Landouria zonifera, Nurinsiyah & Neiber & Hausdorf, 2019

Nurinsiyah, Ayu Savitri, Neiber, Marco T. & Hausdorf, Bernhard, 2019, Revision of the land snail genus Landouria Godwin-Austen, 1918 (Gastropoda, Camaenidae) from Java, European Journal of Taxonomy 526, pp. 1-73 : 60-61

publication ID

https://doi.org/ 10.5852/ejt.2019.526

publication LSID

lsid:zoobank.org:pub:3FFC527D-E9CA-4D9F-BF59-076FA60171AC

DOI

https://doi.org/10.5281/zenodo.5587352

persistent identifier

https://treatment.plazi.org/id/81A2A657-B12B-43BE-B19D-22AE36E2525C

taxon LSID

lsid:zoobank.org:act:81A2A657-B12B-43BE-B19D-22AE36E2525C

treatment provided by

Plazi

scientific name

Landouria zonifera
status

sp. nov.

Landouria zonifera View in CoL sp. nov.

urn:lsid:zoobank.org:act:81A2A657-B12B-43BE-B19D-22AE36E2525C

Figs 44 View Figs 29–47 , 54 View Figs 52–55 , 68 View Figs 63–71 , 82 View Figs 81–84 ; Tables 3–4 View Table 3 View Table 4

Diagnosis

Landouria zonifera sp. nov. is characterized by a keeled, corneous shell with a brownish zone at the periphery, a wide umbilicus and a flagellum that is more than twice as long as the short epiphallus.

Etymology

Landouria zonifera sp. nov. is named after the brownish zone at its shell periphery (Latin: ʻzonaʼ = ʻzoneʼ, ʻferreʼ = ʻto carryʼ; used as an adjective).

Material examined

Holotype

INDONESIA • holotype (det. anat.); Yogyakarta, Kulonprogo, Sibolong , snakefruit plantation; 7°44′44″ S, 110°08′02″ E; 716 m a.s.l.; 9 Jan. 2013; A.S. Nurinsiyah and T. Setiadi leg.; D = 13.2 mm, H = 7.6 mm; MZB 17727 .

GoogleMaps

Paratypes

INDONESIA • 1 spec.; Yogyakarta, Kulonprogo, Sibolong, Sibolong Hill , agroforestry; 7°45′00″ S, 110°07′00″ E; 721 m a.s.l.; MZB 17762 13 spec.; same data as for holotype; MZB 17720 GoogleMaps 1 spec.; same data as for holotype; ZMH 133420 View Materials GoogleMaps 4 spec.; Kulonprogo, Jonggrangan ; 7°44′20″ S, 110°08′32″ E; 730 m a.s.l.; ZMH 140031 View Materials GoogleMaps .

GoogleMaps

Description

SHELL (Fig. 82; Tables 3–4 View Table 3 View Table 4 ). Depressed conical, with 5.5 –5.75 hardly convex whorls; protoconch almost smooth; teleoconch with fine, irregular wrinkles and indistinct incised spiral lines; scaly processes sparse, mainly at periphery; tuberculate around umbilicus; corneous with a brownish zone at periphery; body whorl keeled; aperture rounded rhombic; upper insertion of peristome slightly descending; peristome expanded, reflexed and slightly thickened; umbilicus slightly eccentric, wide, comprising 40% of shell diameter, hardly obscured by columellar edge.

GENITALIA ( Figs 44 View Figs 29–47 , 68 View Figs 63–71 ; Table 4 View Table 4 ). Atrium short; penis long, cylindrical, slightly dilated at proximal end, without penis coecum; membranaceous sheath around distal part of penis connected with proximal part of epiphallus by tissue; penial retractor runs from diaphragm to proximal part of short, cylindrical epiphallus; flagellum more than twice as long as epiphallus, broadest near its proximal end, then abruptly tapering, slightly crenated, without node; vas deferens narrow and long; vagina very long, cylindrical; oviduct shorter, broad, proximally tapering; peduncle of bursa copulatrix subdivided into a broader, cylindrical distal part and a narrower, longer proximal part, ending in an elongate oval bursa beside albumen gland. Right ommatophoral retractor runs between penis and vagina.

Remarks

Landouria zonifera sp. nov. clusters in the molecular phylogeny ( Fig. 1 View Fig ) with L. abdidalem sp. nov. (Fig. 56) and L. tholiformis sp. nov. (Fig. 51). Landouria zonifera sp. nov. and L. abdidalem sp. nov. have overlapping ranges. Landouria zonifera sp. nov. differs from L. abdidalem sp. nov. in the smaller (D 12.5–13.2 mm vs 13.1–16.4 mm in L. abdidalem sp. nov.), more depressed (D/H 1.73–1.81 vs 1.93–2.14 in L. abdidalem sp. nov.), corneous shell with a brownish zone at the periphery, the shorter penis and a flagellum that is more than twice as long as the short epiphallus. Furthermore, Landouria zonifera sp. nov. occurs sympatrically with L. menorehensis sp. nov. ( Fig. 88 View Figs 85–88 ) from which it can easily be distinguished by the keeled, depressed conical, corneous shell with a brownish zone at the periphery.

Distribution

Landouria zonifera sp. nov. is endemic to the Menoreh karst area (Fig. 54).

MZB

Museum Zoologicum Bogoriense

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