Isometrus (Isometrus) maculatus (DeGeer, 1778)

Kovařík, František, 2003, A review of the genus Isometrus Ehrenberg, 1828 (Scorpiones: Buthidae) with descriptions of four new species from Asia and Australia, Euscorpius 10 (10), pp. 1-19 : 2-4

Isometrus (Isometrus) maculatus (DeGeer, 1778)

? = Scorpio europaeus Linné, 1758: 625 ; Fabricius, 1775: 399; Guérin-Méneville, 1843: 11 (syn. by Thorell, 1876a: 8).

=? Scorpio americus Linné, 1758: 624 (syn. by Thorell, 1876a: 8).

Scorpio americanus: Linné, 1775: 1091 .

Scorpio maculatus DeGeer, 1778: 346 .

= Scorpio dentatus Herbst, 1800: 55 (syn. by Pavesi, 1881: 537).

= Buthus (Isometrus) filum Ehrenberg in Hemprich & Ehrenberg, 1828: pl.1, fig. 3; Moritz & Fischer, 1980: 314; Braunwalder & Fet, 1998: 32 (syn. by Peters, 1862: 515).

= Tityus aethiops C. L. Koch, 1843 : fig. 856; C. L. Koch, 1845a: 11 (syn. by Karsch, 1879 a: 114).

= Tityus longimanus C. L. Koch, 1843 : fig. 857; C. L. Koch, 1845a: 13 (syn. by Thorell, 1888: 330).

= Lychas paraensis C. L. Koch, 1845a: 6 , fig. 963; Moritz & Fischer, 1980: 321 (syn. by Thorell, 1888: 406).

Lychas maculatus: C. L. Koch, 1845b: 1 , fig. 960.

Lychas americanus: C. L. Koch, 1845b: 2 , fig. 961; Lamoral & Reynders, 1975: 510.

= Scorpio (Lychas) gabonensis Lucas, 1858: 430 ; Lamoral & Reynders, 1975: 511 (syn. by Peters, 1862: 515).

= Scorpio (Lychas) guineensis Lucas, 1858: 432 (syn. by Peters, 1862: 515).

Isometrus maculatus: Simon, 1876: 219 ; Thorell, 1876a: 8; Oates, 1888: 250; Pocock, 1890b: 236; Pocock, 1893b: 376; Kraepelin, 1899: 66; Kraepelin, 1905: 334; Kraepelin, 1908: 183; Hirst, 1913: 32; Kraepelin, 1913: 134, 185; Kopstein, 1921: 126; Kopstein, 1923: 184; Fage, 1929: 72; Roewer, 1929: 611; Fage, 1936: 181; Fage, 1944: 71; Lamoral & Reynders, 1975: 508; Vachon, 1982: 90; Kovařík, 1997b: 362; Huber et al., 2002: 53; Soleglad & Fet, 2003: 5.

=? Isometrus sonticus Karsch, 1879 a: 116 ; Moritz & Fischer, 1980: 324 (syn. by Kraepelin, 1891: 245).

=? Lychas mabillianus Rochebrune, 1884: 28 ; Pocock, 1891: 447 (syn. by Simon, 1886: 86).

Isometrus europaeus: Lönnberg, 1897: 180 , 183; Pocock, 1898: 326; Pocock, 1899b: 835; Pocock, 1900: 46; Pocock, 1902: 38; Wu, 1936: 119; Takashima, 1945: 85; Takashima, 1948: 76; Takashima, 1950: 18.

= Isometrus europaeus quinquefasciatus Franganillo, 1931: 118 ; Franganillo, 1934: 166 (syn. by Moreno, 1940: 43).

= Isometrus madagassus Roewer, 1943: 217 ; Vachon, 1969: 417; Probst, 1973: 330; Lamoral & Reynders, 1975: 509 (syn. by Lourenco, 1996: 444).

Isometrus (Isometrus) maculatus: Vachon, 1972: 177 ; Vachon, 1976: 38; Kovařík, 1994: 197; Kovařík, 1995: 187; Kovařík, 1997a: 8; Kovařík, 1998b: 111; Fet & Lowe, 2000: 147; Kovařík, 2002: 8.

Isometrus (Isometrus) madagassus: Vachon, 1972: 177 ; Vachon, 1976: 38; Vachon, 1982: 90; Kovařík, 1994: 202.

Isometrus (Raddyanus) europaeus: Tikader & Bastawade, 1983: 286 .

TYPE LOCALITY AND TYPE REPOSITORY. Suriname and Pennsylvania ; NHRS.

TYPE MATERIAL EXAMINED. Madagascar, Tananarive , 1♂ (holotype of Isometrus madagassus ), SMFD No. 8879/221 .

OTHER MATERIAL EXAMINED. Algeria, Stadt , 1♀, SMFD No. 6697/104. Argentina, Resistencia , 1♀, leg. Balzani, MZUF .? Australia, Mariannen, Saipan , 1♀, leg. Schnee, ZMHB No. 899/1911. Australia, Caivus , 1♂, leg. Felten, SMFD ; Queousland Cooktowon , 1♀, 1938, FKCP . Brazil, Amazonas, Manacapuru , 4♂ 3♀, VIII.1924, leg. W. Ehrhardt, SMFD ; Amazonas, Manacapuru, Sdimoes , 3♂ 6♀ 1juv., VII.1924, leg. W. Erhardt, SMFD ; Olinda (Recifo), St. Penambuco , 2♀, 24.V.1930, SMFD ; Manacapuru , 1♂ 1♀, 1934, FKCP ; Mendes, 100 km v. Rio in der Sierra do Mor , VII– IX.1935, 1♂, leg. Friedländer, ZMUH ; Tiriyo, Maloca am, ob. Rio Parú de Oeste , 1♀, 24–30.I.1961, leg. Sattler, SMFD ; 1♂ 1♀, leg. baron von Dungero, ZMHB No. 344D/25; Paraná, Rio Branco , 1♂, leg. Uhe, ZMHB No. 1360/1912; Alacranes Machorro , 2♂ 1♀, MZUF . Costa Rica, Farm Waldeck, Ebene Limon (in Hotel ), 30.XI.1933, 1♂, leg. F. Nevermann, ZMUH . Democratic Republic of Congo, Mutskatscha Dilolo , 1931, 1♀, leg. F. Haas, SMFD ; Kisangani , III.1975, 1♂, MZUF ; Mombasa Haut Zaire , V–VI.1987, 1♂, leg. M. Borri, MZUF . Ethiopia, ( Abyssinia ), 1905, 1♀, ZMHB . India, Bombay, Mahateron , I.1873, 1♂ 1♀, leg. G. Pilastri, MZUF ; Dekan, Anamalei , 1♂ 1♀ 2juvs., SMFD No. 1086/17; Karnataka State, Mysore , 15 mi. S Chikmagalur, 900 m., 20.II.1962, 1♀, leg. E. S. Ross & D. Q. Cavagnaro, CASC ; Karnataka State, Mysore , 7 mi. SE Sakleshpur, 900 m., 21.II.1962, 1♀, leg. E. S. Ross & D. Q. Cavagnaro, CASC ; Tamil Nadu State, Topslip, Anamalai Hills , 760 m, 18.III.1962, 1juv. (det.?), leg. E. S. Ross & D. Q. Cavagnaro, CASC ; Tamil Nadu State, Marayoor , 27 mi NE Munnar, 1.IV.1962, 1♀ 10juvs. before first ecdysis, leg. E. S. Ross & D. Q. Cavagnaro, CASC ; Karwa , 1965, 1♀, FKCP ; West Bengal State, Khalishani Dt. Howran, 10.VI.1966, 1♂, leg. Isa Santra, CASC ; Meghalaya State, West Garo Hills reg., BAGMARA, 19–21.V.1996, 1♂, alt. 100± 50m, GPS N25.11.5 E90.38.5 (WGS 84), leg. E. Jendek & O. Šauša, FKCP ; Nandi Hills , N of Bangalone, 900–1300 m, 22.VII.1996, 1♂, leg. Werner & Lorenz, FKCP ; Tamil Nadu, Nilgiri Hills, Kunchapani , X.1997, 1♀, FKCP ; Tamil Nadu State, Javadu Hills, Kavalore, Vellore Dist. , 1♀, 2000, coll. T.R. S.N., FKCP . Indonesia, Sumatra, Deli , 1♀, leg. Heyden, SMFD No. 5169; Sumatra, Atjeh (Hügellani Boeloch Blang-Asa über lho Seumawa ), 1♂, III.192?, leg. Rookmarker, SMFD . Kenya, (Brit. E. Africa), Mombasa, VI.1907, 1♂ 1♀, leg. Schauer, ZMHB No. 2013/07. Madagascar, Flot Prune by Tamatave , 1♀, leg. Dr. Friederichs, ZMHB No. ZMB 30803; Kap Diego , 1♀, leg. Dr. Friederichs, ZMHB No. ZMB 30804; Nossibé , 1898, 2♂ 4♀, leg. Ebenar, SMFD No. 5206; 1♂, 1902, FKCP . Maldives, Atollo Mase Sud , II.1976, 1♀ (det.?), leg. L. Azzaroli, MZUF . Micronesia, Caroline Island , 1♀, MZUF . New Guinea, Hollandia , 1♀, leg. Mayr, ZMHB No. 47D/29. Pakistan, E., Baraiadhala Forest Res., 150 ft., 21.IX.1961, 1♀, leg. E. S. Ross & D. Q. Cavagnaro, CASC ; Lahore, 250 m., 13.XII.1961, 1juv., leg. E. S. Ross & D. Q. Cavagnaro, CASC. Peru, Callanga , 1♀, FKCP . Philippines, Sabaan , 23.I.1916, 1♀, leg. S. Böttcher and V. Heyne, ZMHB No. 137D/24; Luzon , Manila, 21.IV.1945, 1♂, leg. T. Aarens, CASC ; I.1951, 2♂ 1♀ 2juvs., leg. Terry, CASC. Seychelles, Mahé Island , 1juv., ZMHB . Somalia, Belet Amin , VII.1934, 1juv., leg. S. Patrizi, MZUF . Surinam, 1♂, Cardua, SMFD . Syria, Damascus, 1870, 1♂, leg. G. Pilastri, MZUF . Tanzania, Zanzibar Island , 29.VII.1889, 1♀, ZMHB ; Dar-es-Salam, Warenkiste , 6.II.1927, 1♀, ZMUH ; Dar-es-Salam , 1929, 1♂ 1♀, leg. H. Brauns, ZMUH ; Dar-es-Salaam, 1♂, SMFD No. 8868/210; 1♂ 1♀, FKCP . Thailand, Kohsamui , I.1984, 1♂, leg. E. Talenti, FKCP ; Phuket, 1997, 1♀ (im.) two ecdysis (second and third), FKCP . USA, Hawai, Honolulu , 2♂, SMFD No. 4942/23. Venezuela, San Estaban, Puerto Cabello , 1♀, 19.X.1948, leg. Schöffer, SMFD ; Rancho la Grande , 1♀, II.1955, leg. R. Mertens, SMFD . Vietnam, Annam, Quinhon , X.1934, 1♂, MNHN RS 1537 .?, 1♀, coll. Simon, MNHN No. RS 1473; Au Bord der “Marie“, 1908, 1juv., leg. H. Merton, SMFD No. 5166; 1♂ 2♀, SMFD No. 5168; Aroe, Dobo , Wammer , 22.III.1908, 2♀ 1im., leg. H. Merton, SMFD No. 5165; Elat, Gross-key , IV.1908, 1♂, leg. H. Merton, SMFD No. 5207; Banana, Kongo-Mündung , 1885, 1♂ 1♀ 1im., leg. Hesse, SMFD No. 5124; Kawieng , 1juv., 5.IX.1909, leg. E. Wolf (Hanseatische Südsee Expedition 1909), SMFD ; O. Afrika , 1♀, leg. Knippes, SMFD ; Siam, Chiang-Mai , 1938, 1♀, ZMUH ; 1♂ 1♀, FKCP ; Mt. Makiling , Laguna P.I., 5000 ft, 16.VIII.1932, 2♂, leg. F.C.Hadden, CASC .

DISTRIBUTION. Cosmopolitan.

COMMENTS. Records of this species are so numerous that a complete listing would be unreasonably long. The included records are a representative selection.