).

— Uhlig and Vogler, 1981: 88 (list of some species collected in Germany; habitat and collecting notes).

— Tóth, 1982: 89 (characters; habitus; key to species of Hungary).

— Ádám, 1985: 251 (list of species from southeast Hungary).

— Muona and Viramo, 1986: 16 (list of species of northeastern Finland).

— Segers, 1986: 23 (checklist of species; Belgium).

— Zanetti, 1987: 387 (characters; habitus; key to species of Italy).

— Angelini, 1991: 190 (list of species from high elevations of Calabria, Italy).

— Silfverberg, 1992: 22 (list of species of Fennoscandia, Denmark, and the Baltic States).

— Siitonen, 1993: 230 (list of Finnish species).

— Boháč, 1993: 41 (list of species of Czech Republic and Slovakia).

— Zanetti, 1995: 9 (list of species of Italy).

— M. Hansen, 1996: 95 (list of Danish species).

— M. Hansen, Palm, Pedersen, and Runge, 1998: 70 (list of species collected in 1997; Denmark).

— Newton, Thayer, Ashe, and Chandler, 2000: 339 (1 undescribed Nearctic species; notes; characters in key).

Phaganthus Mulsant and Rey, 1880: 12 , 42 (subgenus of Anthophagus ; species included: scutellaris , testaceus , praeustus , rotundicollis ). TYPE SPECIES: Anthophagus testaceus Gravenhorst , fixed by subsequent designation by Blackwelder, 1952: 299.

— Ganglbauer, 1895: 704 (subgenus of Anthophagus ).

— Bernhauer and Schubert, 1910: 78 (subgenus of Anthophagus ).

— Joy, 1932: 99 (key to British species).

— Scheerpeltz, 1933: 1067 (subgenus of Anthophagus ).

— Tottenham, 1945: 70 (subgenus of Anthophagus ; cited as Plaganthus [Note: Corrected to Phaganthus by Tottenham, 1949c: 41]; type species: Staphylinus caraboides Linné, 1785 = Anthophagus testaceus Gravenhorst, 1802 [Note: caraboides was not an originally included species]).

— Kloet and Hincks, 1945: 161 (list of British species).

— Tottenham, 1949: 358, 408 (subgenus of Anthophagus ; type species: caraboides Linné = testaceus Gravenhorst ; list of British species).

— Horion, 1963: 138 (cited as Phagantus; subgenus of Anthophagus ; list of central European species).

— Lohse, 1964: 64 (subgenus of Anthophagus ; characters; key to central European species).

— Kasule, 1968: 133, 136 (larval characters).

— Zanetti, 1987: 387 (subgenus of Anthophagus ; characters; key to species of Italy).

Dimorphoschelus Blackwelder, 1952: 126 . TYPE SPECIES: Anthophagus alpestris Heer , fixed by subsequent designation by Blackwelder, 1952: 126.

— C. Koch, 1933a: 140, 142 (characters; subgenus of Anthophagus ; species included: angusticollis , fulvus , gracilis , obscuriceps , fascifer , apfelbecki , schneideri , omalinus , arpedinus , arrowi , melanocephalus , fenestrellanus , hummleri , schatzmayri , dinaricus , transversus , alpestris , austriacus , decoratus ; [Note: This is the original description for Dimorphoschelus but the name was unavailable because the author failed to designated a type species (Article 13.3). A type species was designated by Blackwelder, 1952]).

— Zanetti, 1987: 388, 413 (subgenus of Anthophagus ; characters; habitus; key to species of Italy).

abbreviatus Fabricius, 1779 , see: angusticollis Mannerheim, 1830 .

advena Gistel, 1857: 89 ( Anthophagus ; [Note: Nomen dubium]; Type locality: Alpes Longobardiae).

- DISTRIBUTION:?.

aemulus Rosenhauer, 1847 , see: Hygrogeus .

aeneicollis Fauvel, 1873b: 120 [= 1873c: 13] ( Anthophagus ; Type locality: Gap; [Note: See lectotype designation by Zanetti, 1986]).

— Mulsant and Rey, 1880: 17 ( Anthophagus ; characters; France).

— Ganglbauer, 1895: 706 ( Anthophagus ; characters; western Alps).

— Luze, 1902d: 514 ( Anthophagus ; characters; western Alps).

— Bernhauer and Schubert, 1910: 78 ( Anthophagus ; catalog).

— Portevin, 1929: 434 ( Anthophagus ; characters; France).

— Scheerpeltz, 1933: 1068 ( Anthophagus ; catalog).

— C. Koch, 1933a: 142, 143 ( Anthophagus ; subgenus Anthophagus ; figure).

— C. Koch, 1934a: 181 ( Anthophagus ; subgenus Anthophagus ; characters; France; Switzerland; Austria). — Horion, 1963: 143 ( Anthophagus ; subgenus Anthophagus ; France; Italy; Switzerland; Germany; Austria). — Lohse, 1964: 65 ( Anthophagus ; subgenus Anthophagus ; characters; central Europe). — Jarrige, 1968b: 76 ( Anthophagus ; France). — Zanetti, 1986: 97 ( Anthophagus ; lectotype designation). — Zanetti, 1987: 403 ( Anthophagus ; subgenus Anthophagus ; characters; Italy). — Lucht, 1987: 90 ( Anthophagus ; checklist; Germany; Austria). — Zanetti, 1995: 9 ( Anthophagus ; Italy). - DISTRIBUTION: Germany, France, Switzerland, Austria, Italy.

albanicus Bernhauer, 1926 , see: alpinus Paykull, 1790 .

alpestris Heer, 1839: 196 ( Anthophagus ; Type locality: Alp Manigorio Ct. Tessin). — Redtenbacher, 1849: 745 ( Anthophagus ; synonym of austriacus ). — Kraatz, 1857d: 918 ( Anthophagus ; synonym of austriacus ). — Fauvel, 1871a: 141 [= 1872: 115] ( Anthophagus ; characters; France; Germany; Switzerland; Italy; Austria). — Mulsant and Rey, 1880: 37 ( Anthophagus ; characters; France). — Ganglbauer, 1895: 708 ( Anthophagus ; characters; Alps; Sudeten; Carpathians). — Luze, 1902d: 512 ( Anthophagus ; characters; Alps; Sudeten; Carpathians; Jura). — Reitter, 1909: 183 ( Anthophagus ; characters; Germany). — Bernhauer and Schubert, 1910: 78 ( Anthophagus ; catalog). — Portevin, 1929: 434 ( Anthophagus ; characters; France). — Scheerpeltz, 1933: 1068 ( Anthophagus ; catalog). — C. Koch, 1933a: 142, 143 ( Anthophagus ; subgenus Dimorphoschelus ; figure). — C. Koch, 1934a: 234 ( Anthophagus ; subgenus Dimorphoschelus ; characters; Czechoslovakia; Alps; Hungary; Bosnia; Italy; France). — Horion, 1963: 149 ( Anthophagus ; subgenus Dimorphoschelus ; Czech Republic; Slovakia; Romania; Bulgaria; Bosnia Hercegovina; Serbia; Albania; Slovenia; Austria; Italy; France; Germany; Switzerland). — Lohse, 1964: 67 ( Anthophagus ; subgenus Dimorphoschelus ; characters; central Europe). — Tóth, 1982: 95 ( Anthophagus ; characters; Hungary). — Zanetti, 1984: 79 ( Anthophagus ; Bulgaria). — Zanetti, 1987: 420 ( Anthophagus ; subgenus Dimorphoschelus ; characters; Italy). — Lucht, 1987: 90 ( Anthophagus ; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland). — Schülke and Uhlig, 1988: 3 ( Anthophagus ; Germany). — Boháč, 1993: 42 ( Anthophagus ; subgenus Anthophagus ; [Note: Incorrect subgeneric assignment; A. alpestris is the type species of subgenus Dimorphoschelus ]; subspecies alpestris ; Slovakia; Czech Republic). — Zanetti, 1995: 9 ( Anthophagus ; Italy). - DISTRIBUTION: France, Germany, Switzerland, Italy, Austria, Hungary, Czech Republic, Slovakia, Bulgaria, Bosnia Hercegovina.

austriacus Erichson, 1840: 849 ( Anthophagus ; Type locality: Austria; Carinthia). — Kiesenwetter, 1845: 23 ( Anthophagus ; notes). — Kiesenwetter, 1847a: 55 ( Anthophagus ; characters; Germany; Switzerland). — Märkel and Kiesenwetter, 1848a: 327 ( Anthophagus ; Carpathian Alps). — Redtenbacher, 1849: 745 ( Anthophagus ; characters; Austria). — Fairmaire and Laboulbène, 1856: 628 ( Anthophagus ; characters; France). — Redtenbacher, 1857: 242 ( Anthophagus ; characters; Austria). — Kraatz, 1857d: 918 ( Anthophagus ; characters; Austria). — Fauvel, 1865a: 314 [= 1865b: 64] ( Anthophagus ; habitat; France). — Fauvel, 1871a: 141 [= 1872: 115] ( Anthophagus ; synonym of alpestris ). — Redtenbacher, 1874: 264 ( Anthophagus ; characters; Austria). — Mulsant and Rey, 1880: 38 ( Anthophagus ; synonym of alpestris ). — Ganglbauer, 1895: 708 ( Anthophagus ; synonym of alpestris ). — Bernhauer and Schubert, 1910: 79 ( Anthophagus ; synonym of alpestris ). — Scheerpeltz, 1933: 1068 ( Anthophagus ; synonym of alpestris ). — C. Koch, 1933a: 142 ( Anthophagus ; synonym of alpestris ). — Zanetti, 1987: 420 ( Anthophagus ; synonym of alpestris ).

decoratus C. Koch, 1933: 30 ( Anthophagus ; named for an aberrant form of transversus Motschulsky and cited as alpestris decoratus ; [Note: Nomen nudum (Articles 1.3.4, 45.5); although Koch did not explicitly state that decoratus was an aberration he did say that is was named for an aberrant form of transversus and in later articles cited it as an aberration]; locality cited: Alpes de la Carniole). — C. Koch, 1933a: 142 ( Anthophagus ; aberration of alpestris ; name is replacement for transversus sensu auctorum). — C. Koch, 1934a: 234 ( Anthophagus ; subgenus Dimorphoschelus ; aberration of alpestris ). — Tóth, 1982: 95 ( Anthophagus ; aberration of alpestris ; characters). — Zanetti, 1987: 420 ( Anthophagus ; synonym of alpestris ).

kochi Bernhauer, 1935 a: 39 ( Anthophagus ; Type locality: Italien: Mte. Arazecca). — Zanetti, 1986: 96 ( Anthophagus ; subspecies of alpestris ). — Zanetti, 1987: 420 ( Anthophagus ; subgenus Dimorphoschelus ; subspecies of alpestris ; characters; Italy). — Zanetti, 1995: 9 ( Anthophagus ; subspecies of alpestris ; Italy).

alpinus Paykull, 1790: 134 ( Staphylinus ; [Note: This name is older than alpinus Fabricius , however, Paykull cited alpinus again in 1800 but attributed it to Fabricius so the Fabricius and Paykull species may be the same]; Type locality: Dalicarlia, Lapponia). — Turton, 1802: 512 ( Staphylinus ; characters; Lapland). — Latreille, 1806: 297 ( Lesteva ; characters). — Fauvel, 1871a: 147 [= 1872: 121] ( Anthophagus ; characters; France; Scandinavia; Britain; Germany; Switzerland; Italy; Austria). — J. Sahlberg, 1876: 206 ( Anthophagus ; Finland). — Fowler, 1888: 399 ( Anthophagus ; characters; Britain). — Pope, 1977: 23 ( Anthophagus ; Britain). — Lucht, 1987: 90 ( Anthophagus ; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France). — Herman, 2001: 19 ( Anthophagus ; synonymy). - DISTRIBUTION: Europe, Russia.

alpinus Fabricius, 1793: 526 ( Staphylinus ; [preoccupied]; Type locality: Lapponia). — Olivier, 1795:(42): 32 ( Staphylinus ; characters). — Paykull, 1800: 387 ( Staphylinus ; [Note: Paykull attributed the species in this reference to Fabricius and did not mention alpinus Paykull, 1790 ]; characters; Sweden). — Fabricius, 1801: 598 ( Staphylinus ; characters; Lapland). — Gravenhorst, 1802: 188 ( Anthophagus ; characters). — Latreille, 1804: 368 ( Lesteva ; characters; Lapland). — Gravenhorst, 1806: 220 ( Anthophagus ; characters). — Latreille, 1806: 297 ( Lesteva ; characters). — Gyllenhal, 1810: 194 ( Anthophagus ; characters; Sweden). — Latreille, 1825a: 216 ( Lesteva ; characters; Lappland). — Gyllenhal, 1827: 460 ( Anthophagus ; characters of variety). — Zetterstedt, 1828: 46 ( Anthophagus ; characters; Lapland). — Curtis, 1830: pl. 303 ( Lesteva ; habitat; Britain). — Curtis, 1829: 29 ( Lesteva ; catalog; Britain). — Mannerheim, 1830: 56 ( Lesteva ; Finland; Sweden; Lapland). — Mannerheim, 1831: 470 ( Lesteva ; Finland; Sweden; Lapland). — Runde, 1835: 24 ( Lesteva ; characters; Germany). — Zetterstedt, 1838: 48 ( Anthophagus ; characters; habitat; Lapland). — Heer, 1839: 196 ( Anthophagus ; characters; Switzerland).

— Stephens, 1839: 431 ( Lesteva ; characters; England; Scotland). — Erichson, 1840: 848 ( Anthophagus ; characters; Sweden; Alpibus Tyrolensibus; Switzerland; Silesia). — Kiesenwetter, 1845: 22 ( Anthophagus ; notes). — Kiesenwetter, 1847a: 53 ( Anthophagus ; characters; Europe). — Märkel and Kiesenwetter, 1848a: 326 ( Anthophagus ; notes; Carpathian Alps). — Hochhuth, 1849: 201 ( Anthophagus ; characters; Caucasus; Akhaltzik). — Redtenbacher, 1849: 745 ( Anthophagus ; characters; Austria). — Kiesenwetter, 1851a: 433 ( Anthophagus ; Pyrenees). — Hardy, 1851: 54 ( Anthophagus ; notes; England). — Fairmaire and Laboulbène, 1856: 626 ( Anthophagus ; characters; France). — Redtenbacher, 1857: 242 ( Anthophagus ; characters; Austria). — Kraatz, 1857d: 918 ( Anthophagus ; characters; Germany). — Thomson, 1859: 48 (type species of Anthophagus ). — Thomson, 1861: 179 ( Anthophagus ; characters; Scandinavia). — Fauvel, 1865a: 314 [= 1865b: 64] ( Anthophagus ; habitat; France). — Redtenbacher, 1874: 264 ( Anthophagus ; characters; Austria). — Seidlitz, 1875: 242 ( Anthophagus ; characters; Baltic region). — Mulsant and Rey, 1880: 19 ( Anthophagus ; characters; France). — Ganglbauer, 1895: 706 ( Anthophagus ; characters; north and middle Europe; Italy; Bosnia). — Luze, 1902d: 518 ( Anthophagus ; characters; northern and central Europe; northern Italy). — Reitter, 1909: 182 ( Anthophagus ; characters; Germany). — Bernhauer and Schubert, 1910: 79 ( Anthophagus ; catalog). — Portevin, 1929: 434 ( Anthophagus ; characters; France). — Scheerpeltz, 1933: 1068 ( Anthophagus ; catalog). — C. Koch, 1933a: 142 ( Anthophagus ; subgenus Anthophagus ). — C. Koch, 1934a: 196 ( Anthophagus ; subgenus Anthophagus ; characters; Lapland; Russia; Norway; Sweden; Sudeten; Tatra; Hungary; Alps; Italy; England; Scotland; Ireland; Switzerland; France). — Palm, 1948: 124 ( Anthophagus ; characters; Sweden; Norway; Finland). — Horion, 1963: 144 ( Anthophagus ; subgenus Anthophagus ; Britain; Norway; Sweden; Finland; Russia; Czech Republic; Slovakia; Romania; Italy; France; Austria; Germany). — Lohse, 1964: 65 ( Anthophagus ; subgenus Anthophagus ; characters; central Europe). — Hammond, 1980: 136 ( Anthophagus ; Ireland). — Tóth, 1982: 93 ( Anthophagus ; characters; Hungary). — Zanetti, 1987: 407 ( Anthophagus ; subgenus Anthophagus ; characters; Italy). — Boháč, 1993: 41 ( Anthophagus ; subgenus Dimorphoschelus ; [Note: Incorrect subgeneric assignment; A. alpinus is the type species of subgenus Anthophagus ]; Slovakia; Czech Republic). — Zanetti, 1995: 9 ( Anthophagus ; Italy). — Herman, 2001: 19 ( Anthophagus ; synonym of alpinus ).

mandibularis Gyllenhal, 1827: 460 ( Anthophagus ; Type locality: Lapponia). — C. Sahlberg, 1827: 275 ( Anthophagus ; characters; Lapland). — Erichson, 1840: 848 ( Anthophagus ; synonym of alpinus ). — Kraatz, 1857d: 919 ( Anthophagus ; synonym of alpinus ). — Thomson, 1861: 179 ( Anthophagus ; synonym of alpinus ). — J. Sahlberg, 1876: 206 ( Anthophagus ; synonym of alpinus ). — Mulsant and Rey, 1880: 20 ( Anthophagus ; synonym of alpinus ). — Ganglbauer, 1895: 706 ( Anthophagus ; synonym of alpinus ). — Luze, 1902d: 518 ( Anthophagus ; synonym of alpinus ). — Bernhauer and Schubert, 1910: 79 ( Anthophagus ; synonym of alpinus ). — C. Koch, 1933a: 142 ( Anthophagus ; synonym of alpinus ). — Pope, 1977: 23 ( Anthophagus ; synonym of alpinus ).

lapponicus C. Sahlberg, 1827: 275 ( Anthophagus ; Type locality: Lapponia). — Mannerheim, 1830: 56 ( Lesteva ; [Note: Mannerheim cited lapponica as his but at the same time attributed it to C. Sahlberg]; Lapland). — Zetterstedt, 1838: 47 ( Anthophagus ; characters; Lapland).

— Erichson, 1840: 849 ( Anthophagus ; synonym of alpinus ).

— Redtenbacher, 1849: 745 ( Anthophagus ; synonym of alpinus ).

— Kraatz, 1857d: 919 ( Anthophagus ; synonym of alpinus ).

— J. Sahlberg, 1876: 206 ( Anthophagus ; synonym of alpinus ).

— Ganglbauer, 1895: 706 ( Anthophagus ; synonym of alpinus ).

— Luze, 1902d: 518 ( Anthophagus ; synonym of alpinus ).

— Bernhauer and Schubert, 1910: 79 ( Anthophagus ; synonym of alpinus ).

— C. Koch, 1933a: 142 ( Anthophagus ; synonym of alpinus ).

maxillosus Zetterstedt, 1828: 45 ( Anthophagus ; Type locality: insula Reenoe: Finkrog. Lyngentind. Nordlandiae).

— Zetterstedt, 1838: 48 ( Anthophagus ; characters; habitat; Lapland).

— Erichson, 1840: 848 ( Anthophagus ; synonym of alpinus ).

— Kraatz, 1857d: 919 ( Anthophagus ; synonym of alpinus ).

— Thomson, 1861: 179 ( Anthophagus ; synonym of alpinus ).

— Ganglbauer, 1895: 706 ( Anthophagus ; synonym of alpinus ).

— Luze, 1902d: 518 ( Anthophagus ; synonym of alpinus ).

— Bernhauer and Schubert, 1910: 79 ( Anthophagus ; synonym of alpinus ).

flavipennis Lacordaire, 1835: 480 ( Lesteva ; Type locality: Paris).

— Erichson, 1840: 849 ( Anthophagus ; synonym of alpinus ).

— Fairmaire and Laboulbène, 1856: 626 ( Anthophagus ; synonym of alpinus ).

— Kraatz, 1857d: 919 ( Anthophagus ; synonym of alpinus ).

— Ganglbauer, 1895: 706 ( Anthophagus ; synonym of alpinus ).

— Luze, 1902d: 518 ( Anthophagus ; synonym of alpinus ).

— Bernhauer and Schubert, 1910: 79 ( Anthophagus ; synonym of alpinus ).

— C. Koch, 1933a: 142 ( Anthophagus ; synonym of alpinus ).

pyrenaeus Brisout, 1863: 41 ( Anthophagus , cited as Antophagus ; Type locality: Mont-Louis).

— Fauvel, 1871a: 146 [= 1872: 120] ( Anthophagus ; characters; France).

— Fauvel, 1873b: 120 [= 1873c: 13] ( Anthophagus ; Spain).

— Redtenbacher, 1874: 265 ( Anthophagus ; characters; Austria).

— Mulsant and Rey, 1880: 22 ( Anthophagus ; characters; France).

— Luze, 1902d: 519 ( Anthophagus ; characters; Pyrenees; Cantabrian region).

— Portevin, 1929: 434 ( Anthophagus ; characters; France).

— Scheerpeltz, 1933: 1069 ( Anthophagus ; catalog).

— C. Koch, 1933a: 142 ( Anthophagus ; subspecies of alpinus ).

— C. Koch, 1934a: 196 ( Anthophagus ; subgenus Anthophagus ; race of alpinus ; Pyrenees).

— C. Koch, 1940: 374 ( Anthophagus ; subspecies of alpinus ; Spain).

— Zanetti, 1987: 407 ( Anthophagus ; subgenus Anthophagus ; subspecies of alpinus ; characters).

— Tronquet, 1998a: 114 ( Anthophagus ; cited as valid species; collected from marmot burrow; France).

bosnicus Bernhauer, 1901: 539 ( Anthophagus ; Type locality: Sarajevo auf der Bjelasnica in Bosnien).

— Luze, 1902d: 519 ( Anthophagus ; characters; Bosnia).

— Bernhauer and Schubert, 1910: 79 ( Anthophagus ; catalog).

— C. Koch, 1933a: 142 ( Anthophagus ; subspecies of alpinus ).

— C. Koch, 1934a: 196 ( Anthophagus ; subgenus Anthophagus ; Bosnia; Hercegowina).

— Zanetti, 1987: 407 ( Anthophagus ; subgenus Anthophagus ; subspecies of alpinus ; characters).

albanicus Bernhauer, 1926e: 267 ( Anthophagus ; Type locality: Korab in Albanien).

— Scheerpeltz, 1933: 1068 ( Anthophagus ; catalog).

— C. Koch, 1933a: 142 ( Anthophagus ; subspecies of alpinus ).

— C. Koch, 1934a: 197 ( Anthophagus ; subgenus Anthophagus ; race of alpinus ; Albania; Bulgaria).

speculicollis C. Koch, 1933: 29 ( Anthophagus ; variety of alpinus ; Type locality: Alpi della Carinzia, Dobratsch). — C. Koch, 1933a: 142 ( Anthophagus ; aberration of alpinus ). — C. Koch, 1934a: 197 ( Anthophagus ; subgenus Anthophagus ; aberration of alpinus ).

pseudosudeticus C. Koch, 1934a: 197, 200 ( Anthophagus ; subgenus Anthophagus ; aberration of alpinus ; [Note: Nomen nudum (Articles 1.3.4, 45.5)]; locality not cited). — Tóth, 1982: 93 ( Anthophagus ; variety of alpinus ; characters).

aprutianus C. Koch, 1934 a: 197, 201 ( Anthophagus ; subgenus Anthophagus ; race of alpinus ; Type locality: Abruzzen, Gran Sasso, Fonte Portella). — Zanetti, 1987: 407 ( Anthophagus ; subgenus Anthophagus ; subspecies of alpinus ; characters; Italy). — Zanetti, 1995: 9 ( Anthophagus ; subspecies of alpinus ; Italy).

paratus Havelka and R. Dvořák, 1953: 27 ( Anthophagus ; aberration of alpinus ; [Note: Nomen nudum (Articles 1.3.4, 45.5)]; locality cited: Carp.: Pol. Borsava).

fischeri Havelka and R. Dvořák, 1953: 27 ( Anthophagus ; aberration of alpinus ; [Note: Nomen nudum (Articles 1.3.4, 45.5)]; locality cited: Carniola: Silesia: Krnov).

alpinus Fabricius, 1793 , see: alpinus Paykull, 1790 .

angusticollis Mannerheim, 1830: 56 ( Lesteva ; Type locality: circa Petropolin). — Mannerheim, 1831: 470 ( Lesteva ; Russia). — Erichson, 1840: 850 ( Anthophagus ; synonym of abbreviatus ). — Erichson, 1840: 850 ( Anthophagus ; [Note: Misidentified as caraboides according to Bernhauer and Schubert, 1916: 78, who cited it as a synonym of angusticollis ]; characters; Europe). — Kraatz, 1857d: 923 ( Anthophagus ; synonym of caraboides ). — Kraatz, 1857d: 922 ( Anthophagus ; [Note: Misidentified as caraboides according to Bernhauer and Schubert, 1916: 78]; characters; Germany). — Thomson, 1861: 180 ( Anthophagus ; [Note: Misidentified as caraboides according to Bernhauer and Schubert, 1916: 78]; characters; Scandinavia). — Fauvel, 1871a: 142 [= 1872: 116] ( Anthophagus ; [Note: Misidentified as caraboides according to Bernhauer and Schubert, 1916: 78]; characters; France; Scandinavia; Britain; Germany; Switzerland; Italy; Austria; Russia). — J. Sahlberg, 1876: 207 ( Anthophagus ; synonym of abbreviatus ). — Mulsant and Rey, 1880: 30 ( Anthophagus ; synonym of caraboides ). — Mulsant and Rey, 1880: 29 ( Anthophagus ; [Note: Misidentified as caraboides according to Bernhauer and Schubert, 1916: 78]; characters; France). — Ganglbauer, 1895: 708 ( Anthophagus ; synonym of abbreviatus ). — Luze, 1902d: 516 ( Anthophagus ; synonym of abbreviatus ). — Bernhauer and Schubert, 1910: 78 ( Anthophagus ; synonym of abbreviatus ). — Portevin, 1929: 435 ( Anthophagus ; [Note: Misidentified as caraboides according to Zanetti, 1987: 414]; characters; France). — C. Koch, 1933a: 142 ( Anthophagus ; subgenus Dimorphoschelus ). — C. Koch, 1934a: 214 ( Anthophagus ; subgenus Dimorphoschelus ; characters; Finland; Russia; Germany; Czechoslovakia; Poland; Hungary; Croatia; Istria; Bosnia; Dalmatia; Italy; Belgium; France). — Horion, 1963: 146 ( Anthophagus ; subgenus Dimorphoschelus ; Caucasus; Siberia; Russia; Finland; Baltic States; Poland; Czechoslovakia; Hungary; Romania; Bosnia Hercegovina; Dalmatia; Croatia; Slovenia; Italy; France; Switzerland; Belgium Holland; Austria; Germany). — Lohse, 1964: 67 ( Anthophagus ; subgenus Dimorphoschelus ; characters; central Europe). — Scheerpeltz, 1968a: 18 ( Anthophagus ; catalog; Austria). — Tikhomirova, 1973a: 141 ( Anthophagus ; subgenus Dimorphoschelus ; checklist; USSR). — Muona, 1979: 18 ( Anthophagus ; Finland; Russia; Baltic States). — Burakowski, Mroczkowski, and Stefańska, 1979: 75 ( Anthophagus ; catalog; Poland). — Tóth, 1980: 98 ( Anthophagus ; subgenus Dimorphoschelus ; Hungary). — Tóth, 1982: 97 ( Anthophagus ; characters; Hungary). — Terlutter, 1984: 34, 43 ( Anthophagus ; notes; Germany).

— Renner, 1985: 99 ( Anthophagus ; collecting notes; Germany).

— Muona and Viramo, 1986: 16 ( Anthophagus ; Finland).

— Segers, 1986: 23 ( Anthophagus ; Belgium).

— Zanetti, 1987: 414 ( Anthophagus ; subgenus Dimorphoschelus ; characters; Italy).

— Lucht, 1987: 90 ( Anthophagus ; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux).

— Köhler, 1988: 163 ( Anthophagus ; notes; Germany).

— K. Koch, 1989: 236 ( Anthophagus ; habitat).

— Dettner and Reissenweber, 1991 ( Anthophagus ; defensive secretions: chemistry and systematics).

— Silfverberg, 1992: 22 ( Anthophagus ; Finland; Russia; Estonia; Latvia; Lithuania).

— K. Koch, 1992: 36 ( Anthophagus ; cited as angulicollis ; Germany).

— Janák, 1992: 85 ( Anthophagus ; Czech Republic).

— Boháč, 1993: 42 (list of species of Czech Republic and Slovakia).

— Kocian, 1993b: 92 ( Anthophagus ; Czech Republic).

— Boháč, 1993: 42 ( Anthophagus ; subgenus Anthophagus ; Slovakia; Czech Republic).

— Zanetti, 1995: 9 ( Anthophagus ; Italy).

— Ádám, 1996: 239 ( Lesteva ; Hungary).

— Böhme, 1996: 46 ( Anthophagus ; humus collection; Germany; Austria).

— Telnov, Bersevskis, Savich, Kovalevsky, Berdnikov, Doronin, Cibulskis, and Ratniece, 1997: 44 ( Anthophagus ; Latvia).

— Köhler, 1998: 159 ( Anthophagus ; collecting notes; Germany).

— Paśnik, 1998: 69 ( Anthophagus ; habitat; Poland).

— Cibuļskis, 1999: 36 ( Anthophagus ; list; Latvia).

— Herman, 2001: 19 ( Anthophagus ; discussion; has two older synonyms; prevailing use as valid maintained pending outcome of application to Commission under Article 23.9.3).

- DISTRIBUTION: Europe, Russia, Georgia.

fulvus De Geer, 1774: 25 ( Staphylinus ; [Note: This name is older than angusticollis but the synonymy needs verification since only Koch in 1933 has listed fulvus as a synonym of angusticollis ]; Type locality: Not cited).

— Paykull, 1789: 18 ( Staphylinus ; synonym of caraboides Linné ).

— Gmelin, 1790: 2029 ( Staphylinus ; synonym of caraboides ).

— Olivier, 1795:(42): 22 ( Staphylinus ; synonym of caraboides Linné ).

— Marsham, 1802: 521 ( Staphylinus ; synonym of caraboides Linné ).

— Gravenhorst, 1806: 221 ( Anthophagus ).

— Gyllenhal, 1810: 192 ( Anthophagus ; synonym of caraboides ).

— Curtis, 1830: pl. 303 ( Lesteva ; synonym of caraboides ).

— Erichson, 1840: 850 ( Anthophagus ; synonym of caraboides ).

— Kraatz, 1857d: 923 ( Anthophagus ; synonym of caraboides ).

— Luze, 1902d: 525 ( Anthophagus ; synonym of caraboides ).

— C. Koch, 1933a: 142 ( Anthophagus ; synonym of angusticollis ).

abbreviatus Fabricius, 1779: 263 ( Carabus ; [Note: This name is older than angusticollis ]; Type locality: Norwegen: Drontheim).

— Panzer, 1795: 367 ( Carabus ; characters; Germany).

— Panzer, 1796d: 2 ( Carabus ; characters; illustrations).

— Latreille, 1804: 367 ( Lesteva ; synonym of caraboides Linné ).

— Panzer, 1808a: 2 ( Carabus ; characters; illustrations).

— Erichson, 1839a: 615 ( Anthophagus ; [Note: Misidentification: Bernhauer and Schubert, 1910: 78 cited caraboides Erichson as a synonym of abbreviatus Fabricius ; Erichson did not describe the species as new, he attributed it to Linné]; characters; Germany).

— Erichson, 1840: 850 ( Anthophagus ; characters; Europe).

— Kiesenwetter, 1847a: 57 ( Anthophagus ; characters; Europe).

— Redtenbacher, 1849: 746 ( Anthophagus ; characters; Austria).

— Fairmaire and Laboulbène, 1856: 629 ( Anthophagus ; characters; France).

— Fauvel, 1869: 493 ( Anthophagus ; synonym of caraboides ).

— Fauvel, 1871a: 143 [= 1872: 117] ( Anthophagus ; synonym of caraboides ).

— Redtenbacher, 1874: 265 ( Anthophagus ; characters; Austria).

— J. Sahlberg, 1876: 207 ( Anthophagus ; Finland).

— Mulsant and Rey, 1880: 30 ( Anthophagus ; synonym of caraboides ).

— Ganglbauer, 1895: 708 ( Anthophagus ; characters; north and middle Europe; Caucasus).

— Luze, 1902d: 516 ( Anthophagus ; characters; central and northern Europe; Caucasus).

— Reitter, 1909: 183 ( Anthophagus ; characters; Germany).

— Bernhauer and Schubert, 1910: 78 ( Anthophagus ; catalog).

— Scheerpeltz, 1933: 1067 ( Anthophagus ; catalog).

— C. Koch, 1933a: 141 ( Anthophagus ; synonym of caraboides ).

— Lohse, 1964: 67 ( Anthophagus ; synonym of angusticollis ).

— Zanetti, 1987: 414 ( Anthophagus ; synonym of angusticollis ).

gracilis Heer, 1841: 573 ( Anthophagus ; Type locality: Helvetia occidentalis: Jorat, Châlet à Gobet).

— Fauvel, 1871a: 143 [= 1872: 117] ( Anthophagus ; synonym of caraboides ).

— Ganglbauer, 1895: 708 ( Anthophagus ; synonym of abbreviatus ).

— Luze, 1902d: 516 ( Anthophagus ; synonym of abbreviatus ).

— Bernhauer and Schubert, 1910: 78 ( Anthophagus ; synonym of abbreviatus ).

— C. Koch, 1933a: 142 ( Anthophagus ; synonym of angusticollis ).

obscuriceps Motschulsky, 1860: 551 ( Anthophagus ; Type locality: Montagnes de Douderhoff aux environs de St. Petersbourg).

— Fauvel, 1871a: 143 [= 1872: 117] ( Anthophagus ; synonym of caraboides ).

— Ganglbauer, 1895: 708 ( Anthophagus ; synonym of abbreviatus ).

— Luze, 1902d: 516 ( Anthophagus ; synonym of abbreviatus ).

— Bernhauer and Schubert, 1910: 78 ( Anthophagus ; synonym of abbreviatus ).

— C. Koch, 1933a: 142 ( Anthophagus ; synonym of angusticollis ).

fascifer Reitter, 1888: 150 ( Anthophagus ; variety of abbreviatus ; Type locality: Circassien).

— Ganglbauer, 1895: 708 ( Anthophagus ; aberration of abbreviatus ).

— Luze, 1902d: 516 ( Anthophagus ; synonym of abbreviatus ).

— Bernhauer and Schubert, 1910: 78 ( Anthophagus ; aberration of abbreviatus ).

— Scheerpeltz, 1933: 1068 ( Anthophagus ; aberration of abbreviatus ).

— C. Koch, 1933a: 142 ( Anthophagus ; aberration of angusticollis ).

— Tóth, 1982: 97 ( Anthophagus ; aberration of angusticollis ; characters).

roubali C. Koch, 1934 a: 214, 218 ( Anthophagus ; subgenus Dimorphoschelus ; race of angusticollis ; Type locality: Kaukasus: Taberda, Abchasie, Kuban, Wladikavkas).

— Zanetti, 1987: 414 ( Anthophagus ; subgenus Dimorphoschelus ; subspecies of angusticollis ; characters; Italy).

apenninus Baudi, 1870: 401 ( Anthophagus ; Type locality: Etruscis Apenninis; [Note: See lectotype designation by Zanetti, 1986]).

— Fauvel, 1871a: 145 [= 1872: 119] ( Anthophagus ; characters; Italy).

— Luze, 1902d: 514 ( Anthophagus ; characters; Apennines).

— Bernhauer and Schubert, 1910: 79 ( Anthophagus ; catalog).

— C. Koch, 1933: 28 ( Anthophagus ; characters).

— C. Koch, 1933a: 142, 143 ( Anthophagus ; subgenus Anthophagus ; figure).

— C. Koch, 1934a: 183 ( Anthophagus ; subgenus Anthophagus ; characters; Italy).

— Zanetti, 1986: 97 ( Anthophagus ; lectotype designation).

— Zanetti, 1987: 404 ( Anthophagus ; subgenus Anthophagus ; characters; Italy).

— Zanetti, 1995: 9 ( Anthophagus ; Italy).

- DISTRIBUTION: Italy.

apfelbecki Bernhauer, 1914d: 1 ( Anthophagus ; Type locality: Schar Dagh , Ljuboten in Albanien).

— Bernhauer, 1923d: 176 ( Anthophagus ; characters; Albania).

— C. Koch, 1933a: 142 ( Anthophagus ; subgenus Dimorphoschelus ).

— C. Koch, 1934a: 220 ( Anthophagus ; subgenus Dimorphoschelus ; characters; Albania).

- DISTRIBUTION: Albania.

aprutianus C. Koch, 1934 , see: alpinus Fabricius, 1793 .

aquatilis Sharp, 1889 , see: Philydrodes .

aquilonaris C. Koch, 1940 , see: caraboides Linné, 1758 .

armiger Gravenhorst, 1802 , see: bicornis Block, 1799 .

arpedinus Hochhuth, 1849 , see: omalinus Zetterstedt, 1828 .

arrowi C. Koch, 1933 , see: omalinus Zetterstedt, 1828 .

austriacus Erichson, 1840 , see: alpestris Heer, 1839 .

baikalensis Yablokov-Khnzorian, 1974: 62 ( Anthophagus ; subgenus Phaganthus ; Type locality: Baikal, Listvennoe, Limnologicheskii Institut).

- DISTRIBUTION: Russia.

bellicanus Gistel, 1857: 74 ( Anthophagus ; [Note: Nomen dubium]; Type locality: Tyrolis).

- DISTRIBUTION: Austria.

bicolor Grimmer, 1841: 34 ( Anthophagus ; [Note: Nomen dubium]; Type locality: Steiermark).

- DISTRIBUTION: Austria.

bicornis Block, 1799: 118 ( Staphylinus ; Type locality: Plauischer Grund bei Dresden).

— Erichson, 1840: 848 ( Anthophagus ; synonym of armiger ).

— Redtenbacher, 1849: 746 ( Anthophagus ; synonym of armiger ).

— Kraatz, 1857d: 916 ( Anthophagus ; synonym of armiger ).

— Fauvel, 1871a: 144 [= 1872: 118] ( Anthophagus ; characters; France; Belgium; Scandinavia; Germany; Switzerland; Italy; Austria).

— Fauvel, 1873b: 119 [= 1873c: 12] ( Anthophagus ; Sardinia).

— Mulsant and Rey, 1880: 13 ( Anthophagus ; synonym of armiger ).

— Ganglbauer, 1895: 706 ( Anthophagus ; characters; middle Europe; Italy).

— Luze, 1902d: 513 ( Anthophagus ; characters; central Europe; northern Italy).

— Reitter, 1909: 182 ( Anthophagus ; characters; Germany).

— Bernhauer and Schubert, 1910: 79 ( Anthophagus ; catalog).

— Portevin, 1929: 434 ( Anthophagus ; characters; France).

— Scheerpeltz, 1933: 1068 ( Anthophagus ; catalog).

— C. Koch, 1933a: 141, 143 ( Anthophagus ; subgenus Anthophagus ; figure).

— C. Koch, 1934a: 176 ( Anthophagus ; subgenus Anthophagus ; characters; Holland; Germany; Czechoslovakia; Austria; Italy; France; Bosnia; Herzegowina; Bulgaria; Hungary; Croatia).

— Horion, 1963: 143 ( Anthophagus ; subgenus Anthophagus ; Belgium; France; Switzerland; Italy; Bosnia Hercegovina; Serbia; Macedonia; Albania; Bulgaria; Romania; Hungary; Czechoslovakia; Poland; Austria; Germany).

— Lohse, 1964: 66 ( Anthophagus ; subgenus Anthophagus ; characters; central Europe).

— Tóth, 1982: 93 ( Anthophagus ; characters; Hungary).

— Zanetti, 1984: 79 ( Anthophagus ; Bulgaria).

— Zanetti, 1987: 401 ( Anthophagus ; subgenus Anthophagus ; characters; Italy).

— Lucht, 1987: 90 ( Anthophagus ; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux).

— Boháč, 1993: 42 ( Anthophagus ; subgenus Dimorphoschelus ; Slovakia; Czech Republic).

— Zanetti, 1995: 9 ( Anthophagus ; Italy).

— Zerche, 2000: 52 ( Anthophagus ).

- DISTRIBUTION: Europe.

armiger Gravenhorst, 1802: 122 ( Anthophagus ; Type locality: Not cited, but presumably Brunsvigae ).

— Latreille, 1804: 368 ( Lesteva ; cited as armigera; characters; Germany).

— Heer, 1839: 197 ( Anthophagus ; characters; Switzerland).

— Erichson, 1840: 848 ( Anthophagus ; characters; Germany).

— Kiesenwetter, 1845: 21 ( Anthophagus ; notes).

— Kiesenwetter, 1848: 64 ( Anthophagus ; characters; Sweden; Germany; Switzerland).

— Märkel and Kiesenwetter, 1848a: 326 ( Anthophagus ; notes; Carpathian Alps).

— Redtenbacher, 1849: 746 ( Anthophagus ; characters; Austria).

— Fairmaire and Laboulbène, 1856: 626 ( Anthophagus ; characters; France).

— Redtenbacher, 1857: 243 ( Anthophagus ; characters; Austria).

— Kraatz, 1857d: 916 ( Anthophagus ; characters; Germany).

— Fauvel, 1865a: 314 [= 1865b: 64] ( Anthophagus ; habitat; France).

— Fauvel, 1871a: 144 [= 1872: 118] ( Anthophagus ; synonym of bicornis ).

— Redtenbacher, 1874: 264 ( Anthophagus ; characters; Austria).

— Mulsant and Rey, 1880: 13 ( Anthophagus ; characters; France).

— Ganglbauer, 1895: 706 ( Anthophagus ; synonym of bicornis ).

— Luze, 1902d: 513 ( Anthophagus ; synonym of bicornis ).

— Bernhauer and Schubert, 1910: 79 ( Anthophagus ; synonym of bicornis ).

— Scheerpeltz, 1933: 1068 ( Anthophagus ; synonym of bicornis ).

— C. Koch, 1933a: 141 ( Anthophagus ; synonym of bicornis ).

— Zanetti, 1987: 401 ( Anthophagus ; synonym of bicornis ).

nivalis Mulsant and Rey, 1880: 15 ( Anthophagus ; Type locality: Chamonix).

— Ganglbauer, 1895: 706 ( Anthophagus ; aberration of bicornis ).

— Luze, 1902d: 513 ( Anthophagus ; aberration of bicornis ).

— Bernhauer and Schubert, 1910: 79 ( Anthophagus ; aberration of bicornis ).

— Rambousek, 1927: 77 ( Anthophagus ; aberration of bicornis ; characters).

— Scheerpeltz, 1933: 1068 ( Anthophagus ; aberration of bicornis ).

— C. Koch, 1933a: 141 ( Anthophagus ; aberration of bicornis ).

— C. Koch, 1934a: 177 ( Anthophagus ; subgenus Anthophagus ; aberration of bicornis ).

— Tóth, 1982: 93 ( Anthophagus ; aberration of bicornis ; characters; Hungary).

marginicollis Eppelsheim, 1889: 181 ( Anthophagus ; variety of bicornis ; Type locality: Croatien).

— Ganglbauer, 1895: 706 ( Anthophagus ; aberration of bicornis ).

— Luze, 1902d: 513 ( Anthophagus ; aberration of bicornis ).

— Bernhauer and Schubert, 1910: 79 ( Anthophagus ; aberration of bicornis ).

— Rambousek, 1927: 77 ( Anthophagus ; aberration of bicornis ; characters).

— C. Koch, 1933a: 141 ( Anthophagus ; aberration of bicornis ).

— C. Koch, 1934a: 177 ( Anthophagus ; subgenus Anthophagus ; aberration of bicornis ).

— Tóth, 1982: 93 ( Anthophagus ; aberration of bicornis ; characters; Hungary).

nigrinus Eppelsheim, 1889: 181 ( Anthophagus ; variety of bicornis ; Type locality: Saas). — Ganglbauer, 1895: 706 ( Anthophagus ; aberration of bicornis ).

— Luze, 1902d: 513 ( Anthophagus ; aberration of bicornis ).

— Bernhauer and Schubert, 1910: 79 ( Anthophagus ; aberration of bicornis ).

— Rambousek, 1927: 77 ( Anthophagus ; aberration of bicornis ; characters).

— Scheerpeltz, 1933: 1068 ( Anthophagus ; aberration of bicornis ).

— C. Koch, 1933a: 141 ( Anthophagus ; aberration of bicornis ).

— C. Koch, 1934a: 177 ( Anthophagus ; subgenus Anthophagus ; aberration of bicornis ).

— Tóth, 1982: 93 ( Anthophagus ; aberration of bicornis ; characters).

subfasciatus Luze, 1902d: 513 ( Anthophagus ; aberration of bicornis ; [Note: Nomen nudum (Articles 1.3.4, 45.5)]; no locality cited).

— Bernhauer and Schubert, 1910: 79 ( Anthophagus ; aberration of bicornis ).

— C. Koch, 1933a: 141 ( Anthophagus ; aberration of bicornis ).

— C. Koch, 1934a: 177 ( Anthophagus ; subgenus Anthophagus ; aberration of bicornis ).

— Tóth, 1982: 93 ( Anthophagus ; aberration of bicornis ; characters).

obenbergeri Rambousek, 1927: 77 ( Anthophagus ; aberration of bicornis ; [Note: Nomen

— C. Koch, 1933a: 141 ( Anthophagus ; aberration of bicornis ).

— C. Koch, 1934a: 177 ( Anthophagus ; subgenus Anthophagus ; aberration of bicornis ). solarii C. Koch, 1934 a: 177, 179 ( Anthophagus ; subgenus Anthophagus ; race of bicornis ;

Type locality: Gubbio; Monte Pollino, Lucania).

— Zanetti, 1987: 402 ( Anthophagus ; synonym of bicornis ). inexspectatus Havelka and R. Dvořák, 1953: 26 ( Anthophagus ; aberration of bicornis ;

[Note: Nomen nudum (Articles 1.3.4, 45.5)]; locality cited: Bohemia: Krkonose,

Iserwiese). carpathicus Havelka and R. Dvořák, 1953: 26 ( Anthophagus ; aberration of bicornis ; [Note:

Nomen nudum (Articles 1.3.4, 45.5)]; locality cited: Carp. or.: Pop Ivan). bimaculatus Lacordaire, 1835 , see: praeustus P. W. J. Müller, 1821 . bosnicus Bernhauer, 1901 , see: alpinus Fabricius, 1793 . brevicornis Kiesenwetter, 1861 , see: rotundicollis Heer, 1839 . brunneus Say, 1823 , see: Geodromicus . caesus Erichson, 1840 , see: Geodromicus . caliginosus Sharp, 1889 , see: Geodromicus . caprai C. Koch, 1933 , see: fauveli Luze, 1902 . caraboides Linné, 1758: 422 ( Staphylinus ; Type locality: Europa).

— Linné, 1760: 233 ( Staphylinus ; characters; Sweden).

— Linné, 1767: 685 ( Staphylinus ; characters; Europe).

— Fabricius, 1775: 267 ( Staphylinus ; characters; Europe).

— Goeze, 1777: 720 ( Staphylinus ; characters).

— Schrank, 1781: 234 ( Staphylinus ; characters; Austria).

— Fabricius, 1781: 337 ( Staphylinus ; characters; Europe).

— Fabricius, 1787: 222 ( Staphylinus ; characters).

— Paykull, 1789: 19 ( Staphylinus ; redescription; Sweden).

— Villers, 1789: 417 ( Staphylinus ; characters; France).

— Gmelin, 1790: 2029 ( Staphylinus ; characters).

— Fabricius, 1793: 530 ( Staphylinus ; characters).

— Olivier, 1795:(42): 22 ( Staphylinus ; characters; France, Sweden).

— Panzer, 1795: 358 ( Staphylinus ; characters; Germany).

— Cederhjelm, 1798: 119 ( Staphylinus ; characters).

— Paykull, 1800: 386 ( Staphylinus ; characters; Sweden).

— Fabricius, 1801: 601 ( Staphylinus ; characters).

— Turton, 1802: 514 ( Staphylinus ; characters; Europe).

— Gravenhorst, 1802: 120 ( Anthophagus ; characters).

— Marsham, 1802: 521 ( Staphylinus ; characters; Britain).

— Latreille, 1804: 367 ( Lesteva ; characters).

— Gravenhorst, 1806: 220 ( Anthophagus ).

— Gyllenhal, 1810: 192 ( Anthophagus ; characters).

— Gyllenhal, 1827: 460 ( Anthophagus ).

— C. Sahlberg, 1827: 274 ( Anthophagus ; characters; Finland).

— Zetterstedt, 1828: 45 ( Anthophagus ; characters; Lapland).

— Curtis, 1829: 29 ( Lesteva ; catalog; Britain).

— Curtis, 1830: pl. 303 ( Lesteva ; type species of Lesteva ; Britain).

— Mannerheim, 1830: 55 ( Lesteva ; Finland).

— Mannerheim, 1831: 469 ( Lesteva ; Finland).

— Stephens, 1834: 360 ( Lesteva ; characters; England; Scotland).

— Runde, 1835: 23 ( Lesteva ; characters; Germany).

— Lacordaire, 1835: 480 ( Lesteva ; characters; France).

— Westwood, 1838a: 18 ( Lesteva ; type species).

— Stephens, 1839: 431 ( Lesteva ; characters; England; Scotland).

— Laporte, 1840: 192 ( Anthophagus ; characters; France).

— Westwood, 1845: 52 ( Lesteva ; subgenus Anthophagus ; characters; notes).

— Kiesenwetter, 1847a: 56 ( Anthophagus ; characters; Europe).

— Redtenbacher, 1849: 746 ( Anthophagus ; characters; Austria).

— Hardy, 1851: 54 ( Anthophagus ; notes; England).

— Fairmaire and Laboulbène, 1856: 627 ( Anthophagus ; characters; France).

— Redtenbacher, 1857: 243 ( Anthophagus ; characters; Austria).

— Fauvel, 1865a: 315 [= 1865b: 65] ( Anthophagus ; habitat; France).

— Redtenbacher, 1874: 265 ( Anthophagus ; characters; Austria).

— Seidlitz, 1875: 242 ( Anthophagus ; characters; Baltic region).

— J. Sahlberg, 1876: 207 ( Anthophagus ; Finland).

— Fowler, 1888: 399 ( Anthophagus ; synonym of testaceus ).

— Ganglbauer, 1895: 709 ( Anthophagus ; subgenus Phaganthus ; characters; middle Europe).

— Luze, 1902d: 526 ( Anthophagus ; characters; central and northern Europe).

— Reitter, 1909: 183 ( Anthophagus ; subgenus Phaganthus ; characters; Germany).

— Bernhauer and Schubert, 1910: 79 ( Anthophagus ; catalog).

— Johansen, 1914: 586 ( Anthophagus ; characters; Denmark).

— Scheerpeltz, 1933: 1068 ( Anthophagus ; catalog).

— C. Koch, 1933a: 141, 143 ( Anthophagus ; subgenus Phaganthus ; figure).

— C. Koch, 1934a: 169 ( Anthophagus ; subgenus Phaganthus ; characters; Lapland; Sweden; Norway; Russia; Great Britain; Holland; Germany; Czechoslovakia; Austria; Hungary; Yugoslavia; Italy; Switzerland; France; Spain).

— Palm, 1948: 126 ( Anthophagus ; characters; Sweden; Denmark; Norway; Finland).

— Horion, 1963: 141 ( Anthophagus ; subgenus Phaganthus ; Russia; Finland; Sweden; Norway; Denmark; Baltic States; Poland; Czechoslovakia; Hungary; Romania; Bosnia; Croatia; Italy; Switzerland; Spain; France; Belgium; Holland; Britain; Austria; Germany).

— Lohse, 1964: 64 ( Anthophagus ; subgenus Phaganthus ; characters; central Europe).

— Kasule, 1968: 136 ( Phaganthus ; larval characters).

— Szujecki, 1968a: 712 ( Anthophagus ; Poland).

— Szujecki, 1969a: 248 ( Anthophagus ; notes; Poland).

— Steel, 1970a: 21 ( Anthophagus ; larval characters; biological notes).

— Pope, 1977: 23 ( Anthophagus ; Britain).

— Hammond, 1980: 137 ( Anthophagus ; doubtfully in Ireland).

— Tóth, 1982: 92 ( Anthophagus ; characters; Hungary).

— Zanetti, 1987: 393 ( Anthophagus ; subgenus Phaganthus ; characters; Italy).

— Lucht, 1987: 90 ( Anthophagus ; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

— Boháč, 1993: 41 ( Anthophagus ; subgenus Phaganthus ; Slovakia; Czech Republic).

— Zanetti, 1995: 9 ( Anthophagus ; Italy).

— Cibuļskis, 1999: 36 ( Anthophagus ; list; Latvia).

- DISTRIBUTION: Europe, Russia, Georgia.

testaceus Gravenhorst, 1802: 121 ( Anthophagus ; Type locality: Not cited, but presumably Brunsvigae ).

— Latreille, 1804: 367 ( Lesteva ; characters).

— Gravenhorst, 1806: 221 ( Anthophagus ).

— Curtis, 1830: pl. 303 ( Lesteva ; Britain).

— Curtis, 1829: 29 ( Lesteva ; catalog; Britain).

— Mannerheim, 1830: 55 ( Lesteva ; Finland).

— Mannerheim, 1831: 469 ( Lesteva ; Finland).

— Stephens, 1834: 361 ( Lesteva ; characters; England).

— Runde, 1835: 23 ( Lesteva ; characters; Germany).

— Erichson, 1839a: 615 ( Anthophagus ; characters; Germany).

— Heer, 1839: 194 ( Anthophagus ; characters; Switzerland).

— Stephens, 1839: 431 ( Lesteva ; characters; England).

— Märkel and Kiesenwetter, 1848a: 327 ( Anthophagus ; notes; Carpathian Alps).

— Redtenbacher, 1849: 746 ( Anthophagus ; characters; Austria).

— Hardy, 1851: 54 ( Anthophagus ; notes; England).

— Fairmaire and Laboulbène, 1856: 628 ( Anthophagus ; characters; France).

— Redtenbacher, 1857: 243 ( Anthophagus ; characters; Austria).

— Kraatz, 1857d: 924 ( Anthophagus ; characters; Germany).

— Hochhuth, 1862: 102 ( Anthophagus ; Russia).

— Fauvel, 1871a: 140 [= 1872: 114] ( Anthophagus ; characters; France; Britain; Germany; Austria; Switzerland; Italy; Russia).

— Fauvel, 1873b: 119 [= 1873c: 12] ( Anthophagus ; Sicily).

— Redtenbacher, 1874: 265 ( Anthophagus ; characters; Austria).

— Seidlitz, 1875: 242 ( Anthophagus ; characters; Baltic region).

— J. Sahlberg, 1876: 207 ( Anthophagus ; synonym of caraboides ).

— Mulsant and Rey, 1880: 46 ( Anthophagus ; subgenus Phaganthus ; characters; France).

— Fowler, 1888: 399 ( Anthophagus ; characters; Britain).

— Ganglbauer, 1895: 709 ( Anthophagus ; synonym of caraboides ).

— Luze, 1902d: 525 ( Anthophagus ; synonym of caraboides ).

— Bernhauer and Schubert, 1910: 79 ( Anthophagus ; synonym of caraboides ).

— Portevin, 1929: 435 ( Anthophagus ; characters; France).

— Scheerpeltz, 1933: 1068 ( Anthophagus ; synonym of caraboides ).

— C. Koch, 1933a: 141 ( Anthophagus ; synonym of caraboides ).

— Pope, 1977: 23 ( Anthophagus ; synonym of caraboides ).

— Zanetti, 1987: 393 ( Anthophagus ; synonym of caraboides ).

hookeri Stephens, 1834: 361 ( Lesteva ; Type locality: Scotland).

— Stephens, 1839: 431 ( Lesteva ; characters; England; Scotland).

— G. Waterhouse, 1858: 30 ( Anthophagus ; synonym of testaceus ).

— Fauvel, 1871a: 143 [= 1872: 117] ( Anthophagus ; synonym of caraboides ).

— Ganglbauer, 1895: 708 ( Anthophagus ; synonym of abbreviatus ).

— Luze, 1902d: 516 ( Anthophagus ; synonym of abbreviatus ).

— Bernhauer and Schubert, 1910: 78 ( Anthophagus ; synonym of abbreviatus ).

— C. Koch, 1933a: 141 ( Anthophagus ; synonym of caraboides ).

palustris Heer, 1841: 572 ( Anthophagus ; Type locality: Mt. Jorat ).

— Kraatz, 1857d: 924 ( Anthophagus ; synonym of testaceus ).

— Mulsant and Rey, 1880: 46 ( Anthophagus ; synonym of testaceus ).

— Fowler, 1888: 399 ( Anthophagus ; synonym of testaceus ).

— Ganglbauer, 1895: 709 ( Anthophagus ; synonym of caraboides ).

— Luze, 1902d: 525 ( Anthophagus ; synonym of caraboides ).

— Bernhauer and Schubert, 1910: 79 ( Anthophagus ; synonym of caraboides ).

— C. Koch, 1933a: 141 ( Anthophagus ; synonym of caraboides ).

trimaculatus Luze, 1902d: 527 ( Anthophagus ; Type locality: Caucasus, Abastuman).

— Bernhauer and Schubert, 1910: 81 ( Anthophagus ; catalog).

— C. Koch, 1933a: 141 ( Anthophagus ; subspecies of caraboides ).

— C. Koch, 1934a: 169 ( Anthophagus ; subgenus Phaganthus ; aberration of caraboides ).

maculipennis Luze, 1902d: 526 ( Anthophagus ; aberration of caraboides ; [Note: Nomen nudum (Articles 1.3.4, 45.5)]; no locality cited).

— Bernhauer and Schubert, 1910: 80 ( Anthophagus ; synonym of caraboides ).

— Scheerpeltz, 1933: 1068 ( Anthophagus ; aberration of caraboides ).

— C. Koch, 1933a: 141 ( Anthophagus ; aberration of caraboides ).

— C. Koch, 1934a: 169 ( Anthophagus ; subgenus Phaganthus ; aberration of caraboides ).

aquilonaris C. Koch, 1940: 373 ( Anthophagus ; subgenus Phaganthus ; subspecies of caraboides ; [Note: Described first by Koch (1934) as an aberration of caraboides and therefore an unavailable name (Articles 1.3.4, 45.5) but made available in 1940]; Type locality: Nordspanien: Espinama, kantabr. Kordilleren).