Iberomaimetsha rasnitsyni Ortega-Blanco, Perrichot & Engel

Perrichot, Vincent, Ortega-Blanco, Jaime, McKellar, Ryan C., Delclos, Xavier, Azar, Dany, Nel, Andre, Tafforeau, Paul & Engel, Michael S., 2011, New and revised maimetshid wasps from Cretaceous ambers (Hymenoptera, Maimetshidae), ZooKeys 130, pp. 421-453 : 429-433

Iberomaimetsha rasnitsyni Ortega-Blanco, Perrichot & Engel sp. n. Figs 57

Type material.

Holotype MCNA 9928 (♀); paratype MCNA 8765 (♂). All specimens deposited in the Museo de Ciencias Naturales de Álava, Vitoria-Gasteiz, Spain.

Type locality.

Peñacerrada I, Moraza, Burgos Province, Basque-Cantabrian Basin, Spain.

Stratigraphic horizon.

Escucha Formation, Lower Albian, Lower Cretaceous.

Diagnosis.

Mandibles moderately large, each with three teeth, with almost straight margins and teeth (when mandibles opened, teeth facing forward, not curved inward); all flagellomeres of similar length (gradual slight reduction of length); protibial distal margin without comb of stiff setae; tarsal plantulae present on female; cell [1M] about as large as cell [2Cu], both almost square or rhomboidal; 2Rs+M much shorter than 1m-cu and shorter than cell [1Rs]; apicalmost sector of Cu arising at midlength of cell [2Cu]; metasoma elongate, about as long as remainder of body; female ovipositor with apex of second valvula 4-toothed.

Description.

Female (holotype): Body length 2.8 mm. Head rounded in anterior view, length 0.4 mm, width 0.85 mm; compound eyes not especially bulging, oval, large, well separated by 0.43 mm dorsally; basal part of palps not visible but maxillary palps with at least five palpomeres; labial palps with at least three palpomeres; slight, triangular, supra-antennal elevation above torulus, intertorular area flattened; toruli closer to each other than to inner margin of compound eyes; antennae 2.4 mm in length, inserted on frons below midpoint of compound eyes; scape and pedicel short and globular, flagellomeres cylindrical; lengths of antennomeres as follows (all in mm): scape 0.15, pedicel 0.10, flagellomeres 0.22, 0.20, 0.20, 0.20, 0.18, 0.15, 0.14, 0.14, 0.12, 0.12, 0.12, 0.12, 0.12, 0.15; malar space short, 0.15 mm in length. Mesosoma only laterally visible, 1.15 mm in length; pronotum 0.20 mm in length; mesoscutum 0.35 mm in length; mesoscutellum strongly convex, 0.30 mm in length; propodeum slightly rugose, 0.30 mm in length. Only left mid and hind legs are complete, but foreleg can be reconstructed by mixing right and left parts; no visible division of metatrochantellus; tarsi pentamerous, tarsomeres I–IV each with apicoventral plantar lobe (well visible on left metatarsus: Figs 5, 6D); pretarsal claws with small preapical tooth; leg measurements (all in mm): profemur (with protrochantellus) 0.85, protibia 0.50, protarsomeres I–V 0.30/0.15/0.12/?/?; mesofemur 0.75, mesotibia 0.72, mesotarsomeres 0.32/0.15/0.10/0.07/0.12; metafemur 0.95, metatibia 0.90, metatarsomeres 0.38/0.15/0.10/0.08/0.14. Forewing hyaline, length 2.4 mm, maximum width 0.95 mm, densely covered by microtrichia; all veins tubular except for nebulous crossveins 2rs-m and 2m-cu, thus only cells [2Rs] and [2M] pseudo-opened apically (Fig. 7A); 1rs-m not aligned with r-rs; 2m-cu curved at each distal end; veins M and Cu reaching wing margin, with distal end of Cu strongly bent posteriorly; cell [1R1] longer than both cells [1Rs] + [2Rs]; cell [1Rs] about half the surface of cells [1M] and [2Rs]; vein r-rs emerging at two-thirds from base of pterostigma; cell [2R1] short, with vein Rs slightly anteriorly bent distad 2rs-m. Hind wing hyaline and densely covered by microtrichia, length 1.65 mm, maximum width 0.45 mm; five strong hamuli present on costal vein in distal half of wing (Fig. 7B); distal sections of veins Rs and Cu reaching nebulous wing margin; vein A lost beyond cu-a. Metasoma pedunculate, elongate and globoid, length 1.3 mm, greatest height 0.5 mm; sterna hard, convex; no apparent metasomal armature on sternum II or III; cerci very short; ovipositor moderately exserted, first valvula 0.65 mm long (as preserved), second valvula 0.35 mm long, apically with four dorsal teeth, with sheaths covered by very short, sparse setae (Figs. 4, 5A, 5C).

Male: Very similar to female except in following minor differences: Body length 3.63 mm; tarsi without plantar lobes; metasoma slightly longer than mesosoma.

Etymology.

The specific epithet is a patronym honoring Prof. Alexandr P. Rasnitsyn for his numerous important contributions to the study of Hymenoptera and his generous friendship with the authors.