Kalophrynus barioensis Matsui and Nishikawa, 2011 Bario Sticky Frog

Zug, George R., 2015, Morphology and Systematics of Kalophrynus interlineatus-pleurostigma Populations (Anura: Microhylidae: Kalophryninae) and a Taxonomy of the Genus Kalophrynus Tschudi, Asian Sticky Frogs, Proceedings of the California Academy of Sciences 62 (5), pp. 135-190 : 155-156

Kalophrynus barioensis Matsui and Nishikawa, 2011 Bario Sticky Frog

Kalophrynus barioensis Matsui and Nishikawa, 2011, Current Herpetology 30:146 [type locality: “Jln. Arul Dalan trail to New Dam, Bario, State of Sarawak, Malaysia (03°45′N, 115°26′E, 1141 m asl)”] .

TYPE MATERIAL.— HOLOTYPE: Sarawak Museum MU455 .16, by original designation . PARATYPE: Kyoto University Graduate School 53128–129, 5371 .

DEFINITION.— Small, adult female 20.5 mm SVL (n = 1), adult males 17.5–19. 8 mm SVL (n = 5); head moderately long 33–36 % HeadL/ SVL; head width equals length 33–36 % HeadW/ SVL; naris closer to snout than to eye 15–21 % NarEye/ SVL; eye moderately large 15–20 % EyeD/ SVL; tympanum visible and about equal to eye 60–140 % Tymp/EyeD; slender moderately long forelimb 56–63 % Forelimb/ SVL; hindlimb moderately long 126–141 % HndlL/ SVL, 42–47 % CrusL/ SVL.

Vomerine teeth presence or absence not reported; palatal folds present, morphology not report- ed; tongue entire.

Fingers with slight basal web; lengths 3>2>1>4, 4 th very short; tips rounded and not dilated; subarticular tubercles round, indistinct, and one on digit 2, two on 3; outer palmar tubercle large, inner indistinct; no nuptial excrescences on fingers. Toes slightly webbed not extending beyond proximal subarticular tubercles of toe 4, lengths 4>3>2>5>1; tips rounded, not dilated; subarticular tubercles indistinct and one on digit 2, two on 3, three on 4, and none on 1 and 5; oval inner and no outer metatarsal tubercles.

Color in life, dorsum orangish brown with faint brown hour-glass mark from between eyes to suprascapular area and largely immaculate from mid-trunk rearward; body; large black inguinal spot, no light edge; loris and temporal area to inguina dark brown, fading ventrally; yellowish white oblique stripe from above eye to lower inguinal area; chin and chest dark brown with indistinct lighter bands, posterior chest and belly yellowish white with scattered black spots anteriorly.

ETYMOLOGY.— The name barioensis derives from the town of Bario in the Kelabit Highlands of Sarawak.

DISTRIBUTION.— Sarawak, Borneo. Presently, K. barioensis is known only from the vicinity of the type locality .

NATURAL HISTORY.— These frogs are ground-dwelling residents of broad-leaf and bamboo forest. All specimens were collected in mid August from the leaf litter along forest trails. Males, hidden beneath the leaf litter, were calling in scattered small chorus and chorused in late afternoon and early evening whether it rained or not.

COMMENTS.— The extreme variance of the Tymp/EyeD data suggest an error in the measurement of one individual.

Preceding information extracted from Matsui and Nishikawa (2011).