Hammatoderus lingafelteri, Botero, Juan Pablo & Santos-Silva, Antonio, 2017

Botero, Juan Pablo & Santos-Silva, Antonio, 2017, Four new species, taxonomic, and nomenclatural notes in Hammatoderus Gemminger & Harold, 1873 (Coleoptera, Cerambycidae, Lamiinae), Zootaxa 4231 (3), pp. 377-397 : 385-387

Hammatoderus lingafelteri sp. nov.

( Figs. 11–15 )

Description. Holotype male. Integument dark-brown, black on some areas; antennomeres gradually reddishbrown toward distal segments, with distal area of antennomeres III–X dark-brown.

Head. Frons moderately coarsely vermiculate throughout; with yellowish-white pubescence, distinctly not obscuring integument, except narrow, dense band with yellow pubescence close to eyes; with short, erect, sparse, yellowish setae. Area between antennal tubercles slightly vermiculate close to frons, smooth toward upper eye lobes; with yellowish-white pubescence not obscuring integument. Area between upper eye lobes with small depression close to antennal tubercles, slightly vermiculate toward posterior margin; with yellowish-white pubescence not obscuring integument. Remaining surface of vertex vermiculate (slightly less coarse than on frons); with yellowish-white pubescence not obscuring integument, except narrow band close to upper eye lobes with narrow, dense band with yellow pubescence. Area between eyes and antennal tubercles with narrow, dense band with yellow pubescence. Tempora finely vermiculate behind upper eye lobes, gradually smoother toward tumid area close to lower eye lobes and finely asperate close to prothorax; with yellowish pubescence, not obscuring integument, except narrow, dense band with yellow pubescence close to eye. Antennal tubercles elevated, close each other on base; minutely, abundantly punctate, interspersed with some fine punctures; with yellowish-white pubescence not obscuring integument. Coronal suture distinct from clypeus to prothoracic margin. Genae 0.65 times width of lower eye lobe; with transverse, shallow sulcus about middle; minutely, abundantly punctate, except smooth, triangular area close to eye; with yellowish pubescence toward ventral side, yellow toward dorsal side close to eye, gradually glabrous toward apex. Gulamentum finely, semi-circularly striate toward prothoracic margin, depressed about central area; with row of coarse punctures on elevated area close to mentum; with yellowish-white pubescence not obscuring integument from depressed area to mentum, glabrous toward prothoracic margin. Postclypeus transversely, moderately deeply sulcate close to frons, except centrally; basal half coplanar with frons, distal half sloped; with yellowish pubescence on coplanar area, not obscuring integument, interspersed with long, erect, yellowish setae; sloped region with short, sparse, yellowish setae centrally, long, erect, yellowish setae laterally. Labrum convex, sub-coplanar with anteclypeus on basal third, sloped on distal 2/3; centrally almost glabrous, laterally with long, erect, yellowish setae directed forward. Distance between upper eye lobes 0.15 times length of scape; distance between lower eye lobes 0.55 times length of scape. Antennae 3.15 times elytral length (antennomere X of left antenna broke after middle, antennomere XI of right antennae broke, apparently after middle), reaching elytral apex at basal fifth of antennomere VI; scape rugose, abundantly, minutely punctate, with yellowish pubescence not obscuring integument, denser on base of cicatrix, with short, erect, sparse, yellowish setae ventrally; antennomere III with short, erect, yellowish and brownish setae on ventral side; antennomeres III–IV with moderately sparse, yellowish pubescence on base and on distal quarter before apex; antennomere IV with some short, erect, yellowish setae; antennomere V with yellowish pubescence along outer side; antennomeres VI–XI with yellowish pubescence, distinctly not obscuring integument; antennal formula (ratio) based on antennomere III: scape = 0.56; pedicel = 0.12; IV = 0.93; V = 0.83; VI = 0.82; VII = 0.85; VIII = 0.87; IX = 0.85; X = 0.85; XI broke.

Thorax. Prothorax with large, conical lateral tubercle, curved upward, with moderately acute apex, about 1.8 times as long as distance between upper eye lobes. Pronotum with two transverse sulci, basalmost sub-straight, narrow, distalmost curved; with a large, elevated gibbosity centrally, from apex of basal quarter to after middle and two transverse gibbosities at each side of apex of central gibbosity; surface coarsely vermiculate, mainly between sulci, including on gibbosities; with yellowish pubescence between vermiculation, partially obscuring integument; with moderately long, erect, sparse, yellowish setae. Sides of prothorax with small, sparse, shiny tubercles between base and lateral tubercle, sparsely vermiculate under lateral tubercle (gradually smoother toward ventral side), smoother between lateral tubercle and distal margin; with yellowish pubescence except on small tubercles and vermiculations. Prosternum finely, sparsely, transversely striate; with yellowish pubescence, almost obscuring integument. Prosternal process longitudinally sulcate; with short, sub-erect, abundant, yellowish setae. Mesosternum with yellowish pubescence centrally interspersed with short, erect, abundant, yellowish setae. Mesosternal process longitudinally tumid, but not forming tubercle on base; with yellowish pubescence interspersed with short, erect, yellowish setae. Mesepisternum, metepisternum and metasternum with yellowish pubescence, partially obscuring integument. Mesepimeron surrounded by band with decumbent, yellow pubescence, centrally with sparser yellowish pubescence. Scutellum with dense, yellowish pubescence. Elytra. Humerus slightly projected; basal seventh with small, abundant, shiny, glabrous tubercles; remaining surface coarsely, abundantly punctate, gradually sparser toward apex; with yellowish pubescence, damaged in the holotype, but apparently with areas with dense patches; apex uniformly rounded with spine at sutural angle. Legs. Femora with yellowish pubescence interspersed with short, erect, yellowish setae. Protibiae with yellowish pubescence (less so on dorsal basal half), interspersed with short, erect, yellowish and brownish setae; with dense, sub-erect, yellowish setae ventrally, obscuring integument. Mesotibiae with dense, sub-erect, yellowish setae obscuring integument dorsally and ventrally (distinctly shorter on basal third); laterally with yellowish pubescence not obscuring integument, interspersed with short, erect, yellowish and brownish setae. Metatibiae with dense, suberect, yellowish setae on distal half of dorsal and ventral side (shorter on ventral side); remaining surface with yellowish pubescence not obscuring integument, laterally interspersed with yellowish setae.

Abdomen. Ventrites with yellowish pubescence partially obscuring integument, except narrow distal band with yellow, short, decumbent setae on ventrites I–IV; with short, erect, sparse, yellowish setae, distinctly more abundant on ventrite V; distal margin of ventrite V distinctly concave.

Dimensions (Holotype male). Total length, 33.9; prothoracic length, 5.0; anterior prothoracic width, 5.9; posterior prothoracic width, 6.3; widest prothoracic width (between apices of lateral tubercles), 8.5; humeral width, 10.5; elytral length, 24.5.

Type material. Holotype male from MEXICO, Guerrero: Zapotitlán Tablas , X.1963, M. Rosas col. ( MNRJ, former Campos Seabra collection).

Etymology. The specific name is a genitive patronym in honor of Steven W. Lingafelter, for his contributions to the knowledge of the cerambycid fauna and for his hospitality and support toward the first author during his visit to the USNM.

Remarks. Hammatoderus lingafelteri sp. nov. differs from H. ornator (Bates, 1885) as follows: elytral pubescence pattern, even damaged, distinctly different; base of the elytra with small tubercles more abundant; pronotum distinctly vermiculate between sulci; meso- and metasternum and abdominal ventrites without patches of pubescence. In H. ornator (see photographs of the holotype at Bezark 2016), the elytral pubescence forms distinct patches, the base of the elytra has sparser small tubercles, the pronotum is not vermiculate between sulci, and meso- and metasternum and abdominal ventrites have patches of pubescence. It differs from male of H. olivescens ( Dillon & Dillon, 1941) ( Figs. 37–38 ) by the body wider (slender in H. olivescens ), the pronotum coarsely vermiculate between sulci (not vermiculate in H. olivescens ), and the elytral pubescence not forming distinct patches (forming in H. olivescens ). Hammatoderus lingafelteri sp. nov. also differs from H. antonkozlovi sp. nov. by the frons coarsely rugose, the mesosternal process without tubercle and the apex of elytra uniformly rounded with spine at sutural angle. In H. antonkozlovi the frons is not rugose, the mesosternal process is tuberculate and the elytral apex is obliquely truncate with spine at outer angle.

Figures

FIGURES 11 – 15. Hammatoderus lingafelteri sp. nov., holotype male: 11, dorsal view; 12, ventral view; 13, lateral view; 14, head, frontal view; 15, tubercle of lateral mesosternal process, lateral view.

FIGURES 32 – 38. 32, Hammoderus lacordairei, holotype female, dorsal view. 33, Hammoderus inermis, holotype male, dorsal view. 34, Hammoderus sallei, syntype female, dorsal view. 35 – 36, Plagiohammus confusor, holotype male: 35, dorsal view; 36, labels. 37 – 38, Plagiohammus olivescens, holotype male: 37, labels; 38, dorsal view.