Grylloblatta chandleri Kamp, 1963

Schoville, Sean D. & Graening, G. O., 2013, Updated checklist of the ice-crawlers (Insecta: Grylloblattodea: Grylloblattidae) of North America, with notes on their natural history, biogeography and conservation, Zootaxa 3737 (4), pp. 351-378 : 356-360

Grylloblatta chandleri Kamp, 1963

U.S.A., California. Lassen Co. Ice caves near Eagle Lake, 1.5 miles WSW of Spaulding Tract, elev. 5,180 ft., 5 adult males and 1 adult female coll. by H.P. Chandler, T.L. Rodgers and J.W. Kamp (paratypes), from 12 June to Sep. 1951 (Kamp 1953, 1963); 1 adult female coll. by J.W. Kamp (allotype), 9 July 1951 (Kamp 1963); on moist rocks in cave, 1 adult male coll. by V.D. Roth (holotype), 5 Sep. 1959; 5,000 ft. elev., 1 nymph coll. by H.P. Chandler, 0 2 May 1952 (CAS); Griffith Meadows Ice Caves, 1962 (Kamp 1970), also referred to as Griffith Ranch Ice Caves by Peck (1973); unidentified ice cave (Kamp 1973); Lassen National Forest, Gray's Flat, Township 32 north, Range 7 east, Section 33, northeast 1/4, southwest ¼, N 40.628 W -121.18932 1825m elev., on rock talus after onset of snowstorm, 1 nymph coll. by D.R. Cluck, 0 6 Nov. 2005 (Schoville & Roderick 2010); Lassen Peak, NE slope in Devastated Area, 1 adult female coll. by S.D. Schoville and R. Y. Dudko, 17 May 2013.

Shasta Co. North Christmas Tree Cave, 1 nymph coll. by G. O. Graening, 21 April 2012; Subway Cave, Lassen National Forest, 1 nymph coll. by G. O. Graening, 0 3 Nov. 2012.

Tehama Co. Wilson Lake Ice Cave, Mt. Lassen (Kamp 1970); Wilson Lake Ice Cave, 1615m elev., 1 adult female coll. by M.D. Terry, T.H. Ogden, and K.J. Jarvis, 22 Aug. 2001 (BYU: Jarvis & Whiting 2006); 1 nymph coll. by K.J. Jarvis, G.T. Svenson, and J. Jones, 18 June 2002 (BYU: Jarvis & Whiting 2006).

Kamp (1963) suggested G. chandleri is widespread in the Modoc Plateau and Basin Ranges of Eastern California, and later stated that “…almost every ice cave explored between Eagle Lake and the Medicine Lake Highlands contains populations of Grylloblatta . ” (Kamp 1973).

Grylloblatta chirurgica Gurney, 1961

U.S.A. Washington. Skamania Co. Ape Cave, 1 adult male and 3 adult females coll. by H. Reese and family (holotype, allotype and 2 paratypes), 22–28 Nov. 1959 (Gurney 1961); 1 adult female and 1 nymph coll. by R.L. Crawford and C.M. Senger, 14–30 May 1983 (BMUW: Jarvis & Whiting 2006); 2 adults coll. by R.L. Crawford and C.M. Senger, 27 Oct. to 11 Nov. 1983 (BMUW); 4 specimens (CMNH); Bat Cave, 1,160 ft. elev., 1 adult female coll. by W.R. Halliday, 0 1 Nov. 1960 (BMUW); 1 adult coll. by R.L. Crawford and C.M. Senger, 22 July to 0 6 Aug. 1983 (BMUW); 1 adult coll. by R.L. Crawford and C.M. Senger, 18 Nov. to 8 Dec. 1983 (BMUW); H. Reese, 0 1 March 1961; 1 adult male coll. by D. Nielsen, 18 Nov. 1972; Bean Creek, Mt. St. Helens National Volcanic Monument, 1 adult coll. by K. Johnson, 15–28 June 1983 (BMUW); Beaver Cave, 1,520 ft. elev., 1 adult coll. by R.L. Crawford and C.M. Senger, 24 July 1983 (BMUW); R.L. Crawford and C.M. Senger, 27 Oct. to 19 Nov. 1983 (BMUW); Butte Camp A on talus slope, Mt. St. Helens National Volcanic Monument, 1 adult coll., 0 5 Oct. 1983 (BMUW); Cave 6 miles NE Cougar, 3 specimens coll. by R. Zack, 31 Jan. 1976 (WSU); Dollar and A Dime Cave, 1,290 ft. elev., 1 adult female coll. by R.L. Crawford and C.M. Senger, 18 Nov. to 0 8 Dec. 1983 (BMUW); 1 adult female coll. by R.L. Crawford and C.M. Senger, 21 July to 0 6 Aug. 1983 (BMUW); Gremlin Cave, 2,980 ft. elev., 1 adult male coll. by R.L. Crawford and C.M. Senger, 07–23 Oct. 1983 (BMUW); 1 adult and 2 nymphs coll. by R.L. Crawford and C.M. Senger, 0 5 July 1983 (BMUW); Little People Cave, 2,840 ft. elev., 1 adult and 1 nymph coll. by R.L. Crawford and C.M. Senger, 23 July to 0 5 Aug. 1983 (BMUW); Manhole Cave, 3,000 ft. elev., 1 nymph coll. by R.L. Crawford and C.M. Senger, 0 8 Oct. 1983 (BMUW); N. Butte Camp, 46.175 N 122.225 W, 4,520 ft. elev., 1 adult coll. by R.L. Crawford, 0 3 Nov. 1982 (BMUW); 1 adult coll. by D. Mann, 1– 16 Oct. 1981 (BMUW); 2 adults coll. by C.J. Becker, 12–24 June 1983 (BMUW); Lake Cave, 1,900 ft. elev., 1 adult female (paratype) coll. by W.R. Halliday, 28 Dec. 1958 (Gurney 1961); 1 nymph coll. by R.L. Crawford and C.M. Senger, 0 7 Dec. 1983 (BMUW); 1 nymph coll. by R.L. Crawford and C.M. Senger, 25 May to 27 June 1983 (BMUW); 1 nymph coll. by R.L. Crawford and C.M. Senger, 23 Nov. 1973 (BMUW); Mt. St. Helens, Timberline Rd., 3,800 ft. elev., 1 specimen coll. by A. Smetana, 4 July 1978 (CMNH); Ole’s Cave, Mt. St. Helens south end lava beds, 1 specimen coll. by R. Zack, 0 2 February 1978 (WSU); 1 adult female coll. by C.M. Senger, 25 Aug. 1965 (BMUW); 2 adults and 2 nymphs coll. by R.L. Crawford and C.M. Senger, 21 May to 26 June 1983 (BMUW); 3 specimens coll. by R.L. Crawford and C.M. Senger, 27 Oct. to 12 Nov. 1983 (BMUW); Nielsen’s Cave, 2 nymphs coll. by W.R. Halliday, 1958 (Gurney 1961); caves on west flank of Mt. St. Helens (Kamp 1970); NE slope Mt. St. Helens, 1,520–1,620 m elev., 1 adult female and 1 nymph coll. by D.H. Kavanaugh, 5 Aug. 1974 (CAS); Pumice Plain, nymphs coll. by P.S. Sugg and J.S. Edwards, 1984–1985 (Sugg & Edwards 1998); NW side of Lava Dome, Mt. St. Helens, observation of C.H. Anderson, Jr. and M.R. Vining, Sep. 1997 (Anderson & Vining 1999); Prince Albert Cave, 1,200 ft. elev., 1 adult coll. by R.L. Crawford and C.M. Senger, 18 Nov. to 0 8 Dec. 1983 (BMUW); 2 adults coll. by. R.L. Crawford and C.M. Senger, 22 July to 0 2 Aug. 1983 (BMUW); Spider Cave, 5.1 miles NE of Cougar, 2,840 ft. elev., 1 adult and 1 nymph coll. by R.L. Crawford and C.M. Senger, 05–31 July 1983 (BMUW); 1 nymph coll. by R.L. Crawford, 0 6 Aug. 1977 (BMUW); 1 adult coll. by G. Ericson, 12 Nov. 1966 (BMUW); Spirit Lake at Bear Creek, 3,200 ft. elev., 1 adult coll. by A. Smetana, 0 6 July 1974 (BMUW); Surprise Cave, 3,240 ft. elev., 1 adult female coll. by R.L. Crawford and C.M. Senger, 23 July 1983 (BMUW; Jarvis & Whiting 2006).

Yakima Co. Chinook Pass, 0.25 mi. SE of Chinook Pass Picnic Area, 5,360ft. elev., Z10 E 613220 N 5192760, 1 adult male coll. by E.A. Lisowski, 10 Nov. 2003 (BYU; Jarvis & Whiting 2006); Goat Peak Trail, 5,880 ft. elev., 1 adult male coll. by E.A. Lisowski, 11 Sep. 2002 (BYU; Jarvis & Whiting 2006); White Pass at summit, under rock in forest north of US 12, specimens coll. by J.C. Bergdahl (J.C. Bergdahl, pers. comm.).

Probable records. Washington. Kittitas Co. Mount Baldy, base of large rocks on ridge, 14.3 miles S-SE Ellensburg, UTM 10Z, 696040mE, 5186640mN, 2,200 ft. elev., 1 adult and 3 nymphs coll. by E.A. Lisowski, 21 Dec. 1997; Wilson Creek, 10 miles N. Ellensburg, 1 nymph coll. by S.D. Smith, 17 Dec. 1982 (BMUW).

Lewis Co. Eagle Peak Trail, 11.7 miles NW Packwood, UTM 10Z, 591340mE, 5177730mN, 2,960 ft. elev., coll. by E.A. Lisowski, 11–25 February 1995 (BMUW); Laughing Water Creek, 46.753' N 121.548 'W, 2,600 ft. elev., 9 nymphs coll. by J. Putera, 08–17 February 1988 (BMUW); 1 adult female and 9 nymphs coll. by J. Putera, 17–24 February 1988 (BMUW); 1 adult male, 7 adult females and 10 nymphs coll. by J. Putera, 24 February to 0 5 April 1988 (BMUW); 1 adult coll. by J. Putera, 15 April to 0 6 May 1988 (BMUW); 1 adult female coll. by J. Putera, 28 Sep. to 27 Oct. 1988 (BMUW); 2 adult males, 2 adult females and 40 nymphs coll. by J. Putera, 27 Oct. to 0 1 Dec. 1988 (BMUW); coll. by J. Putera, 11 Dec. 1988 to 0 5 Jan. 1989 (BMUW); 2 adult males and 1 nymph coll. by J. Putera, 15–22 Jan. 1989 (BMUW); 1 nymph coll. by J. Putera, 15–24 Jan. 1989 (BMUW); 3 adults and 3 nymphs coll. by J. Putera, 21 Feb. to 23 March 1989 (BMUW); 5 adults coll. by J. Putera, 22 Jan. to 21 Feb. 1989 (BMUW); Tongue Creek Trail, 46.426N 121.780W, 2,900 ft. elev., 1 nymph coll. by J. Putera, 17–28 Jan. 1988 (BMUW). Pierce Co. 0.1 mi. East Route 410 Entrance, 13.5 miles S SE Greenwater, Mt. Rainier National Park, UTM 10Z, 611640mE, 5203560mN, 2,800 ft. elev., 2 nymphs coll. by E.A. Lisowski, 22 Dec. 1995 (BMUW); Bear Head Mountain Summit, 47.023N 121.814W, 6,000-6,089 ft. elev., 1 adult coll. by R. Crawford, 15 Aug. 1982 (BMUW); Carbon River, 14.7 miles SE Wilkeson, UTM 10Z, 590400mE, 5201740mN, 2,600 ft. elev., 1 specimen coll. by E.A. Lisowski, 0 4 Feb. 1995 (BMUW); Upper Spray Park, 1 adult coll. by J.S. Edwards, 0 2 June 1979; Westside Road, 2.7 miles N-NE main road and 7.1 miles E-NE Ashford, UTM 10Z, 585120mE, 5181140mN, 2,920 ft. elev., 1 adult coll. by E.A. Lisowski, 0 7 Jan. 1996 (BMUW); 14.7 miles SE Wilkeson, UTM: 10Z, 590400mE, 5201740mN, 1 adult coll. by E.A. Lisowski, 4 Feb. to 11 March 1995 (BMUW).

Yakima Co. 0.5 mi SE Windy Point, 7.5 miles W Tieton, UTM 10Z, 660000mE, 5172300mN, 2,520 ft. elev., 1 specimen coll. by E.A. Lisowski, 27 Feb. 2000 (BMUW); Forest Service Road 417 at elev. 2,960 ft., 9.2 miles W- SW Tieton, UTM 10Z, 657560mE, 5171040mN, 1 specimen coll. by E.A. Lisowski, 24 June 2001; Little Rattlesnake Creek., middle of section 16, 11.9 miles S-SE Cliffdell, UTM 10Z, 656080mE, 5183700mN, 2,480 ft. elev., 1 specimen coll. by E.A. Lisowski, 10 March 2001; Tieton Reservoir Road near Goose Egg Mountain, 1.7 miles SE Rimrock, UTM 10Z, 645720mE, 5167940mN, 2,760 ft. elev., 1 specimen coll. by E.A. Lisowski, 31 July 1997; Thunder Creek, 2.9 mi. S-SE Goose Prairie, UTM 10Z, 633560mE, 5190360mN, 5,480 ft. elev., 2 specimen coll. by E.A. Lisowski, 14 Oct. 2001; 2.7 mi. S-SE Goose Prairie, UTM 10Z, 633460mE, 5190660mN, 1 specimen coll. by E.A. Lisowski, 0 3 Nov. 2001; US 12 between 1st and 2nd Tieton River Bridge, 7.9 miles W Tieton, UTM 10Z, 659300mE, 5173060mN, 2,280 ft. elev., 1 specimen coll. by E.A. Lisowski, 24 Jan. 1998 (BMUW).

Surveys showed that G. chirurgica occupied disturbed habitat in Mt. St. Helens within 4 years of the eruption in 1980 (Sugg & Edwards 1998). This species is sympatric with G. sp. nov. “Sawyer’s Ice Cave #1" at Chinook Pass ( Jarvis & Whiting 2006). Its range also approaches G. sp. nov. “Mt. Rainier” and G. sp. nov. “Trout Lake Caves.”

Grylloblatta gurneyi Kamp, 1963

U.S.A. California. Siskiyou Co. All the following records are from lava tubes in or near Lava Beds National Monument: Merrill Ice Cave, 4,740 ft., 1 adult male and 1 adult female coll. by J. W. Kamp and D. H. Kistner (holotype and allotype), 20 Feb. 1960 (USNM; Kamp 1963); 1 adult male coll. by V.D. Roth (paratype), 1954 (Kamp 1963); R. Crawford, 24 Nov. 1989 (BMUW); 1 adult female and 3 nymphs coll. by K.J. Jarvis, G.J. Svenson, and J. Jones, 0 2 June 2002 (BYU; Jarvis & Whiting 2006); 1 adult female coll. by Fuhrman, K.J. Jarvis, G.J. Svenson, and J. Jones, 0 1 Jan. 2003 (BYU; Jarvis & Whiting 2006); 1 adult female and 1 nymph coll. by S.D. Schoville and R. Y. Dudko, 12 May 2013; 3-level Ice Cave, 1 adult female coll. by C.M. Senger, 23 June 1984 (BMUW); 2 adults coll. by C.M. Senger, 23–29 June 1984 (BMUW); 1 adult male coll. by T. Briggs, 19 June 1973 (CAS); 1 specimen coll. by H. MacDonald et al., 1997; Angleworm Cave/Lost Pinnacle Cave, 1 nymph coll. by R.L. Crawford, 27 May to 23 June 1990 (BMUW); R. Crawford, 24 Nov. 1989 (BMUW); Arch Cave, R. Crawford, 20 Nov. 1989 (BMUW); Catacombs Cave, 5,000 ft. elev., 5 adults coll. by R.L. Crawford, 21 Nov. to 9 Dec. 1989 (BMUW); Cox Ice Cave, 4,580 ft. elev., 1 adult female coll. by R.L. Crawford, 27 May 1990 (BMUW); 1 adult female coll. by K.J. Jarvis, G.J. Svenson, and J. Jones, 20 June 2002 (BYU; Jarvis & Whiting 2006); Deep Ice Cave, NE of McCloud, 5,560 ft. elev., 1 specimen coll. by D. Cowan, and 3 nymph coll. by C.M. Senger, 23–29 June 1984 (BMUW); Frozen River Ice Cave, coll. by S.B. Peck (Peck 1973); Heppe Cave, SE grotto, 5,280 ft. elev., 1 nymph coll. by C.M. Senger, 22–29 June 1984 (BMUW); Indian Wells Ice Cave, 3 miles south of Merrill, 2 adult males and 4 adult females coll. by J.W. Kamp (paratypes), Sep. 1951 and 1954 (Kamp 1963); 2 nymphs coll. by S.D. Schoville and R. Y. Dudko, 12 May 2013; Mushpot Cave, 1 nymph coll. by R.L. Crawford, 21 Nov. to 9 Dec. 1989 (BMUW); Skull Cave, D. Cowan, 23 Nov. 1989 (BMUW).

Probable records. California. Siskiyou Co. Bray Ice Cave, 2 adult females coll. by T. Briggs, 19 June 1973; Lava cave near Hambone, 1 adult male coll. by T. Briggs, 28 May 1977 (CAS); Jack Jones Ice Cave (Halliday 1962); Jake Bell Ice Cave(s) (Halliday 1962); Mayfield Ice Cave (Halliday 1962); Mt. Shasta, “high elevations on snowbanks,” coll. by S.B. Peck (Peck 1973); Mt. Shasta, 1 specimen, coll. unknown, 21 May 1978 (BME: not identified to species); Constance Creek, Mt. Shasta, 1 adult male, 1 adult female and 5 nymphs coll. by S.D. Schoville and R. Y. Dudko, 13 May 2013; Starr Cave (Halliday 1962, Peck 1973).

Grylloblatta marmoreus Schoville, 2012

U.S.A. California. Siskiyou Co. Bigfoot Cave, Marble Mountains Wilderness Area, 1 adult male G. O. Graening and Guy Graening (holotype), 0 3 Sep. 2010 (Schoville 2012); Brokedown Palace Cave, collections in 1977 and 1978, B. Her (Suggett 1982); Marble Valley Guard Station, 16 specimens coll. by D. Hemphill, M. Winning, and R.L. Suggett, 0 1 Feb. 1980 (Suggett 1982); 3 specimens coll. by M. Winning and R.L. Suggett, 15 Feb. 1981 (Suggett 1982); trail to Marble Valley from Lover’s Camp, 1 specimen coll. by R.L. Suggett, 23 Feb. 1980 (Suggett 1982); Marble Valley, 30+ specimens coll. by S.D. Schoville and R. Y. Dudko, 14 May 2013; Planetary Dairy Cave, 1 nymph coll. by G. O. Graening and D. Weaver, 4 July 2010 (Schoville 2012).

Grylloblatta oregonensis Schoville, 2012

U.S.A. Oregon. Josephine Co. Oregon Cave, Oregon Caves National Monument, 1 adult female coll. by J. Roth (holotype), 15 June 2010 (Schoville 2012); 1 adult, 10 Oct. to 15 Nov. 1992 (BMUW); one nymph, 30 Nov. 2006 (Schoville 2012); one nymph, 11 July 2008 (Schoville 2012); one nymph, 14 Jan. 2009 (Schoville 2012); one nymph coll. by S.D. Schoville, 20 March 2010 (Schoville 2012); 1 specimen coll. by C. Davis, April 1983; Kincaid’s dancehall, 5 nymphs, 31 May 1993 (BMUW); Wedding Cake room, 15 nymphs, 31 May 1993 (BMUW); 1 adult, 10 May 1993 (BMUW).

This species is currently a single-site endemic and potentially cave-restricted (Schoville 2012). It is parapatric or sympatric with G. siskiyouensis at Oregon Caves National Monument.

Grylloblatta rothi Gurney, 1953

U.S.A. Oregon. Deschutes Co. Happy Valley, 15 to 20 miles south of Sisters on Century Drive, 6,450 ft. elev., 1 adult male coll. by V.D. Roth (holotype), 12 Sep. 1918 (Gurney 1953); McKenzie Pass, Willamette National Forest, 1 specimen coll. by P.S. Ward, 19 May 1985 (BME); 2 specimens (CMNH; Kamp 1973); 44.25˚ N 121.81 ˚W, 5,220 ft. elev., 1 adult female coll. by J.S. Edwards, 24 June 1985 (BMUW); N 44.260 W -121.809, 1 specimen coll. by SSE, 23 June 1981; 3 adult males, 12 adult females and 8 nymphs coll. by K.J Jarvis, G.J. Svenson, and J. Jones, 24 June 2002 (BYU; Jarvis & Whiting 2006); Edison Ice Cave (Kamp 1970).

Hood River Co. Mt. Hood, junction of Phlox Point and Timberline roads, 5,900 ft. elev., 2 adult males and 5 adult females coll. by J.W. Kamp, 18 June 1970 (Kamp 1973); mileage post 68 on Mt. Hood Loop Road, one nymph (NMNH); Tilly Jane Creek, Mt. Hood National Forest, N 45.39896 W -121.64813 1,772 m elev., 1 adult male coll. by S.D. Schoville, 18 June 2009 (CAS).

Klamath Co. Crater Lake, 2 adult females coll. by J.E. Elsea (paratype), 27-28 Nov. 1936 (CAS; USNM; Elsea 1937, Gurney 1953).

Probable records. Oregon. Deschutes Co. 1 mi. NW Mt. Bachelor, 1 specimen coll. by J.G. Todd, 12 March 1983 (CMNH); Bachelor Butte (Kamp 1973); Belknap crater lava fields (T. 15S, R. 8E, USGS Three Sisters Quad.), 5,384 ft. elev., 5 adult males and 12 adult females coll. by J.W. Kamp, 18 June 1970 (Kamp 1973); one adult female coll. by J.M. Taylor, 18 July 1971 (NMNH); Crossbill Ice Cave, ~ 20 mi. SE of Bend, 2 adult females coll. by B. McGregor, 5 May 2013; Ice Cave N Newberry Crater; caves near Bend (Peck 1973); Lava Top Butte Ice Cave, 1 adult obs. by B. McGregor, April 2012; South Ice Cave at Fort Rock, 1 adult female coll. by J.W. Kamp (Kamp 1970).

Linn Co. Lebanon (Kamp 1963). Marion Co. Boca Cave near Triangulation Peak, one adult obs. by B McGregor, Nov. 2011; Silver Falls State Park near Stayton, N 44.87877 W

-122.6590, several larvae obs. by M. Leppin, 13 Dec. 2008.

There appears to be extensive sympatry of G. rothi and G. sculleni , including sites at Edison Ice Cave (Kamp 1970) and throughout the Three Sister region (Kamp 1973). Kamp (1973) identified a third species at the Belknap lava fields that is sympatric with both G. sculleni and G. rot hi at this site. However, the species description was never published. He also suggested samples from Mt. Hood form a new species (Kamp 1973), but our genetic and morphological analysis assign this population to G. rot hi.

Grylloblatta scudderi Kamp, 1979

Canada. British Columbia. Whistler Mountain, Garibaldi Provincial Park, 1,951 m elev., 1 adult male, 1 adult female, and 9 nymphs coll. by L. Bartlett (holotype and allotype), 13 July 1970 (Kamp 1979).

Probable records. Canada. British Columbia. Blowdown Pass in Stein Valley, near Pemberton, N 50.3626 W- 122.15, 1 adult coll. by R. Bennett and D.R. Copley, 15 Sept. 2012 (RBCM); Grouse Mountain (Spencer 1945); Mission Ridge above Seton Portage, near Lillooet, N50.76356 W-122.169806, 2,174m elev., 4 specimens coll. by R. Bennett, C.R. Copley and D.R. Copley, 3 August 2011 (RBCM); Mount Garibaldi (Kamp 1979).

Kamp (1973) lists the Grouse Mountain record as “doubtful” due to extensive glaciations in this area.

Grylloblatta sculleni Gurney, 1937

U.S.A. Oregon. Benton Co. Mary’s Peak (T. 12 S., R. 7 W., Section 28 NW1/4 of the NE1/4 of the NW1/4), 3,960 ft. elev., 1 adult female coll. by J. LaBonte, 3 April 1993 ( Jarvis & Whiting 2006); 1 adult female coll. by J. LaBonte, 3 April 1995; 3 adult females coll. by J. LaBonte, 10 Feb. 1996; Mary’s Peak, 14 miles west of Corvallis (Kamp 1973).

Deschutes Co. Scott Camp, Three Sisters, 6,600 ft. elev., 3 adult females and 6 nymphs coll. by H.A. Scullen and R. Rider (holotype and paratypes), 12 July 1936 (CAS; USNM; Gurney 1937); 1 adult female and 2 nymphs coll. by H.A. Scullen and G. Ferguson (paratype), 0 6 Aug. 1935 (Gurney 1937); 1 adult female coll. by H. Scullen, 1 Aug. 1959 (CAS; Gurney 1961); Sunshine Meadow, 6,600 ft. elev., 1 adult female coll. by H. Crowell, 1 Aug. 1959 (Gurney 1961); Belknap crater lava fields, Scott Pass, south slope of North Sister, 6,500 ft. elev., 4 adult males coll. by J.W. Kamp, 22 July 1970 (Kamp 1973); Edison Ice Cave, Deschutes National Forest, 5,200 ft. elev., 2 adult males and 6 adult females (Kamp 1973); McKenzie Pass, 2 adult males coll. by J.W. Kamp, 18 July 1971 (CMNH; Kamp 1973).

Probable record. Linn Co. Quartzville Creek ½ mile E of Green Creek (Sec. 26, T 11S, R 4E), 1,900 ft. elev., 1 nymph coll. by H. Hacker, 14 Nov. 1959 (Gurney 1961).

See comments on sympatry with G. rot hi. Kamp (1973) proposed subspecific status to a morphological variant of G. sculleni from Edison Ice Cave, but the description was never published.

Grylloblatta siskiyouensis Schoville, 2012

U.S.A. Oregon. Josephine Co. Big Tree Loop Trail, Oregon Caves National Monument, 1 adult female coll. by A. Y. (holotype), 6 March 2007 (Schoville 2012); Oregon Cave exit, 1 nymph, 23 April 1994 (BMUW); 1 adult female coll. by Annette Rasch, 21 Nov. 2003 ( Jarvis & Whiting 2006); Blind Leads Cave, 3 nymphs coll. by S.D. Schoville, 20 March 2010 (Schoville 2012); slope above Oregon Cave, N 42.098 W -123.407, 2 nymphs coll. by S.D. Schoville, 20 March 2010 (Schoville 2012).

Probable records. Oregon. Josephine Co. Oregon Caves National Monument, one larva coll. by J. Roth, 26 Feb. 1989; 5 nymphs coll. by J. Roth, 27 Feb. 1993 (BMUW); one adult coll. by J. Roth, 0 2 April 1993 (BMUW); Mt. Ashland (Kamp 1973, Schoville 2012).

Grylloblatta siskiyouensis is parapatric or sympatric with G. oregonensis at Oregon Caves National Monument (Schoville 2012).

Grylloblatta washoa Gurney, 1961

U.S.A. California. El Dorado Co. Echo Summit, 4 miles south of Meyers on Highway 50 near Echo Peak, 7,382 ft. elev., 1 adult male (holotype) and 3 nymphs coll. by W.J. Gertsch and V.D. Roth, 9 Sep. 1959 (Gurney 1961); Camp Sacramento, 2 nymphs coll. by T.S. Briggs, 3 Jan. 1981 (CAS), and 1 specimen coll. by R.L. MacDonald, 19 Dec. 1962 (BME); Smith Lake, Desolation Wilderness, N 38.85681 W -120.18805 2,678 m elev., 1 adult female and 1 nymph coll. by S.D. Schoville, 23 July 2005 (Schoville & Roderick 2010); Susie Lake, Desolation Wilderness, N 38.88574 W -120.13450 2,520 m elev., 3 adult males, 11 adult females, and 9 nymphs coll. by S.D. Schoville, 12 May 2007 (Schoville & Roderick 2010).

Nevada Co. Carpenter Ridge, 39º25´N 120º20´W 2,610m. elev., 1 adult female coll. by P.S. Ward, 30 June 2000 ( Jarvis & Whiting 2006), and another specimen coll. by P.S. Ward on 5 July 2002 (BME).

Placer Co., SE Barker Pass on Ellis Peak Trail, N 39.06661 W -120.22999 2,479 m elev., 1 adult male, 12 adult females, and 4 nymphs coll. by S.D. Schoville, 0 2 June 2007 (Schoville & Roderick 2010). Sierra Co. Sierra Buttes (Kamp 1973); 16 nymphs coll. by S.D. Schoville and R. Y. Dudko, 11 May 2013.

Probable records. Plumas Co. Mt. Elwell, 60 miles southeast of Mt. Lassen (Kamp 1973).

Sierra Co. Avalanche Cave, near Henness Pass Road, 1 specimen coll. by D. Cowan, mid-1980s.