Goliathopsis ferreroi Antoine, 1991

Xu, Hao & Qiu, Jian-Yue, 2020, Three new synonyms within the flower chafer genus Goliathopsis Janson, 1881 (Coleoptera: Scarabaeidae: Cetoniinae) from China, Zootaxa 4789 (1), pp. 91-131 : 107-110

Goliathopsis ferreroi Antoine, 1991

( Figs. 9 , 17–18 , 30–31 , 46–47 , 73–76 , 95–96 , 138–143 , 164 )

Goliathopsis ferreroi Antoine, 1991: 10 (type locality: Lamphun, N. Thailand), plate I, fig. 3 (parameres), plate III, fig. 3 ♂; Sakai & Ikeda 1996: 40 (Doi Pui, Chiang Mai; Huay Gaew, Chiang Mai; Mae Tha, Lampan, Thailand); Sakai & Nagai 1998: 156 (Mae Tha, Lamphun, Thailand), plate 2, figs. 38- 1 ♂, 38- 2 ♀; Krajčík 1999: 39; Krajčík 2012: 116.

Type material. This species was described based on 27 males and 27 females, and all of them were collected from the type locality, except one male from Laos. Only holotype and six paratypes were examined, the remaining should be preserved in private collections as stated in the original publication ( Antoine 1991). Holotype ( Figs. 138–141 ) is labeled: Thailand, Lamphun, 12.VI.[19]90 [white label] // HOLOTYPE ♂, Goliathopsis ferreroi, Ph. ANTOINE det. 1990 [white label]. Six examined paratypes (3♂♂, 3♀♀): 1♀ (allotype, MNHN, Figs. 142–143 ), Thailand, Lamphun, 12. VI. [19]90 [white label] // ALLOTYPE ♀, Goliathopsis ferreroi, Ph. ANTOINE det. 1990 [white label]; 1♀ ( BMNH), Thailand, Lamphun, 15.VII.[19]90 [white label] // Paratype , Goliathopsis ferreroi, Ph. ANTOINE det. 1990 [white label]; 1 ♂ ( BMNH), Thailand, Lamphun, 12. VI. [19]90 [white label] // Paratype , Goliathopsis ferreroi Ph. ANTOINE det. 1990 [white label]; 1 ♂ ( BMNH), Laos B.N.Pot, 13-V-1920, R. Vitalis de Salvaza // Laos, Ban Nam Pot, 13-V-1920 [yellow label] // Bourgoin Coll. B.M. 1938-252. // Pilinurgus despectus Westw. // Paratype , Goliathopsis ferreroi, Ph. ANTOINE det. 1990 [white label]; 1♂, 1♀ ( KSCJ), 1990.VI.27, Meatha , Lampang Prov., N. Thailand // Paratype , Goliathopsis ferreroi Ph. ANTOINE det. 1990 [white label].

Other material examined (27♂♂, 12♀♀). Thailand: 1♂, 1♀ ( MNHN), [19]75.VII.20, Ngao, Thailand, G. Minet // Goliathopsis ferreroi, S. Rojkoff det. 2010 ; 1♂ ( BMNH, Krajčík Coll.), 1997.IV.16-V.1, Pai , [Mae Hong Son], M. Zyka leg. ; 2♂♂ ( MYNU), 1986.VI, Chiang Mai // Goliathopsis ferreroi Antoine, 1991 , det. K. Sakai, 2013 ; 1♀ ( MYNU), 1984.VIII.2, Doi Pui , Chiang Mai // Goliathopsis ferreroi Antoine, 1991 , det. K. Sakai, 2013 ; 3♂♂ ( MYNU), 2016.V, Chiang Mai, native leg ; 3♂♂, 3♀♀ ( KSCJ), 1990.VII.9, Meatha , Lampang Prov., N. Thai- land ; 2♂♂, 2♀♀ ( KSCJ), 1990.VI.27, Meatha , Lampang Prov., N. Thailand ; 4♂♂, 4♀♀ ( KSCJ), 1985.VIII.18, Huay Gaew , Chiang Mai, N. Thailand ; 1♂, 1♀ ( KSCJ), 1984.VII.5, Doi Pui , Chiang Mai, N. Thailand ; 1♂ ( KSCJ), 1984.VI.27, Doi Pui , Chiang Mai, N. Thailand ; 1♂ ( KSCJ), 1984.VII.23, Doi Pui , Chiang Mai, N. Thailand ; 1♂ ( KSCJ), 1984.VII.15, Doi Pui , Chiang Mai, N. Thailand ; 1♂ ( KSCJ), 2000.IV.23, Huay Gaew , Chiang Mai, N. Thailand, T. Chiku leg. ; 3♂♂ ( KSCJ), 2013.V.5, Chiang Mai University, Chiang Mai, Thailand , Toshiaki Chiku leg. ; 3♂♂ ( KSCJ), 2013.V.13, Huay Gaew, Chiang Mai, Thailand , T. Chiku leg.

Diagnosis. Body length 11.5–12.5 mm; width 5.8–6.7 mm. Dorsal surface and pygidium clad with light yellow and brown tomentum ( Figs. 73, 75 ). Pronotum with four indistinct, small, tomentose maculae ( Fig. 75 ); disc darker, lateral portion light yellow or brown ( Figs. 73, 75 , 95–96 , 138, 140, 142 ); midline on pronotum and scutellum yellow, distinct. Elytron usually with 4 light brown maculae: a small one on posthumeral area; a small one on the mediodiscal area near sutural costa; a small one on the median portion of lateral declivity, sometimes enlarged ( Fig. 96 ); a large one on the postdiscal areas near sutural costa. Few other small tomentose maculae scattered on declivity of elytra ( Fig. 140 ). Disc of elytron with U- and drop shaped, setiferous punctures ( Fig. 9 ); setae very long. Pygidium with rounded punctures ( Figs. 30–31 ); longitudinal ridge without tomentum in few individuals ( Fig. 31 ). Cephalic horns of male slender, not constricted at base; without tomentum ( Fig. 17 ), sometimes with few tomentum on inner surfaces and proximal portion ( Fig. 18 ). Male genitalia small ( Figs. 46–47 ), proximal and distal parts of parameres expanded in apical view, medially constricted, outer line of distal part obtusely angled, apex sharp; parameres almost flat in lateral view, basal piece expanded.

This species resembles Goliathopsis despectus , and the differences are already provided in the diagnosis section of G. despectus . Goliathopsis ferreroi is also similar to G. velutinus , but can be differentiated by inner surface of cephalic horns glabrous, at most tomentose in proximal portion ( Figs. 17–18 ; totally tomentose in G. velutinus , Fig. 21 ). Besides, the shape of parameres of these three species is distinctly different as shown in Figs. 42, 46, 48 .

Distribution. Thailand, Laos.

Figures

FIGURE 164. Known distribution of Goliathopsis Janson, 1881 species. Syncretic markers indicate identical or very close localities; type localities of the known species are marked, that of G. despectus (Westwood, 1873), G. cervus Janson, 1881, and G. esquiroli Pouillaude, 1913 are missing because the precise type localities are unknown.

FIGURES 1–12. Characters of Goliathopsis Janson, 1881 species. 1–4, mouthparts of Goliathopsis duponti Antoine, 1991: 1, epipharynx; 2, mandibles; 3, maxillae; 4, labium. 5–12, setiferous punctures on the disc of elytron: 5, G. cervus Janson, 1881; 6, G. despectus (Westwood, 1873), paralectotype of G. capreolus Gestro, 1888 in BMNH; 7, G. duponti Antoine, 1991; 8, G. esquiroli Pouillaude, 1913; 9, G. ferreroi Antoine, 1991; 10, G. gressitti Ruter, 1978; 11, G. lameyi Fairmaire, 1893; 12, G. velutinus Pouillaude, 1913.

FIGURES 13–21. Cephalic horns of Goliathopsis Janson, 1881 species (dorsal view). 13, G. cervus Janson, 1881; 14, G. despectus (Westwood, 1873), paralectotype of G. capreolus Gestro, 1888 in MNHN; 15, G. duponti Antoine, 1991; 16, G. es- quiroli Pouillaude, 1913; 17–18, G. ferreroi Antoine, 1991; 19, G. gressitti Ruter, 1978; 20, G. lameyi Fairmaire, 1893; 21, G. velutinus Pouillaude, 1913.

FIGURES 22–37. Pygidium of Goliathopsis Janson, 1881 species (male in left, female in right; posterior view). 22–23, G. cer- vus Janson, 1881; 24–25, G. despectus (Westwood, 1873), lectotype and paralectotype of G. capreolus Gestro, 1888 in MSNG; 26–27, G. duponti Antoine, 1991; 28–29, G. esquiroli Pouillaude, 1913; 30–31, G. ferreroi Antoine, 1991; 32–33, G. gressitti Ruter, 1978; 34–35, G. lameyi Fairmaire, 1893; 36–37, G. velutinus Pouillaude, 1913.

FIGURES 38–53. Male genitalia of Goliathopsis Janson, 1881 species (apical and lateral view). 38–39, G. lameyi Fairmaire, 1893; 40–41, G. esquiroli Pouillaude, 1913; 42–43, G. velutinus Pouillaude, 1913; 44–45, G. duponti Antoine, 1991; 46–47, G. ferreroi Antoine, 1991; 48–49, G. despectus (Westwood, 1873), paralectotype of G. capreolus Gestro, 1888 in MNHN; 50–51, G. gressitti Ruter, 1978; 52–53, G. cervus Janson, 1881.

FIGURES 73–76. Habitus of Goliathopsis ferreroi Antoine, 1991 (dorsal and ventral view). 73–74, male; 75–76, female.

FIGURES 89–102. Habitus of Goliathopsis Janson, 1881 species (male in left, female in right; oblique view). 89–90, G. cervus Janson, 1881; 91–92, G. duponti Antoine, 1991; 93–94, G. esquiroli Pouillaude, 1913; 95–96, G. ferreroi Antoine, 1991; 97–98, G. gressitti Ruter, 1978; 99–100, G. lameyi Fairmaire, 1893; 101–102, G. velutinus Pouillaude, 1913.

FIGURES 138–143. Type specimens of Goliathopsis ferreroi Antoine, 1991 in MNHN. 138–141, male holotype and labels; 142–143, female paratype (allotype) and labels.