Laccophilus nodieri Regimbart , 1895

Bistroem, Olof, Nilsson, Anders N. & Bergsten, Johannes, 2015, Taxonomic revision of Afrotropical Laccophilus Leach, 1815 (Coleoptera, Dytiscidae), ZooKeys 542, pp. 1-379 : 104-106

Taxon classification Animalia Coleoptera Dytiscidae

Laccophilus nodieri Regimbart, 1895 Figs 77-78, 268, 424, 547

Laccophilus nodieri Régimbart 1895: 134 (original description, faunistics); Guignot 1946c: 270, 273, 277 (description, faunistics, discussion); Guignot 1959a: 562, 567 (description, faunistics); Bruneau de Miré and Legros 1963: 873, 888 (faunistics); Bilardo and Pederzani 1978: 119: (faunistics, description); Biström 1979: 22 (faunistics); Medler 1980: 155 (faunistics, list.); Nilsson 2001: 247 (catalogue, faunistics); Pederzani and Reintjes 2002: 38, 40 (discussion, faunistics); Reintjes 2004: 68 (faunistics); van Vondel 2005: 131 (faunistics); Nilsson 2015: 215 (catalogue, faunistics).

Type locality.

Mali: Badoumbé.

Type material studied

(2 exs.). Lectotype (by present designation) male: "Ht. Sénégal Badoumbé Dr. Nodier I à V 1882 / male symbol /Co-Type / Laccophilus nodieri Rég. cotype" (MNHN; habitus in Fig. 424). - Paralectotype: female: “Badoumbé Ht. Senegal / Museum Paris coll. Maurice Régimbart 1908 / nodieri Reg." (1 ex. MNHN). [Comment: most probably the female paralectotype belongs to Laccophilus modestus .]

Additional material studied

(70 exs.). Gambia: "W Div. Abuko Nat. Res. 27.11. 2003 Vondel / Laccophilus nodieri det. Rocchi" (1 ex. CSR); "Abuko Nat. Res., at light at the Bamboo pool 18.30-20.30 18.11. 1977, UTM 28PCK2181 / Cederholm et. al leg. / Laccophilus nodieri Rég. det. Brancucci 85" (1 ex. MZLU). - Senegal: "Casamance Tabor" (1 ex. MNHN); "Dakar 5. 1939" (2 exs. MNHN). - Mali: "Goundaka, Bandiagara river 14°29'N, 3°56'W, 12.2. 2000 Komarek & Mayer / Laccophilus nodieri Rég. det. Wewalka 2001" (1 ex. NMW); "W Bandiagara, pools 14°22'N, 3°41'W, 12.2. 2000 Komarek & Mayer / Laccophilus nodieri Rég. det. Wewalka 2001" (1 ex. NMW); "W50 km E Djenne 13°50'N, 4°25'W, 12.2. 2000 Komarek & Mayer / Laccophilus nodieri Rég. det. Wewalka 2001" (1 ex. NMW). - Sudan: "Aluakulak 30,5E 6,30N 14.5. 1954 / Laccophilus nodieri Reg. det. J. Omer-Cooper" (1 ex. AMGS); "Alel rock pool 30,56 E 6,11N 18.I. 1954 JJOC." (7 exs. AMGS); "Sandy river 50 mi. NW of Juba 29.1. 1954 JJOC." (4 exs. AMGS); "Stream from hot springs Nyangwara 30,51E 4,39N, 29.I. 1954 JJOC." (2 exs. AMGS); "Sandy river 50 mi. NW of Juba 29.I. 1954 JJOC." (2 exs. AMGS). - Ivory Coast: “Comoé N8,5° - W3,5°, 22.3. 1999 Reintjes" (6 exs. NMW); same and " Laccophilus nodieri det. Rocchi" (3 exs. CSR). - Burkina Faso: "Pundu, Mte Volta 1927-28 Dez-Juni Olsufiejev" (3 exs. NHRS). - Ghana: "17 mi S Palbe 1.9. 1971, filtered black light, Gruwell" (1 ex. USNM). - Nigeria: "Stream crossing Kaduna rd. nr Zaria 4.4. 1963 JOC." (1 ex. AMGS); "Trib. of R. Gagere en route Zaria-Katsina 5.IV. 1963 JOC." (2 exs. AMGS); "Stream 64 mi. from Bida on Jebba rd.15.IV. 1963 JOC." (1 ex. AMGS); "Detritus pond 45 mi. from Jos on Bauchi rd.9.IV. 1963 JOC." (1 ex. AMGS); "R. Niger, bridge Kontagora-Kaduna rd. 3.IV. 1963 JOC." (1 ex. AMGS); "Kontagora stream 3.IV. 1963 JOC." (1 ex. AMGS); "Stream nr. Zaria 4.IV. 1963 JOC." (8 exs. AMGS); "Zaria Pr., Zaria 5-6.3. 1949 Malkin" (1 ex. BMNH, 1 ex. MZH); "NC St. Zaria 2-3.8. 1973 Linnavuori" (1 ex. MZH); "NW St. Yelwa 23.7. 1973 Linnavuori" (1 ex. MZH); "Kano St., Kano-Wudil 17.5. 1973 Linnavuori" (1 ex. MZH); "R. Ogun, Olokomeji nr. Ibadan 24.III. 1963 JOC." (2 exs. AMGS); "Pond, road Dawria-Kano 6.4. 1963 JOC." (4 exs. AMGS). - Zaire: "PNG, Utukuru 14/s, 22.7. 1952 De Saeger 3812" (2 exs. NHMB).

Diagnosis.

Externally Laccophilus nodieri resembles very much of Laccophilus flaveolus . Both species have elytral irrorations in part reduced, which makes coverage of it uneven and patchy. Shape of penis, however, is peculiar with a distinct, sharp knob in inner curvature. Corresponding sharp knob lacks in Laccophilus flaveolus .

Description.

Body length 4.0-4.3 mm, width 2.2-2.4 mm. Elytral colour pattern consists of rather fine, in part unevenly distributed irrorations (Fig. 424).

Head: Pale ferrugineous; no distinct colour pattern. Impunctate, except at eyes; with fine, dense and somewhat irregular punctures. Areas of punctures extend a little towards middle of head. Slightly mat, finely microsculptured. Reticulation almost simple; only indistinct fragments of large meshes discernible. Large meshes, when discerned, contain 3-6 fine meshes.

Pronotum: Pale ferrugineous; lacks colour pattern. Rather shiny, although very finely microsculptured. Reticulation double, but difference between size classes small. Large meshes contain 3-6 fine meshes. At margins with fine, scattered and irregular punctures, which basally in middle are indistinct.

Elytra: Pale ferrugineous, with distinctly reduced, somewhat vague dark ferrugineous irrorations. Irrorations are sparsest basally and in a transverse area posterior to middle (Fig. 424). Slightly mat; finely microsculptured. Reticulation indistinctly double; difference between size classes small. Large meshes, when discernible, contain 3-6 fine meshes. Mesh-organization in part slightly vague. Almost impunctate except for discal row of punctures; consist of fine, irregularly placed fine punctures. Dorsolateral and lateral row of punctures reduced to a few irregular fine punctures.

Ventral aspect: Pale ferrugineous to ferrugineous; no distinct colour pattern exhibited. Almost impunctate. Submat to rather shiny, very finely, in part indistinctly microsculptured. Metacoxal plates in frontal half with some transversely located very shallow furrows. Abdomen basally with fine curved striae. Prosternal process rather slender, apex slightly extended and pointed. Apical ventrite asymmetric; provided with a minute but sharp lateral knob (Fig. 77).

Legs: Pro- and mesotarsus slightly enlarged, somewhat extended and provided with distinct suckers.

Male genitalia: Penis in lateral aspect slightly curved, broad, provided with a sharp, minor process, approximately in middle of the inner side of penis. Extreme apex of penis hooked (Fig. 268).

Female: Externally as male but pro- and mesotarsus slender. Apical ventrite not distinctly asymmetric; lacks lateral knob (Fig. 78).

Distribution.

Gambia, Senegal, Mali, Sudan, Ivory Coast, Burkina Faso, Ghana, Nigeria, Zaire (Fig. 547). Personally verified records only accepted in map. Additional country records are Chad ( Bruneau de Miré and Legros 1963), Guinea ( Reintjes 2004) and Benin (van Vondel 2005 a).

Collecting circumstances.

Sampled e.g. in pools with loamy bottom and in stagnant remain of brooklet (van Vondel 2005 a). Label data give the species to be collected from various pools, at light and in a sandy river.