Landouria tholiformis, Nurinsiyah & Neiber & Hausdorf, 2019

Nurinsiyah, Ayu Savitri, Neiber, Marco T. & Hausdorf, Bernhard, 2019, Revision of the land snail genus Landouria Godwin-Austen, 1918 (Gastropoda, Camaenidae) from Java, European Journal of Taxonomy 526, pp. 1-73 : 37-38

publication ID

https://doi.org/ 10.5852/ejt.2019.526

publication LSID

lsid:zoobank.org:pub:3FFC527D-E9CA-4D9F-BF59-076FA60171AC

DOI

https://doi.org/10.5281/zenodo.11238072

persistent identifier

https://treatment.plazi.org/id/C083E997-420A-4CAA-8A67-088C02177B83

taxon LSID

lsid:zoobank.org:act:C083E997-420A-4CAA-8A67-088C02177B83

treatment provided by

Plazi

scientific name

Landouria tholiformis
status

sp. nov.

Landouria tholiformis View in CoL sp. nov.

urn:lsid:zoobank.org:act:C083E997-420A-4CAA-8A67-088C02177B83

Figs 7 View Figs 6–11 , 20 View Figs 15–24 , 34 View Figs 29–47 , 51–52 View Figs 48–51 View Figs 52–55 ; Tables 3–4 View Table 3 View Table 4

Diagnosis

Landouria tholiformis sp. nov. is characterized by a brownish-corneous, sharply keeled shell with scaly processes all over the shell, with a bulge at the proximal end of the penis forming a short coecum, and a crenated flagellum, which tapers abruptly at its proximal end.

Etymology

Landouria tholiformis sp. nov. is named after its dome-shaped shell (Latin: ʻtholusʼ = ʻdomeʼ, ʻformaʼ = ʻshapeʼ; used as an adjective).

Material examined

Holotype

INDONESIA • holotype (det. anat.); Gunung Kidul , Tepus, near Poktunggal Beach, agroforestry; 8°08′42″ S, 110°37′52″ E; 137 m a.s.l.; 9 Aug. 2014; A.S. Nurinsiyah, F.L.H. Irsyad, F.J. Sari and E. Nurlaela leg.; D = 8.7 mm, H = 5.2 mm; MZB 17999 .

GoogleMaps

Paratypes

INDONESIA • 8 spec.; Yogyakarta, Gunung Kidul, Tepus, Pego Hill , agroforestry; 8°06′10″ S, 110°37′50″ E; 230 m a.s.l.; MZB 19299.

GoogleMaps

Description

SHELL ( Figs 7 View Figs 6–11 , 51 View Figs 48–51 ; Tables 3–4 View Table 3 View Table 4 ). Depressed conical, with a dome-shaped spire, with 6–6.25 slightly convex whorls; protoconch almost smooth; teleoconch with fine, irregular wrinkles; apical side with indistinct incised spiral lines, umbilical side with more distinct spiral lines; scaly processes all over shell; tuberculate around umbilicus; brownish-corneous; body whorl sharply keeled; aperture rounded rhombic; upper insertion of peristome descending; peristome expanded, reflexed and thickened; umbilicus slightly eccentric, wide, comprising 32–40% of shell diameter, hardly obscured by columellar edge.

GENITALIA ( Figs 20 View Figs 15–24 , 34 View Figs 29–47 ; Table 4 View Table 4 ). Atrium short; penis long, becoming slightly broader proximally, with a bulge at proximal end forming a short coecum; membranaceous sheath around distal part of penis connected with epiphallus by tissue; penial retractor runs from diaphragm to proximal part of cylindrical epiphallus; flagellum moderately long, broadest in its middle part, crenated, without node, abruptly tapering at its proximal end; vas deferens narrow and long; vagina moderately long; oviduct short, broad, proximally tapering; peduncle of bursa copulatrix subdivided into a broader, almost cylindrical distal part that abruptly passes into a narrower and longer proximal part that ends in an elongate oval bursa beside albumen gland. Right ommatophoral retractor runs between penis and vagina.

Remarks

Landouria tholiformis sp. nov. clusters in the phylogeny ( Fig. 1 View Fig ) with L. abdidalem sp. nov. (Fig. 56 View Figs 56–59 ). It differs from that species in the more depressed (D/H 1.77–1.88 vs 1.93–2.14 in L. abdidalem sp. nov.), less sharply keeled shell with scaly processes all over the teleoconch ( Fig. 7 View Figs 6–11 ), in the flagellum, which is slightly longer than the short epiphallus, and in having a bulge at the proximal end of the penis (perhaps this is an individual abnormity).

Landouria tholiformis sp. nov. occurs sympatrically with L. sewuensis sp. nov. (Fig. 58), from which it differs in the less sharply keeled shell with a dome-shaped spire, a more distinct granular microsculpture, with scaly processes all over the shell ( Fig. 7 View Figs 6–11 ) and less distinct spiral lines at the umbilical side, the bulge at the proximal end of the penis that forms a short coecum, and a narrow, crenated flagellum. The flagellum of L. sewuensis sp. nov. is broader than that of L. tholiformis sp. nov. and equipped with two rows of tubercles. There is a globular bulge in the distal part of the vagina in L. sewuensis sp. nov. which is absent in L. tholiformis sp. nov.

Distribution

Landouria tholiformis sp. nov. is endemic to the Gunung Sewu karst region in Gunung Kidul Regency (Fig. 52).

MZB

Museum Zoologicum Bogoriense

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