Hlavaciellus bifoveatus, Jałoszyński, Paweł, 2011

Jałoszyński, Paweł, 2011, Hlavaciellus Jałoszyński, 2006: eleven new species, detailed morphology and systematic position within Cephenniini (Coleoptera, Staphylinidae, Scydmaeninae), Zootaxa 2763, pp. 1-33 : 12-13

Hlavaciellus bifoveatus sp. n.

( Figs. 9 , 24 , 31, 32)

Type material. Holotype: MALAYSIA: ɗ, " SABAH: Crocker Ra. \ 1600 m, km 51 rte \ Kota Kinabalu– \ Tambunan, 18.V.87 \ Burckhardt - Löbl" [white, printed], " HLAVACIELLUS \ bifoveatus m. \ det. P. JAŁOSZYŃSKI, '09 \ HOLOTYPUS " [red, printed] ( MHNG). Paratype: MALAYSIA: ɗ, same data as holotype ( PCPJ).

Diagnosis. Vertex in males with pair of shallow transverse lateral impressions broadly separated in middle; antennomere III nearly twice as long as broad.

Description. BL 2.05–2.18 (2.11). Body of male ( Fig. 24 ) strongly convex, pigmentation light brown, vestiture yellowish.

Head ( Fig. 9 ) very large, HL 0.35–0.38 (0.36), HW 0.53–0.55 (0.54); vertex long and convex, its anterior margin broadly subtriangular and delimited from supraantennal tubercles and frons by pair of large, shallow and subtriangular impressions broadly separated in middle; frons and clypeus confluent, distinctly convex; supraantennal tubercles prominent, indistinctly separated from median part of frons; eyes large, strongly convex and coarsely faceted. Punctures and setae as in Fig. 9 . Antennae very long and slender, as long as 0.78 BL, gradually thickened up to antennomere XI, uniformly covered with thin, long, dense and suberect setae, AnL 1.60–1.68 (1.64), antennomere I twice as long as broad; II much narrower and shorter than I, 1.3x as long as broad; III slightly broader and much longer than II, 1.8x as long as broad; IV slightly larger than III, 1.8x as long as broad; V distinctly broader but only minimally longer than IV, 1.7x as long as broad; VI distinctly longer and broader than V, 1.6x as long as broad; VII–IX nearly equal in length and width, each distinctly larger than VI and about 1.6x as long as broad; X slightly larger than IX, 1.6x as long as broad; XI slightly broader than X, distinctly shorter than IX–X together.

Pronotum in dorsal view subrectangular with weakly convex anterior margin, nearly equally broad from base up to anterior fourth, PL 0.63–0.68 (0.65), PW 0.85–0.88 (0.86); sides strongly rounded in anterior fifth or fourth, then nearly straight and parallel with barely discernible constriction near posterior third; hind angles slightly sharp but broadly rounded; posterior margin deeply bi-emarginate; ante-basal pits very shallow and diffused, each equally distant from posterior and lateral margin of pronotum; lateral edges not swollen. Pronotal disc glossy, in middle covered with very fine but distinct punctures separated by spaces 1–1.5x as wide as puncture diameters, punctures close to margins of pronotum are smaller. Setae long, moderately dense, suberect to erect.

Elytra oval, relatively long, broadest near anterior third, EL 1.08–1.13 (1.10), EW 0.93, EI 1.16–1.22; basal pit on each elytron small but distinct, located close to middle of elytral base within adsutural half of elytron; subhumeral lines very distinct, as long as 0.3x EL, very slightly divergent, each developed as sharply marked border between higher humeral region and lower adsutural area; apices of elytra separately rounded. Punctures on basal third of each elytron only slightly more distinct than those on pronotum, nearly equally dense but slightly larger, gradually reducing in diameter and depth toward lateral margins and apices; setae similar to those on pronotum. Hind wings entirely reduced, absent.

Legs slender and long, tibiae straight or nearly straight.

Metaventrite with very large and deep postmesocoxal impressions, occupying 2/3 of its length.

Aedeagus (Figs. 31, 32) very slender, AeL 0.43; median lobe with subtriangular, rounded apical part; internal armature asymmetrical; parameres in lateral view moderately broad, with moderately strongly curved apices.

Female. Unknown.

Distribution. East Malaysia: Borneo, Sabah.

Etymology. The name refers to the pair of impressions on the head.

Remarks. See remarks for H. smetanai .

Figures

FIGURES 4 – 9. Modifications of head in frontal view. 4, Hlavaciellus peninsularis sp. n. 5, H. cirrus sp. n. 6, H. angustifrons sp. n. 7, H. decoratus sp. n. 8, H. smetanai sp. n. 9, H. bifoveatus sp. n. Scale bars: 0.1 mm.

FIGURES 24 – 26. Dorsal view of holotypes of Hlavaciellus, males. 24, H. bifoveatus sp. n. 25, H. metrios sp. n. 26, H. miser sp. n. Scale bars: 1 mm.