Hydraena accurata Orchymont

PERKINS, PHILIP D., 2014, A revision of the water beetle genus Hydraena Kugelann for southern Africa (Coleoptera: Hydraenidae), Zootaxa 3758 (1), pp. 1-92 : 51-56

Hydraena accurata Orchymont

Figs. 38 (habitus), 40 (aedeagus), 77 (map)

Hydraena accurata Orchymont 1948: 45 .

Type material. Holotype (male): South Africa: Gauteng, Pretoria distr., "Fountain Grove." Labels : "[male symbol]//Fount. Grove. Pret. Dist. 27.4.1910 C. J. Swierstra. // A. d'Orchymont det. Hydraena (s. str.) accurata m.// TYPE [red label]// Holotypus Hydraena accurata d'Orchimon [sic][label with red border]//AEDEAGUS DRAWN BY P. D. PERKINS//DIGITAL IMAGE captured 2012 P. D. Perkins. "

Material examined (4284): South Africa: Eastern Cape Province, Ngagane River Station 6; NAT 420F, marginal vegetation, 29° 46' S, 30° 1' E, 23 vii 1974 (2 AMG); Aliwal North, in tank, 30° 42' S, 26° 43' E, 18-25 viii 1954, H. Andreae (5 SAMC); Transkei, Umtata, 31° 25' S, 28° 45' E, 18 ii 18 iii 1923, R. E. Turner (4 BMNH); Grahamstown, 33° 18' S, 26° 32' E, 1-30 v 1939, J. Omer-Cooper (10 BMNH); Albany District, Gowie's Kloof, 33° 18' S, 26° 32' E, 6 viii 1939, J. Omer-Cooper (4 BMNH); Piggott Bridge Road, 33° 18' S, 26° 32' E, 20 iii 1939, J. Omer-Cooper (7 BMNH); Amatole, Pirie Forest Station, shore washing, 32° 43' S, 27° 17' E, 8 xii 1987, Endrödy-Younga (#2563) (11 TMSA); 10 miles N Matatiele, 30° 10' S, 28° 50' E, 7 iii 1951, Brinck & Rudebeck (211) (12 LUM); near Wildebees River, Small muddy pool, fringed with sedges, at side of road, marginal vegetation out of current, ECR 115, 31° 12' S, 28° 5' E, 26 iii 1993 (44 AMG); Antelope Park Spruit, below bridge, MVIC, ECR 5N, 30° 49' S, 28° 12' E, 4 xii 1990 (1 AMG); Bell river tributary, MVIC, ECR 60D, 30° 43' S, 28° 6' E, 22 iii 1991 (1 AMG); Upper Wildebees River (Gatberg Rd.), marginal vegetation in current, 31° 12' S, 28° 5' E, 12 v 1990 (2 AMG); Nqancule River, trib. at Two Streams, MVIC; ECR 85S, 31° 20' S, 28° 12' E, 25 iii 1991 (9 AMG); KuDidwayo River at Marinus, marginal vegetation in current; ECR 84K, 31° 25' S, 28° 6' E, 25 iii 1991 (6 AMG); Seep near Ben Macdhui, bedrock and small stones, some trailing algae and small plants, elev. 2500 m, 30° 59' S, 28° 15' E, 1 iv 1993 (ECR 177) (2 AMG); Rock pools, summit of Prentjiesberg, rock pools, elev. 1900 m, 31° 8' S, 28° 8' E, 30 iii 1993, F.C. de Moor, H.M. Barber-James, K. Martens (ECR 163) (3 AMG); Mooi River at Keat's Drift, stones in current, 28° 51' S, 30° 30' E, 8 viii 1985 (1 AMG); Naudes Nek, 12 mi ENE Rhodes, 30° 40' S, 28° 5' E, 9 iii 1951, Brinck & Rudebeck (SSAE 220) (15 LUM); Drakensburg, about 13 miles ENE Rhodes, 30° 40' S, 28° 5' E, 9 iii 1951, Brinck & Rudebeck (219) (1 LUM); Eastern Cape, Amatola Mts. 20 km NNE Alice, 32° 47' S, 26° 50' E, 9 iii 1997, Hess & Heckes (26) (23 NMW); Stormberg River south of Bargersdorp, 30° 38' S, 26° 29' E, 16 v 2005, G. L. Challet (1 NMW); Great Brak River Hwy R56 at Schoombee, 31° 27' S, 25° 30' E, 17 v 2005, G. L. Challet (1 NMW). 11 mi. from Matatiele, Kokstad road, in roadside pond, 30° 10' S, 28° 50' E, 28 iii 1954, J. Balfour-Browne (112) (10 BMNH); Albany District, Gowie’s Kloof, 33° 18' S, 26° 32' E, 6 viii 1939, J. Omer-Cooper (4 BMNH); Albany District, Grahamstown, small stream, silty sandy edges, elev. 570 m, 33° 18' S, 26° 32' E, 22 iii 1954, J. Balfour-Browne (stn 90) (1 BMNH); Farm Dam nr. Mooi River, mud with Juncus and Nitella , elev. 1295 m, 31° 0' S, 28° 30' E, 2 iv 1954, J. Balfour-Browne (134) (10 BMNH); Grahamstown, 33° 18' S, 26° 32' E, 1-30 v 1939, J. Omer-Cooper (10 BMNH); Grahamstown, Teafontein Farm, 33° 18' S, 26° 32' E, 5 viii 1939, J. Omer-Cooper (7 BMNH); Kingwilliamstown, Maden dam, boggy pond close to Maden dam, 32° 53' S, 27° 24' E, 25 iii 1954, J. Balfour-Browne (93) (10 BMNH); Kingwilliamstown, Maden dam, edges of Maden dam, deep water, 32° 53' S, 27° 24' E, 25 iii 1954, J. Balfour-Browne (94) (10 BMNH); Lady Frere District, ca. 12 mi from Queenstown on road to Lady Frere, in small grassy dam, elev. 1158 m, 31° 54' S, 26° 53' E, 26 iii 1954, J. Balfour-Browne (98) (10 BMNH); Lahlanguto, nr. Mt. Fletcher, at side stream, elev. 1524 m, 30° 41' S, 28° 30' E, 28 iii 1954, J. Balfour-Browne (108) (2 BMNH); Maclear District, near Ugle, flooded pasture along deep river, elev. 1340 m, 31° 5' S, 28° 22' E, 27 iii 1954, J. Balfour-Browne (105) (2 BMNH); Matatiele, shallow muddy roadside ditch, very weedy with grasses and Juncus , elev. 1463.04 m, 30° 20' S, 28° 49' E, 28 iii 1954, J. Balfour- Browne (111) (29 BMNH); Piggott Bridge Road, 33° 18' S, 26° 32' E, 20 iii 1939, J. Omer-Cooper (1 BMNH); Transkei, Umtata, 31° 25' S, 28° 45' E, 18 ii–18 iii 1923, R. E. Turner (4 BMNH); Free State Province, Drakensberg, Cathedral Peak area, Alpine meadows, elev. 2590.8-2956.56 m, 27° 50' S, 29° 41' E, 19-23 iii 1955, G. Rudebeck (1 LUM); Vaal River System, VAL Stn. 0083F, 26° 40' S, 27° 58' E (3 AMG); Vaal River System, VAL Stn. 0085F, 26° 42' S, 28° 0' E (2 AMG); Vaal River System, VAL Stn. 0098E, 26° 42' S, 27° 53' E (5 AMG); Vaal River System, VAL Stn. 0099H, 26° 42' S, 28° 0' E (5 AMG); Vaal River System, VAL Stn. 0100F, 26° 40' S, 27° 58' E (1 AMG); Vaal River System, VAL Stn. 0112C, 26° 42' S, 28° 0' E (4 AMG); Vaal River System, VAL Stn. 0124B, 26° 42' S, 28° 0' E (1 AMG); Vaal River System, VAL Stn. 0212A, 26° 40' S, 27° 58' E (1 AMG); Vaal River System, VAL Stn. 0238D, 26° 42' S, 28° 0' E (1 AMG); Vaal River System, VAL Stn. 0283B, 26° 40' S, 27° 58' E (1 AMG); Vaal River System, VAL Stn. 0289A, 26° 40' S, 27° 58' E (3 AMG); Vaal River System, VAL Stn. 0320D, 26° 42' S, 28° 0' E (2 AMG); Vaal River System, VAL Stn. 0321, 26° 42' S, 28° 0' E (3 AMG); Vaal River System, VAL Stn. 0333, 26° 42' S, 28° 0' E (2 AMG); Vaal River System, VAL Stn. 0346D, 26° 40' S, 27° 58' E (5 AMG); Vaal River System, VAL Stn. 0361B, 27° 17' S, 28° 29' E (3 AMG); Vaal River System, VAL Stn. 0444H, 28° 20' S, 29° 16' E (3 AMG); Vaal River System, VAL Stn. 0463C, 27° 16' S, 28° 29' E (1 AMG); Vaal River System, VAL Stn. 0467G, 27° 16' S, 28° 29' E (2 AMG); Vaal River System, VAL Stn. 0471JL, 27° 48' S, 28° 47' E (1 AMG); Vaal River System, VAL Stn. 0504U, 28° 16' S, 29° 7' E (2 AMG); Vaal River System, VAL Stn. 0548NN, 28° 16' S, 29° 7' E (1 AMG); Vaal River System, VAL Stn. 0550E, 28° 1' S, 28° 59' E (2 AMG); Vaal River System, VAL Stn. 0579F, 28° 16' S, 28° 51' E (4 AMG); Vaal River System, VAL Stn. 0607E, 27° 21' S, 29° 10' E (1 AMG); Vaal River System, VAL Stn. 0609G, 27° 49' S, 29° 38' E (1 AMG); Vaal River System, VAL Stn. 0628U, 27° 48' S, 28° 40' E (2 AMG); Vaal River System, VAL Stn. 0635F, 28° 20' S, 29° 16' E (1 AMG); Vaal River System, VAL Stn. 0644H, 28° 11' S, 29° 33' E (4 AMG); Vaal River System, VAL Stn. 0687D, 27° 48' S, 28° 47' E (1 AMG); 40 km S Bethlehem, 28° 35' S, 28° 18' E, 30 xii 1993, Wewalka (10) (6 NMW). Harrismith, 28° 16' S, 28° 51' E, 1 iii 1927, R. E. Turner (1 BMNH); Kareefontein nr. Nylstroom, rapid stream over rock and gravel, elev. 1460 m, 28° 25' S, 26° 53' E, 16 iv 1954, J. Balfour-Browne (164) (8 BMNH); O.F.S. nr. Harrismith, farm dam, very grassy, elev. 1676 m, 28° 16' S, 28° 51' E, 7 iv 1954, J. Balfour-Browne (10 BMNH); Vredefort, nr. Homing Spruit, 27° 1' S, 27° 23' E, 29 viii 1949, J. Omer-Cooper (10 BMNH); Gauteng, Hennops River, river, 25° 51' S, 27° 55' E, 28 v 1952, E. J. Pearce (1 BMNH); Suikerbosrand Rivier, near Heidelberg, fast flowing stream with gritty sand, elev. 1530 m, 26° 31' S, 28° 21' E, 8 iv 1954, J. Balfour-Browne (10 BMNH); Gauteng, Natalspruit; REA 085C, Typha capense swamp, 26° 17' S, 28° 8' E (3 AMG); Pretoria, light trap, 25° 45' S, 28° 12' E, 3-5 ii 1957 (5 SANC); Vaal River System, VAL Stn. 45F, 26° 40' S, 27° 57' E (4 AMG); Vaal River System, VAL Stn. 0084E, 26° 40' S, 27° 56' E (2 AMG); Vaal River System, VAL Stn. 0111C, 26° 42' S, 27° 53' E (1 AMG); Vaal River System, VAL Stn. 0122B, 26° 42' S, 27° 53' E (1 AMG); Vaal River System, VAL Stn. 0127B, 26° 40' S, 27° 57' E (1 AMG); Vaal River System, VAL Stn. 0137K, 26° 45' S, 27° 41' E (1 AMG); Vaal River System, VAL Stn. 0142C, 26° 44' S, 27° 36' E (1 AMG); Vaal River System, VAL Stn. 0160H, 26° 47' S, 27° 46' E (1 AMG); Vaal River System, VAL Stn. 0204C, 26° 44' S, 27° 36' E (1 AMG); Vaal River System, VAL Stn. 0237, 26° 44' S, 27° 52' E (1 AMG); Vaal River System, VAL Stn. 0259C, 26° 44' S, 27° 42' E (2 AMG); Vaal River System, VAL Stn. 0271B, 26° 40' S, 27° 56' E (1 AMG); Vaal River System, VAL Stn. 0273B, 26° 45' S, 27° 41' E (1 AMG); Vaal River System, VAL Stn. 0286, 26° 44' S, 27° 36' E (2 AMG); Vaal River System, VAL Stn. 0288A, 26° 45' S, 27° 41' E (1 AMG); Vaal River System, VAL Stn. 0301C, 26° 40' S, 27° 57' E (2 AMG); Vaal River System, VAL Stn. 0312, 26° 44' S, 27° 42' E (1 AMG); Vaal River System, VAL Stn. 0313, 26° 42' S, 27° 59' E (1 AMG); Vaal River System, VAL Stn. 0315B, 26° 44' S, 27° 52' E (2 AMG); Vaal River System, VAL Stn. 0317B, 26° 47' S, 27° 46' E (1 AMG); Vaal River System, VAL Stn. 0318B, 26° 40' S, 27° 56' E (4 AMG); Vaal River System, VAL Stn. 0330A, 26° 41' S, 27° 44' E (1 AMG); Vaal River System, VAL Stn. 0338B, 26° 44' S, 27° 36' E (2 AMG); Vaal River System, VAL Stn. 0391, 26° 41' S, 27° 44' E (1 AMG); Vaal River System, VAL Stn. 0397B, 26° 45' S, 27° 41' E (1 AMG); Vaal River System, VAL Stn. 0415C, 26° 44' S, 27° 42' E (1 AMG); Vaal River System, VAL Stn. 0437A, 26° 44' S, 27° 36' E (1 AMG); Vaal River System, VAL Stn. 0472, 26° 44' S, 27° 42' E (2 AMG); Crocodile River Jukskei River System, Stn. CRO 0264A, 26° 2' S, 28° 6' E (1 AMG); Crocodile River Jukskei River System, Stn. CRO 0264B, 26° 2' S, 28° 6' E (3 AMG); Crocodile River Jukskei River System, Stn.CRO 0265A, 26° 0' S, 28° 2' E (2 AMG); Crocodile River Jukskei River System, Stn.CRO 0269A, 26° 6' S, 27° 58' E (1 AMG); Crocodile River Jukskei River System, Stn. CRO 0270A, 26° 2' S, 27° 59' E (1 AMG); Crocodile River Jukskei River System, Stn. CRO 0272A, 25° 57' S, 27° 57' E (1 AMG); Crocodile River Jukskei River System, Stn. CRO 0272B, 25° 57' S, 27° 57' E (1 AMG); Crocodile River Jukskei River System, Stn. CRO 0273A, 25° 50' S, 27° 55' E (1 AMG); Olifantsvlei, Stn. OLF 0097B, 26° 20' S, 27° 54' E (2 AMG); Olifantsvlei, Stn. OLF 0014E, 26° 18' S, 27° 50' E (1 AMG); Benoni Sewage Works and Dam system, Stn. BEN 8F, 26° 12' S, 28° 18' E (1 AMG); Pretoria, 25° 45' S, 28° 12' E, 26 iv 1951, Brinck & Rudebeck (282) (12 LUM). KwaZulu- Natal Province, Umzimkulu River at The Rocks, marg. veg., NAT 67T-V, 29° 49' S, 29° 31' E, 15 vii 1972 (15 AMG); Tugela River, Jamesons Drift; TUG66Q4, fringing vegetation, 28° 54' S, 31° 2' E (3 AMG); Polela River, Himeville; NAT 056K-M, marg. veg., 29° 44' S, 29° 31' E, 13 vi 1972, F. M. Chutter (1 AMG); Umzimkulu River, the Rocks near Underberg; NAT 067T-V, marg. veg., 29° 49' S, 29° 31' E, 15 vi 1972 (15 AMG); Albany Museum catalog NAT 156Y-Z; Untentweni R., marg. veg., 30° 42' S, 30° 29' E, 7 vi 1972 (1 AMG); Ngagane River Stn 6; NAT432J, marginal vegetation, 27° 46' S, 30° 1' E, 27 viii 1974, H. Metz (3 AMG); Ngwagwane River near Riverside above confluence with Umzimkulu River; NAT 030AB-AD, marg. veg., 30° 5' S, 29° 44' E, 13 vi 1972, H. Metz (32 AMG); Zululand, Nqutu, 28° 12' S, 30° 40' E, 2-15 vi 1951, A. H. Newton (2 SAMC); Zululand, Nqutu, 28° 12' S, 30° 40' E, 2-15 vi 1951, A. H. Newton (1 SAMC); Zululand, Nqutu, 28° 12' S, 30° 40' E, 2-15 vi 1951, A. H. Newton (1 SAMC); Zululand, Nqutu, mountain stream, 28° 12' S, 30° 40' E, 14-30 ix 1950, A. H. Newton (15 BMNH); Cathedral Peak, air plankton, forest, 28° 57' S, 29° 12' E, 18 iii 1976, Endrödy-Younga (#1099) (3 TMSA); Weza, Impetyene Forest, forest, +/ 17 km from Station, shore washing, 30° 38' S, 29° 39' E, 26 xi 1989, Endrödy-Younga & Klimaszewski (#2719) (25 TMSA); Weza, Impetyene Forest, lower forest, shore washing, large and rapid forest river with stony banks, 30° 37' S, 29° 42' E, 19 ii 1989, Endrödy-Younga (#2697) (2 TMSA); Karkloof Grassveld, floating debries, 29° 19' S, 30° 15' E, 12 ix 1989, Endrödy-Younga & Klimaszewski (#2753) (5 TMSA); Tugela River, Stn. TUG 66Q4, 28° 54' S, 31° 2' E ,, W. D. Oliff (8 AMG); Tugela River, Stn. TUG 85Q28, 28° 46' S, 28° 53' E (1 AMG); Buffalo River, near Newcastle, Stn. BF 18Q10, 27° 38' S, 29° 59' E (3 AMG); Umkomaas River, top of Sani Pass, site 1, ex. stones in current, elev. 2865 m, 29° 35' S, 29° 17' E, 18 xii 1996, F. de Moor & C. Dickens (Umk 133 B) (2 AMG); Royal Natal National Park. The Hostel, 28° 41' S, 28° 56' E, 5 iv 1951, Brinck & Rudebeck (263) (2 LUM); Royal Natal National Park. Tugela Valley, 28° 45' S, 28° 53' E, 3 iv 1951, Brinck & Rudebeck (258) (2 LUM); KwaZulu/Natal, Drakensberge: 40 km W Mooi River, Bushman's River unterhalb Nationalpark Camp, elev. 1700 m, 29° 13' S, 30° 0' E, 22 ii 1997, Hess (4 NMW); Drakensberge: 40 km W Mooi River, Bushman's River unterhalb Nationalpark Camp, elev. 1700 m, 29° 13' S, 30° 0' E, 21 ii 1997, Hess (33 NMW); Drakensberg Gardens, 29° 45' S, 29° 15' E, 22 iii 1968, P. J. Spangler (5 USNM); Alfred District, Staffords Post, shallow stony stream, 30° 31' S, 29° 45' E, 29 iii 1954, J. Balfour-Browne (115) (1 BMNH); Bergville Dist., Oliviershoek Pass, clear stream with mud silt bottom, elev. 1524 m, 28° 41' S, 29° 10' E, 7 iv 1954, J. Balfour-Browne (158) (7 BMNH); Bergville, Mkombe River, fine gravel at edges, elev. 1036 m, 28° 41' S, 29° 10' E, 6 iv 1954, J. Balfour-Browne (149) (4 BMNH); Bergville, Mont-aux-sources, farm dam, thick weed at edges, elev. 1220 m, 28° 41' S, 29° 10' E, 6 iv 1954, J. Balfour-Browne (147) (3 BMNH); Bergville, Tugela River, elev. 1158 m, 28° 44' S, 29° 21' E, 6 iv 1954, J. Balfour-Browne (148) (10 BMNH); betw. Lidgetton & Balgowan, trib. of Umgeni River, in marshy edges of river, elev. 1220 m, 29° 45' S, 30° 34' E, 2 iv 1954, J. Balfour-Browne (132) (1 BMNH); Drake Spruit, nr. Estcourt, in side gravel of gritty stream, elev. 1220 m, 29° 17' S, 29° 30' E, 2 iv 1954, J. Balfour-Browne (135) (1 BMNH); Drakensberg Mts., Cathedral Peak, in running water of small brooks, elev. 1980 m, 28° 57' S, 29° 12' E, 3 iv 1954, J. Balfour-Browne (138) (4 BMNH); Harding, Umzimkulwana River, rapid muddy and weedy stream, 30° 35' S, 29° 53' E, 29 iii 1924, J. Balfour-Browne (1 BMNH); nr. Bergville, small unnamed spruit, in side pools among muddy stones and sand, 28° 44' S, 29° 21' E, 4 iv 1954, J. Balfour-Browne (142) (13 BMNH); nr. Bergville, small unnamed spruit, pools in dry bed, slab rock, muddy grit, elev. 1143 m, 28° 44' S, 29° 21' E, 6 iv 1954, J. Balfour-Browne (150) (10 BMNH); nr. Drummond, small stream, stony and weedy pools, elev. 457.2 m, 30° 50' S, 30° 25' E, 1 iv 1954, J. Balfour-Browne (128) (5 BMNH); Tugela River nr. Olivershoek road, washed from stony and sandy banks, elev. 1128 m, 28° 41' S, 29° 10' E, 6 iv 1954, J. Balfour- Browne (6 BMNH); Zululand, Nqutu, mountain stream, 28° 12' S, 30° 40' E, 2-15 vi 1951, A. H. Newton (9 BMNH). Limpopo Province, Transvaal, Shingwidze River, pools in dry sand bed, elev. 550 m, 23° 53' S, 32° 17' E, 26 iv 1954, J. Balfour-Browne (198) (1 BMNH); Mpumalanga Province, Olifants River Stn N5. on farm Aasvogelkrans; OKS 140R, marginal vegetation, 25° 42' S, 29° 16' E, 31 viii 1965 (1 AMG); Klein Olifants River Stn N6 near Botsabelo mission stationOKS 146T, marginal vegetation, 25° 40' S, 29° 24' E, 1 ix 1965 (2 AMG); Olifants River Stn N2, on farm Donkerhoek; OKS 131B, marginal vegetation, 25° 29' S, 29° 16' E, 30 viii 1965 (4 AMG); Van Reenen, Drakensberg, 28° 22' S, 29° 23' E, 1-30 xi 1926, R. E. Turner (2 BMNH); Mpumalanga, Kopees Kraal.Lydenbg., shore washing, 25° 7' S, 30° 12' E, 6 xii 1996, Endrödy-Younga (3255) (1 TMSA); Nelshoogte gallery forest below St., shore washing, 25° 51' S, 30° 53' E, 2 xii 1986, Endrödy-Younga (#2343) (54 TMSA); S. Transvaal, Roberts Drift, Vaal R., sifted flood debris, 27° 2' S, 29° 2' E, 8 x 1973, Endrödy-Younga (#190) (~3,100 TMSA); S. Transvaal, Roberts Drift, Vaal R., flood debris, 27° 2' S, 29° 2' E, 8 x 1973, Endrödy- Younga (#189) (1 TMSA); Bundu Inn, shore washing, 25° 28' S, 28° 55' E, 24 iii 1974, Endrödy-Younga (#304) (2 TMSA); c. Transvaal, Roodeplaat Dam, shore washing, 25° 37' S, 28° 23' E, 14 viii 1974, Endrödy-Younga (#359) (12 TMSA); Uitsuk, window trap, 25° 15' S, 30° 33' E, 15 xii 1986, Endrödy-Younga (#2390) (23 TMSA); Uitsoek, Grootkloof ind. for., intercept trap, 62 days, 25° 15' S, 30° 33' E, 6 ii 1987, Endrödy-Younga (#2422) (19 TMSA); Olifants-Klip River System, Stn. OKS 31W, 25° 57' S, 28° 52' E (1 AMG); Olifants-Klip River System, Stn. OKS 35C, 25° 50' S, 28° 59' E (1 AMG); Olifants-Klip River System, Stn. OKS 44R, 25° 50' S, 28° 59' E (1 AMG); Olifants-Klip River System, Stn. OKS 115U, 25° 34' S, 29° 6' E (1 AMG); Vaal River System, VAL Stn. 0362B, 26° 16' S, 29° 6' E (4 AMG); Vaal River System, VAL Stn. 0363C, 26° 58' S, 29° 14' E (1 AMG); Vaal River System, VAL Stn. 0443R, 26° 51' S, 29° 41' E (1 AMG); Vaal River System, VAL Stn. 0451V, 26° 17' S, 30° 3' E (3 AMG); Vaal River System, VAL Stn. 0452G, 26° 46' S, 29° 32' E (1 AMG); Vaal River System, VAL Stn. 0506J, 26° 55' S, 29° 52' E (1 AMG); Vaal River System, VAL Stn. 0552U, 27° 16' S, 29° 47' E (3 AMG); Vaal River System, VAL Stn. 0656P, 26° 17' S, 30° 3' E (1 AMG); Vaal River System, VAL Stn. 0701J, 26° 51' S, 28° 53' E (1 AMG); Vaal River System, VAL Stn. 0741C, 27° 0' S, 29° 1' E (3 AMG); Vaal River System, VAL Stn. 0765G, 27° 0' S, 29° 1' E (2 AMG); Vaal River System, VAL Stn. 0767H, 27° 5' S, 30° 5' E (1 AMG); Vaal River System, VAL Stn. 0771D, 27° 0' S, 29° 1' E (2 AMG); Vaal River System, VAL Stn. 0773G, 27° 5' S, 30° 5' E (3 AMG); Vaal River System, VAL Stn. 0791E, 26° 33' S, 30° 9' E (3 AMG); Vaal River System, VAL Stn. 0798F, 26° 37' S, 29° 50' E (4 AMG); Vaal River System, VAL Stn. 0799E, 26° 33' S, 30° 12' E (2 AMG); Vaal River System, VAL Stn. 0802F, 26° 17' S, 30° 3' E (1 AMG); Vaal River System, VAL Stn. 0821K, 26° 17' S, 30° 3' E (6 AMG); Vaal River System, VAL Stn. 0825H, 26° 37' S, 29° 50' E (1 AMG); Vaal River System, VAL Stn. 0826F, 27° 1' S, 30° 9' E (2 AMG); Vaal River System, VAL Stn. 0829G, 27° 5' S, 30° 5' E (1 AMG); Vaal River System, VAL Stn. 0829G, 27° 5' S, 30° 5' E (3 AMG); Vaal River System, VAL Stn. 0834R, 26° 42' S, 30° 5' E (3 AMG); Vaal River System, VAL Stn. 0881AC, 27° 5' S, 30° 5' E (2 AMG); Vaal River System, VAL Stn. 0884J, 26° 49' S, 30° 8' E (2 AMG); Vaal River System, VAL Stn. 0894H, 26° 51' S, 29° 41' E (7 AMG); Vaal River System, VAL Stn. 0897E, 26° 33' S, 30° 12' E (2 AMG); Vaal River System, VAL Stn. 0988S, 26° 37' S, 29° 1' E (2 AMG); Vaal River System, VAL Stn. 0990D, 26° 51' S, 29° 41' E (2 AMG); Vaal River System, VAL Stn. 0993D, 27° 5' S, 30° 5' E (3 AMG); Vaal River System, VAL Stn. 1025N, 26° 33' S, 30° 12' E (5 AMG); Vaal River System, VAL Stn. 1041M, 26° 17' S, 30° 3' E (3 AMG); Vaal River System, VAL Stn. 1045L, 27° 5' S, 30° 5' E (3 AMG); Vaal River System, Vaal River where crossed by Ermelo-Lake Chrissie Road, VAL Stn. 1070P, 26° 21' S, 30° 6' E, 21 iii 1960, F. M. Chutter (5 AMG); Vaal River System, VAL Stn. 1089H, 26° 33' S, 30° 12' E (11 AMG); Vaal River System, VAL Stn. 1103K, 26° 37' S, 29° 50' E (7 AMG); Vaal River System, VAL Stn. 1105E, 26° 37' S, 29° 1' E (2 AMG); Vaal River System, VAL Stn. 1111F, 26° 49' S, 30° 8' E (1 AMG); Vaal River System, VAL Stn. 1114C, 26° 51' S, 29° 41' E (4 AMG); Vaal River, Stn. VD1: VAL 1119E, marginal vegetation, 26° 17' S, 30° 3' E (2 AMG); Vaal River System, VAL Stn. 1126N, 27° 5' S, 30° 5' E (2 AMG); Vaal River System, VAL Stn. 1138E, 26° 37' S, 29° 1' E (1 AMG); Vaal River System, VAL Stn. 1145J, 26° 33' S, 30° 12' E (1 AMG); Vaal River System, VAL Stn. 1146R, 27° 1' S, 30° 9' E (6 AMG); Vaal River System, VAL Stn. 1151J, 27° 1' S, 30° 9' E (4 AMG); Vaal River System, VAL Stn. 1169G, 26° 49' S, 30° 8' E (3 AMG); Vaal River System, VAL Stn. 1182H, 27° 1' S, 30° 9' E, (9 AMG); Vaal River System, VAL Stn. 1191C, 26° 58' S, 29° 14' E, (2 AMG); Vaal River System, VAL Stn. 1212E, 26° 17' S, 30° 3' E, (2 AMG); Vaal River System, VAL Stn. 1339S, 26° 17' S, 30° 3' E, (4 AMG); Vaal River System, VAL Stn. 0735M, 26° 34' S, 30° 12' E, (1 AMG); Vaal River System, VAL Stn. 0733Y-AA, 26° 49' S, 30° 8' E, (3 AMG); Olifants-Klip River System, Stn. OKS 0005G, 25° 40' S, 29° 10' E, (1 AMG); Olifants-Klip River System, Stn. OKS 0007Y, 25° 34' S, 29° 6' E, (1 AMG); Olifants-Klip River System, Stn. OKS 0092B, 25° 37' S, 29° 13' E, (1 AMG); Olifants-Klip River System, Stn. OKS 0111D, 25° 40' S, 29° 10' E, (1 AMG); Pretoria Sewage Works, Stn. PTA 46A, 25° 42' S, 28° 9' E (1 AMG); Pretoria Sewage Works, Stn. PTA 47, 25° 42' S, 28° 9' E (1 AMG); 11 mi. SE Middelburg, 31° 30' S, 25° 1' E, 27 iii 1968, P. J. Spangler (25 USNM). nr. Standerton, pool by muddy stream, algae and grass, 26° 58' S, 29° 13' E, 8 iv 1954, J. Balfour-Browne (161) (10 BMNH); Transvaal, Ermelo Dist., Warburton, elev. 1740 m, 26° 15' S, 30° 25' E, 7 xii 1948, J. Omer-Cooper (3 BMNH); Vaal River System, VAL Stn. 743X, 26° 17' S, 30° 3' E, 22 vii 1959, F. M. Chutter (10 BMNH); Van Reenen, Drakensberg, 28° 22' S, 29° 23' E, 1-30 xi 1926, R. E. Turner (1 BMNH); Waterval River, fast stony stream, rather muddy, 26° 36' S, 29° 2' E, 8 iv 1954, J. Balfour-Browne (162) (5 BMNH). Western Cape Province, Hex River Mts., 7 km SW Ceres, Oberhalb Michell's Pass-Strasse: Restionaceen-Sumpf in feuchter Rinne durchBerg-Fynbos, elev. 500 m, 33° 23' S, 19° 19' E, 25 ii 1997, Hess (1 NMW); Du Toits Mts., 8 km SE Franschhoek, 33° 55' S, 19° 8' E, 28 ii 1997, Hess & Heckes (1 NMW). Lesotho: Drakensberg, Sani Pass Valley, floated tussocks, 29° 36' S, 29° 20' E, 10 iii 1976, Endrödy-Younga (#1063) (16 TMSA); Drakensberg, Sani Pass Valley, shore washing, 29° 36' S, 29° 20' E, 14 iii 1976, Endrödy-Younga (#1084) (4 TMSA); Drakensberg, Sani Pass Valley, shore washing, 29° 36' S, 29° 20' E, 10 iii 1976, Endrödy-Younga (#1062) (31 TMSA); Drakensberg, Sani Pass Valley, water collection, 29° 36' S, 29° 20' E, 8 iii 1976, Endrödy-Younga (#1053) (14 TMSA); Luteni River, flood debris, 29° 24' S, 29° 29' E, 12 iii 1976, Endrödy-Younga (#1073) (6 TMSA); Qachas Nek, 20 miles N Matatiele, 30° 7' S, 28° 41' E, 7 iii 1951, Brinck & Rudebeck (213) (7 LUM); Mamathes, 5 mi. E. Teyateyneng, 29° 9' S, 27° 50' E, 29 iii 1951, Brinck & Rudebeck (SSAE 252) (4 LUM); Nazareth M. S., 20 mi ESE Maseru, 29° 30' S, 27° 30' E, 24 iii 1951, Brinck & Rudebeck (SSAE 245) (3 LUM); Basutoland, Hensley's Dam, 8 mi. SW Leribe, 29° ' S, 28° ' E, 30 iii 1951, Brinck & Rudebeck (SSAE 254) (19 LUM); Dikolobeng River, 11 mi. NE Quthing, river, 30° 23' S, 27° 43' E, 16 iii 1951, Brinck & Rudebeck (SSAE 237) (2 LUM).

Differential diagnosis. This species is very similar in habitus, size, and ventral characters to H. neoaccurata ( Figs. 38 , 39 ). The aedeagi, which also show a relationship, cannot be confused with those of other species, and must be examined for reliable determinations ( Figs. 40, 41 ).

Description. Size: holotype (length/width, mm): body (length to elytral apices) 1.60/0.70; head 0.25/0.42; pronotum 0.40/0.53, PA 0.45, PB 0.50; elytra 1.02/0.70. Dorsum of head dark brown to piceous; pronotum brown around dark brown macula that extends, in lighter color, to margin in middle; elytra and legs testaceous to brown; maxillary palpi testaceous.

Frons punctures ca. 1-2xef, slightly larger and denser near eyes than medially; interstices medially weakly shining, 1–3xpd, laterally mcr and dull, 0.5–1xpd. Clypeus mcr laterally, very finely sparsely punctulate medially. Mentum finely sparsely punctulate, interstices shining; postmentum contiguously punctulate. Genae raised, weakly shining, without posterior ridge.

Pronotum laterally slightly emarginate behind middle; anterior margin straight behind eyes, emarginate behind frons, scintilla absent; punctures on disc much larger and deeper than those of frons, interstices shining, 1–2xpd, punctures slightly larger and denser at anterior and posterior; PF1 absent; PF2 very shallow; PF3 moderately deep; PF4 very shallow; area between PF3 and PF4 weakly convex.

Elytra weakly arcuate laterally; summit of posterior declivity at ca. midlength; lateral explanate margin narrow; on basal 1/3 punctures slightly smaller than largest pronotal punctures, punctures becoming finer and more widely spaced toward posterior. Intervals not raised, very weakly shining, on basal 1/3 ca. 1–2xpd, as are interstices between punctures of a row. Apices in dorsal aspect conjointly rounded, in posterior aspect margins forming deep angle with one another.

Ratios of P2 width and plaque shape (P2/w/l/s) ca. 2/1.5/4/5. P1 laminate; median carina sinuate in profile. P2 l/w ca. 4/1, sides parallel, apex blunt, apex raised slightly above mesoventral intercoxal process. Plaques very narrow, parallel, weakly raised, margins not strongly demarcated, located at sides of moderately deep median depression. Metaventrite without midlongitudinal carina between intercoxal process and median depression. AIS width at slightly arcuate posterior margin ca. 2x P2. All tibiae slender, protibiae slightly arcuate, meso- and metatibiae straight. Abdominal apex slightly asymmetrical; last tergite oval, with depression in distal ½, and minute off-center tooth at distal margin. Female (microslide mount, n=1): last tergite moderately sharply rounded, with rather deep apicomedian notch, ca. 10 hooked setae set back from margin ca. ½ length of setae, setae not quite contiguous, median gap rather wide; gonocoxite not midlongitudinally divided, apical margin moderately sharply rounded, transverse ridge evident at border of microreticulation; spermatheca type simple C shape.

Distribution. A very common species, widely distributed in eastern and northeastern South Africa, and Lesotho. There are two disjunct localities known, at the Cape ( Fig. 77 ).

Figures

FIGURE 38. Hydraena accurata, holotype, dorsal and lateral habitus.

FIGURE 39. Hydraena neoaccurata, holotype, dorsal and lateral habitus.

FIGURES 40–41. Aedeagi of holotypes.

FIGURES 77–78. Geographical distributions of Hydraena species.