Fortipalpa yucatanica Kasparyan & Ruíz-Cancino, 2007

Supeleto, Fernanda A., Santos, Bernardo F. & Aguiar, Alexandre P., 2022, Revision of Fortipalpa Kasparyan & Ruíz-Cancino, (Ichneumonidae, Cryptinae), Zootaxa 5219 (6), pp. 501-533 : 526-529

Fortipalpa yucatanica Kasparyan & Ruíz-Cancino, 2007

( Figs 16–21 )

Fortipalpa yucatanica Kasparyan & Ruíz-Cancino, 2007 . Original description, figures, ♀. Holotype UATM, paratype ZIN. Fortipalpa yucatanica: Yu et al., 2016 . Catalog.

Redescription. ♀. Fore wing 7.10 mm. Head ( Figs 17A–C , 18 ). MLW 1.44. MWW 0.50. MSM 0.28. CHW 0.47. Clypeus anterior margin centrally with two small tubercles. Antenna with 21–23 flagellomeres. Supra-antennal area polished smooth; with slight median line.

Thorax ( Figs 16 , 17C , 18A ). Epomia approximately straight. Mesoscutum length to width 0.95 × as long as wide. Notaulus reaching about 0.32 of mesoscutum length. Scuto-scutellar groove with vestigial longitudinal wrinkles. Epicnemial carina reaching about 0.7 of distance to subalar ridge; ending above pronotoal spiracle; slightly curved. Sternaulus moderately strong, wide and shallow at posterior 0.25; strongly sinuous. Mesopleuron polished smooth, centro-anteriorly with transversal striation, ventral half punctate. Mesosternum medio-longitudinal suture with short transverse wrinkles. Transverse furrow anterior to propodeum entirely polished smooth. Metapleuron densely punctate-rugulose; moderately pilose.

Propodeum ( Figs 16 , 17C , 18A&C ). Area behind anterior transverse carina transversally strigate, arched posteriorwards. Spiracle elliptic, SLW 1.88. Anterior transverse carina medially slightly arched forwards.

Wings ( Fig. 17D ). Fore wing crossvein 1m-cu more or less uniformly curved; limit between 1m-cu and 1-Rs+M more or less traceable. Vein 2Cua 0.74 × as long as crossvein 2cu-a. APH 0.54. Areolet subpentagonal; distinctly wider than high, AWH 0.84. Crossvein 3r-m spectral. Crossvein 2r-m and 3r-m about same length. Vein 3-M distinctly shorter than 2-M. HW1C 1.76. Hind wing vein Cub almost straight, very slightly convex. Vein 2-1A ending at 0.23 of distance to wing margin.

Metasoma ( Figs 16 , 17C&E , 18A&D ). T1LW 2.54. T1WW 1.92. T1 spiracle at 0.55 of its length. Median depression of T1 elongate, very shallow. T2LW 1.02. T2WW 1.75. T2–8 from distinctly coriarious T2 to smooth T8. Ovipositor straight, OST 1.37; ventral valve apex with 15–16 teeth ( Fig. 18D ).

Color. Head whitish (243,239,227) with supra-antennal area and vertex black (37,32,24). Orbital band complete. Mandible whitish (243,239,227), apical 0.25 dark brown (48,31,18).Antenna black(39,24,12); scape ventral 0.80 whitish; white band composed by fl5 apical 0.40, fl6–13, and fl14 dorsally. Pronotum black, dorsally and collar whitish.Propleuron whitish. Mesoscutum black, median lobe centrally with whitish spot. Tegula, subalar ridge and epicnemium whitish. Mesopleuron light orange, anteriorly black; hypoepimeron, mesepimeron, dorsal division of metapleuron, scutellum and postscutellum whitish. Propodeum orange, apically whitish (244,232,205). Fore leg: coxa whitish (243,239,227), trochanter whitish (241,230,204), trochantellus, femur, tibia and t1–3 light orange; t4–5 dark brown (48,31,19). Mid leg light orange, coxa anteriorly with whitish area; t1 pale yellow, apical 0.45 dark brown (87,68,55); t2 dark brown; t3–5 dark brown (48,31,19). Hind leg light orange, coxa dorsally with whitish spot; t1 whitish (244,243,240), basal 0.25 pale yellow (237,219,153); t2–4 whitish; t5 whitish, apically dark brown. Metasoma striped, dark brown and whitish (243,239,227). T1 tricolored, basal 0.35 pale orange, centrally dark brown, posterior 0.20 whitish. T2–8 bicolored, dark brown with posterior 0.10–0.45 whitish. S2–4 anteriorly dark brown, posteriorly whitish.

Morphometry. See Table 1, Figs 20–21 , and Appendix.

Variation. Fore wing vein 3-M distinctly shorter than 2-M, almost indistinct. Specimen from Trinidad with epicnemium almost entirely blackish (vs. yellowish in holotype). Overall , the Trinidad specimen is more extensively orange, while the specimens from Mexico are yellowish.

Male. Unknown.

Comments. Structure. Most similar to F. panamensis , but differs by having the following features: Sternaulus strongly sinuous (vs. weakly sinuous); epicnemial carina reaching about 0.7 of distance to subalar ridge (vs. reaching about 0.8 of distance to subalar ridge); T1LW 2.54 (vs. 2.67–2.85); HW1C 1.76 (vs. 1.30–1.49); MSM 0.28 (vs. 0.33–0.39); MLW 1.44 (vs. 1.55–1.67); OST 1.37 (vs. 1.58–1.59).

Also similar to F. frida , from which it differs by having the following features: Clypeus anterior margin centrally with two small tubercles (vs. without teeth); mesopleuron polished smooth, centro-anteriorly with transversal striation, ventral half punctate (vs. polished smooth, ventral half punctate); AWH 0.84 (vs. 0.71–0.75); APH 0.54 (vs. 0.69); MSM 0.28 (vs. 0.34); notaulus reaching about 0.32 of mesoscutum length (vs. 0.45–0.49).

Also similar to F. exelysae , but differs by having the following features: Sternaulus strongly sinuous (vs. weakly sinuous); scuto-scutellar groove with vestigial longitudinal wrinkles (vs. with distinct longitudinal wrinkles); area behind anterior transverse carina transversally strigate, arched posteriorwards (vs. transversally strigate, generally weak, anterolaterally faint); notaulus reaching about 0.32 of mesoscutum length (vs. 0.39–0.54); T2–8 from distinctly coriarious T2 to smooth T8 (vs. faintly coriarious); median depression of T1 elongate, very shallow (vs. drop-shaped, distinct, large); epicnemial carina reaching about 0.7 of distance to subalar ridge (vs. reaching about 0.8 of distance to subalar ridge.

Color pattern. Closest to F. exelysae from which it differs by having scape ventrally whitish (vs. dark brown); mid coxa dorsally with whitish area (vs. entirely light orange); and hind coxa with distinct whitish spot dorsally (vs. without or with a faint whitish spot dorsally).

Also similar to F. panamensis , from which it differs by having propodeum posteriorly with pale yellow mark (vs. without pale yellow mark); hind coxa with distinct whitish spot dorsally (vs. without whitish spot); and scape ventrally whitish (vs. dark brown, or with yellow spot).

Distribution records. Mexico, and Trinidad (New record) ( Fig. 19 ).

Material examined. One specimen, 1♀. TRINIDAD, 1♀, 2?, Simla nr. Arima ,?, 250 m, 10°41′33″N, 61°17′22″W, 1977-12-3 /10, W.R.M.Mason, Malaise, FAS 4634,?, CNCI .

Figures

FIGURE 16. Fortipalpa yucatanica Kasparyan & Ruíz-Cancino. Female FAS4634. Habitus (without the head).

FIGURE 17. Fortipalpa yucatanica Kasparyan & Ruíz-Cancino. Female paratype at ZIN. Pictures by Andrey Khalaim. A, Head, frontal. B, Antennae. C, Head, mesosoma and petiole, dorsal. D, Left fore wing (flipped). E. Hind legs and metasoma, left.

FIGURE 18. Fortipalpa yucatanica Kasparyan & Ruíz-Cancino. Female FAS4634. A, Dorsal view (except head). B, Head, frontal. C, Propodeum, dorsal. D, Ovipositor tip, left side.

FIGURE 19. All known distribution records for Fortipalpa spp. (first map) and individual records for each species. Taxa presented in alphabetical order.

FIGURE 20. All results for morphometric ratios calculated for specimens of Fortipalpa spp., shown comparatively. On each plot, the species are sorted on the Y-axis from highest (top) to lowest (bottom) average. Graphs organized by alphabetical order of ratios.

FIGURE 21. Continuation of Figure 20.