Kalophrynus subterrestris Inger, 1966 Labang Sticky Frog

Zug, George R., 2015, Morphology and Systematics of Kalophrynus interlineatus-pleurostigma Populations (Anura: Microhylidae: Kalophryninae) and a Taxonomy of the Genus Kalophrynus Tschudi, Asian Sticky Frogs, Proceedings of the California Academy of Sciences 62 (5), pp. 135-190 : 180

Kalophrynus subterrestris Inger, 1966 Labang Sticky Frog

Kalophrynus subterrestris Inger, 1966, Fieldiana, Zoology 52:137 [type locality: “ Sungei Seran, Labang, Bintulu District, Fourth Division, Sarawak”, East Malaysia (Borneo)] .

TYPE MATERIAL.— HOLOTYPE: Field Museum of Natural History 150421 . PARATYPES: FMNH 140238 , 154022–026 , 157652–656 . All by original designation .

DEFINITION.— Small, adult females 25.8–27.0 mm SVL (n = 4), adult males 21.0– 23. 4 mm SVL (n = 6); head moderately long NA % HeadL/ SVL; head slightly wider than long NA % HeadW/HeadL; snout moderately broad NA % SnW/HeadL; naris closer to snout than to eye NA NA NarEye/SnEye; eye moderately large NA % EyeD/HeadL; tympanum visible and smaller than eye 50–66 % Tymp/EyeD; moderate length forelimb NA % Forarm/ SVL and forearm equal crus length NA % Forarm/CrusL; hindlimb moderately long NA % HndlL/ SVL, 40–45 % CrusL/ SVL, and NA % CrusL/ThghL; hindfoot well developed 85 % HndfL/CrusL.

Vomerine teeth presence or absence not reported; palatal fold morphology, vomerine fold not reported, postorbital fold angular and strongly notched, buccal fold crenulated; tongue size and shape not reported.

Fingers with slight basal web; lengths 3>2>1>4, fourth nub-like; tips rounded and not dilated; single subarticular tubercle between palmar tubercle and tip of finger 4, number of subarticular tubercles not reported for other fingers; fleshy palmar tubercle. Toes lightly webbed to center of distal subarticular tubercles of toe 3, lengths 4>3>5>2>1; tips rounded; conspicuous subarticular tubercles on first four toes, number not reported, none on toe 5; low inner and outer metatarsal tubercles.

In life, dorsum dark gray without darker or lighter markings; usually no light diagonal line of side or dorsolaterally on head; inguinal region with or without a dark spot, if present without light border; side of body orange in groin; venter dusky mottling anteriorly, white belly.

ETYMOLOGY.— The name subterrestris was selected because two individuals were discovered in burrows.

DISTRIBUTION.— This species occur broadly, if not commonly in northern Borneo ( Sarawak and Sabah).

NATURAL HISTORY.— Most individuals of this species derived from the floor of primary rain forest in hillside situations, and most were found beneath the floor litter and two in long burrows.

COMMENTS.— Preceding information extracted from Inger (1966).