Gugnelve butleri Ramos-González & Parra, 2024

Ramos-González, Mario I., Venegas-González, María Francisca, Zamora-Manzur, Carlos & Parra, Luis E., 2024, Revision of the genus Digonis (Lepidoptera, Geometridae): new species and new genera, ZooKeys 1216, pp. 173-200 : 173-200

Gugnelve butleri Ramos-González & Parra sp. nov.

Figs 2 Q, R , 11

Erosina cervinaria ( Blanchard, 1852); Butler 1882: 347 pl. 16, fig. 4; Bartlett-Calvert 1886: 331; Angulo and Casanueva 1981: 12 [misidentification of Ennomos cervinaria Blanchard].

Type material.

Holotype. Chile • 1 male; Cordillera Prov., Santiago, Guayacan ; XII-1950; “ Holotype Gugnelve butleri ” [red handwritten label], “ Mirg- 017 ” [genitalia slide] ( MZUC-UCCC) . Allotype. Chile • 1 female, Cordillera Prov., Santiago, Guayacan ; I-1951; “ Allotype Gugnelve butleri ” [red handwritten label], “ Mirg- 018 ” [genitalia slide] ( MZUC-UCCC) . Paratypes. Chile — Limarí Prov . • 2 males, Ovalle, Quebrada Seca ; 28 / 29-XI-1997; ZSM . — Cordillera Prov. • 4 females and 1 male, same data as Holotype ; • 1 female and 1 male, same data as allotype .

Other material examined.

Chile — Cachapoal Prov. • 1 female; Termas de Cauquenes ; 11-I-1953; “ Museo 4516 ” [Museum ID]; MNNC .

Additional records.

Chile. — Cachapoal Prov. • Mountains of the hacienda of Cauquenes; January; T. Edmonds leg.; NHMUK ( Butler, 1882) — Talca Prov. • Parque Natural Tricahue; - 35.70869903, - 71.08727243; 5 - I- 2020; observed by Vicente Pantoja and submitted to iNaturalist in: https://inaturalist.mma.gob.cl/observations/37300513. • Cipreses hydroelectric plant; - 35.8099272997, - 70.8359745; 17 - XII- 2022; observed by César Picar and submitted to iNaturalist in: https://iNaturalist.mma.gob.cl/observations/144651054. • Cipreses hydroelectric plant; - 35.8096577, - 70.8353316; 10 - I- 2024; observed by César Picar and submitted to iNaturalist in: https://inaturalist.mma.gob.cl/observations/196337433.

Diagnosis.

As for the genus.

Description.

Male (Fig. 2 Q ). Head: antennae slightly serrated; palpi short, subequal to the eye diameter, porrect; frons and vertex covered with juxtaposed brownish scales. Thorax: Patagia covered with elongated scales of same color as background; tegulae covered with piliform scales of same color as background; tibial spur formula 0-2 - 4. Forewings: slightly falcate, subtriangular; fovea absent; background color yellowish brown; antemedial band dark yellowish brown, externally bordered by pale brown, oblique, extending toward the medial area, near the end of the discal cell where it curves toward the antemedial sector to the costa, forming an acute arc; postmedial band dark yellowish brown, externally bordered by pale brown, slightly sinuous, oblique, and slightly curved, extending to the subapical sector; discal spot not visible. Hindwings: suboval; background color pale brown without bands; discal spot not visible. Male genitalia (Fig. 11 A, B ). Uncus conical; socii slightly setose; gnathos V-shaped with sinuous arms, expanded apex with 11 small denticles of varying sizes toward the center; valvae sub-rhomboidal, costa and anterior margin sclerotized; transtilla fork-shaped; saccus suboval; juxta sub-halberd-shaped, laterally extended into a pair of tri-spined strongly sclerotized processes; anellus sclerite weakly defined. Tubular aedeagus, straight; vesica armed with cornuti, formed by 11 grouped spines of different sizes.

Female (Fig. 2 R ). Similar to male but with simple filiform antennae, dark yellowish brown falcate forewings, and slightly crenulated pale brownish gray subrectangular hindwings. Female genitalia (Fig. 11 C ). Ductus bursae half the length of the corpus bursae; antrum funnel-like; corpus bursae membranous, subpyriform, signum with a long blade-like ridge on a sclerotized patch; posterior apophyses three times longer than anterior apophyses.

Etymology.

The specific name is dedicated to Arthur Gardiner Butler, an ornithologist, arachnologist, and entomologist of the 19 th century, for his contribution to the study of Lepidoptera in Chile.

Distribution.

This species is found between the provinces of Limarí and Talca. It is distributed in parts of the biogeographic provinces of Coquimbo and Santiago, Central Chilean subregion; Maule, Subantarctic subregion, in the Andean region.

Flight period.

Specimens were captured in November, December, and January. There are no records for other months.

Figures

Figure 2. Habitus of Digonis adults A Digonis aspersa (male in dorsal view) B Digonis aspersa (male brown-morpho in dorsal view) C Digonis aspersa (male brown-morpho in ventral view) D Digonis aspersa (female in dorsal view) E Digonis cervinaria, stat. rev. (male olivaceous-morpho in dorsal view) F Digonis cervinaria, stat. rev. (male olivaceous-morpho in ventral view) G Digonis cervinaria, stat. rev. (male fuscous-morpho in dorsal view) H Digonis cervinaria, stat. rev. (female in dorsal view) I Digonis cervinaria, stat. rev. (female olivaceous-morpho in ventral view) J Digonis cervinaria, stat. rev. (female brown-morpho in ventral view) K Digonis punctifera (male in dorsal view; photo courtesy of A. Hausmann) L Digonis punctifera (female in dorsal view) M Digonis gungnir Ramos-González & Parra, sp. nov. (holotype in dorsal view) N Digonis apocrypha Ramos-González & Parra, sp. nov. (holotype in dorsal view) O Digonis apocrypha Ramos-González & Parra, sp. nov. (holotype in ventral view) P Phasmadigonis alba, comb. nov. (male in dorsal view) Q Gugnelve butleri Ramos-González & Parra, sp. nov. (holotype in dorsal view) R Gugnelve butleri Ramos-González & Parra, sp. nov. (allotype in dorsal view). Scale bar: 10 mm.

Figure 11. Genitalia of Gugnelve butleri Ramos-González & Parra, sp. nov. (holotype and allotype) A male genitalia in ventral view B aedeagus in lateral view C female genitalia in ventral view. Scale bars: 1 mm.