Ladda Grishin, 2019

Cong, Qian, Zhang, Jing, Shen, Jinhui & Grishin, Nick V., 2019, Fifty new genera of Hesperiidae (Lepidoptera), Insecta Mundi 731 (731), pp. 1-56 : 18-20

Ladda Grishin , new genus

http://zoobank.org/ 4BBF25C1-3C31-4B69-9499-516C8251D74E

Type species. Cyclopides eburones Hewitson, 1877 .

Definition. A sister genus to Dalla Mabille, 1904 (type species Cyclopides eryonas Hewitson, 1877 ), where these species were placed previously ( Fig. 8 ). Consists of the ibhara, part of quadristriga and part of caenides groups of Evans (1955) and keys to H.6.56b, H.6.44b, or H.6.53b. Distinguished by the combination of straight (non-concave) costa of forewing with unmarked, dark brown above hindwing. If hindwing is spotted above, then the spot is single, centered around the cell, ventral hindwing not plain yellow and without a pale ray from base to margin and if a pale spot is present in M 3 -CuA 1 cell, then it is either joined with the spot in CuA 1 -CuA 2 cell, or closer the discal cell spot than to margin and ventral hindwing, and the pale spot in CuA 1 -CuA 2 cell on ventral hindwing is not connected with distal margin by a broadening pale area (but could be connected with the area of the same width as the spot or narrower). In DNA, a combination of the following base pairs is diagnostic: aly103.37.1:A134T, aly638.15.11:T5060C, aly23605.1.35:A566G, aly1139.25.1:G552T, aly23605.1.35:C580G.

Etymology. The name is a feminine noun in the nominative singular, a play of letters, from Dalla .

Species included. The type species, Dalla tona Evans, 1955 , Dalla decca Evans, 1955 , Butleria morva Mabille, 1898 , Thanaos ibhara Butler, 1870 , Carterocephalus plancus Hopffer, 1874 , Cyclopides crithote Hewitson, 1874 , Dalla calima Steinhauser, 1991 , Dalla rosea Evans, 1955 , Butleria granites Mabille, 1898 , Dalla parma Evans, 1955 , Dalla mars Evans, 1955 , Pamphila cuadrada Weeks, 1901 , Butleria quadristriga Mabille, 1889 , Dalla rubia Evans, 1955 , Argopteron xicca Dyar, 1913 , Dalla pedro Steinhauser, 2002 , Dalla connexa Draudt, 1923 , Dalla disconnexa Steinhauser, 2002 , Dalla vista Steinhauser, 2002 , Butleria ticidas Mabille, 1898 , Cyclopides caenides Hewitson, 1868 , Dalla bos Steinhauser, 1991 , Dalla pura Steinhauser, 1991 , Dalla mora Evans, 1955 , Dalla carnis Evans, 1955 , Butleria monospila Mabille, 1898 , Dalla simplicis Steinhauser, 1991 , Dalla seirocastnia Draudt, 1923 , Dalla celsus Steinhauser, 2002 , Dalla pantha Evans, 1955 , Dalla puracensis Steinhauser, 1991 , Dalla ochrolimbata Draudt, 1923 .

Parent taxon. Tribe Heteropterini Aurivillius, 1925 .

Comments. Although this genus does not disrupt the monophyly of Dalla when combined with it, Ladda is split from Dalla , due to the large genetic distance between the two genera, each forming a tight cluster of species well separated from the other. Genetic differentiation between these two genera is similar to

that between traditional genera such as Heteropterus Duméril, 1806 and Leptalina Mabille, 1904 , or Carterocephalus Lederer, 1852 and Metisella Hemming, 1934 .

Figures

Figure 8. Genomic trees of Heteropterinae and Trapezitinae. See Fig. 2 legend for notations.