Fidia rileyorum Strother, 2008

Strother, M. S. & Staines, C. L., 2008, A revision of the New World genus Fidia Baly 1863 (Coleoptera: Chrysomelidae: Eumolpinae: Adoxini), Zootaxa 1798 (1), pp. 1-100 : 56-58

publication ID	https://doi.org/ 10.11646/zootaxa.1798.1.1
persistent identifier	https://treatment.plazi.org/id/039887A6-FF9D-745C-A1C3-7BF60BD4DEB6
treatment provided by	Felipe
scientific name	Fidia rileyorum Strother
status	sp. nov.

Treatment

Fidia rileyorum Strother , New Species

( Figs. 19 View FIGURES 17–20 , 25 View FIGURES 25–30 , 33 View FIGURES 31–34 , 47 View FIGURES 42–47 , 61 View FIGURES 61–70 , 72 View FIGURES 71–75 , 94 View FIGURES 93–95 ; Map 3)

Fidia longipes: Blatchley 1910: 1143 (faunal treatment); Clavareau 1914: 76 (catalog), ex parte; Leng 1920: 293 (catalog), ex parte; Schultz 1970: 245 (dissertation), ex parte; Rouse & Medvedev 1972: 79 (checklist); Wilcox 1975: 57 (checklist), ex parte; Riley & Enns 1979: 65 (checklist).

Holotype ♂ ( MCZC): " 1 mi. E Moberly Mo. Randolph Co. 28 ⋅ June '72 [date handwritten] E. G. Riley / HOLOTYPE Fidia rileyorum M.S.Strother 1993 [red]". The specimen is glued on a point and is in excellent condition with all appendages intact. The genitalia were not dissected . Paratypes (14): each bears the label, " PARATYPE Fidia rileyorum M.S.Strother 1993 [yellow]" (see specimens examined for label data and deposition of each paratype).

Description. Males: TL = 3.68–4.84 mm, HW = 1.64–2.40 mm. Females: TL = 4.08–5.35 mm, HW = 2.04–2.76 mm. Description as for F. longipes with the following exceptions: males with protibial apex not noticeably expanded laterally nor dorsoventrally compressed; protibial spurs separated by slightly more than the width of one spur, but less that the length of one spur; apical emargination of penis with width subequal to depth, relatively wider and deeper than in F. longipes , giving apical lobes longer, slightly more tapered appearance.

Etymology. Named in honor of Dr. T. J. Riley and E. G. Riley for their constant support and advice throughout this study.

Diagnosis. Small to medium-sized (3.68–5.35 mm); fulvous to black, with at least one pair of femora distinctly lighter basally (some specimens from southern portion of the distribution have all femora entirely flavous to fulvous); pronotal punctation coarse. Male with protibial apex not distinctly dorsoventrally compressed nor laterally expanded; protibial spurs separated by less than the length of one spur ( Fig. 47 View FIGURES 42–47 ); apical emargination of penis with width subequal to depth ( Fig. 94 View FIGURES 93–95 ), usually with small medial tooth.

Fidia rileyorum is similar to F. convexicollis , F. delilahae , and F. longipes . Characters distinguishing it from these three species are given in the diagnosis for each, respectively.

Distribution (Map 3). Fidia rileyorum occurs west of the Appalachian Mountains, from western Ohio to western Alabama, west to southeastern Minnesota and east Texas.

Specimens examined (473). Paratypes: " 1 mi. E Moberly Mo. Randolph Co. 28 ⋅ June '72 [date handwritten] E. G. Riley ( EGRC:3); " 1 mi. E Moberly Mo. Randolph Co. 13 July 75 E. G. Riley " ( EGRC:1); " Sugar Creek June 17, 72 E. G. Riley " ( EGRC:1); " Sugar Creek L. June 15. 72 E. G. Riley " ( EGRC:1); " Sugar Creek L. June 23, 72 E. G. Riley " ( EGRC:1); " Randolph Co. , Mo. Sugar Creek Lake , Moberly: VI-14-80 Coll. E.G.Riley " ( EGRC:1); " Sugar Creek Lake Mo. Moberly, Randolph Co. 16 - June '74 E. G. Riley " ( AJG:1, JEWC:1); " Sugar Creek Lake Mo. Moberly, Randolph Co. 22' June '74 E. G. Riley " ( AJG:1, EGRC:2, JEWC:1) .

Non-type material: ALABAMA: Choctaw Co., Bladon Spgs. St. Pk., vii:31:1963 (AUEM:1), viii:1:1963 (AUEM:1); Lawrence Co., County Road 41, v:23:1992 (MSS:2). ARKANSAS: Benton Co., Bentonville, vi:5:18 (USNM:2), vi:10:18 (USNM:1); Clay Co., Knobel, 10:6:94 (UMRM:2); Johnson Co., Clarksville, vii:16:1974 (UADE:1); Madison Co., [county only], vi:18:1965 (UADE:1); Polk Co., 7.1 mi. NW of Acorn, vii:24:73 (UADE:1); Washington Co., Fayetteville, vii:6:1893 (CSUC:1), White Rock Mt. Winslow, vi:23:44 (INHS:1); No county: South West Ark., no date (AMNH:1). ILLINOIS: Alexander Co., Horseshoe Lake, vi:18:1947 (INHS:1), vi:22:50 (INHS:1); Calhoun Co., Hamburg, vi:8:1949 (INHS:8), Hardin, vi:5–9:1932 (INHS:1); Champaign Co., 3.5 mi. NE Mohamet, vii:2:1960 (INHS:1), vii:7:1960 (INHS:1), Cham., 7:8:15 (INHS:1, MCZC:1), vi:20:15 (MCZC:1), vii:15:15 (INHS:1), Champaign, vi:11:1911 (INHS:10, MCZC:1), U. Ill. Woods, vi:28:37 (USNM:1), Urbana, vi:10:51 (CMNC:3), vi:12:1905 (INHS:3), vi:13:13 (INHS:1), vi:18:14 (INHS:1), vi:2:05 (INHS:2), vi:24:1942 (INHS:1), vi:24:24 (INHS:1), vi:5:1905 (INHS:6), vi:9:08 (INHS:3, USNM:1), vii:5:1926 (INHS:1), Urbana, University Campus, vii:9:1923 (INHS:1); Clark Co., Rocky Branch Creek, N of Clarksville, viii:8–20:1987 (PES:1), Rocky Branch Dolson, (Clarksville), vi:14:1933 (INHS:3); Coles Co., county only, viii:29:1984 (PES:1); Fayette Co., Mulberry Grove, vii:6:1950 (INHS:1), Vandalia, vii:6:1950 (INHS:1); Gallatin Co., Shawneetown, vi:23:1950 (INHS:1) Hancock Co., Kibbe Life Sci. Sta., vii:9:1965 (NDSU:1), A Kebbe [sic] Fd Sta, vi:26:78 (NDSU:1); Jackson Co., Giant City S.P., vii:5:1944 (INHS:2); La Salle Co., Starved Rock St. Pk., vi:27:1938 (UAIC:1), vi:12:1939 (UAIC:1), Utica, no date (UAIC:1); Lawrence Co., Red Hills St. Pk., vi:13:64 (FSCA:1), county only, v:30:38 (CASC:1), vii:13:1937 (UCDC:2); Marion Co., Iuka, vi:21:1950 (INHS:1); Piatt Co., Allerton Park, nr. Monticello, vii:1:1956 (INHS:2); Pope Co., Golconda, vi:22:1932 (INHS:2); Pulaski Co., county only, v:24:08 (INHS:1), vi:28:09 (INHS:1); Putnam Co., county only, vi:24:1932 (INHS:1), vii:4:1932 (INHS:1); Saline Co., Harrisburg, vi:23:1950 (INHS:3); St. Clair Co., Kahokia, vi:30:04 (UMRM:2), county only, vi;12 (MCZC:1), vi:12:1904 (UMRM:2), vi:15:1908 (UMRM:11), vi:20:1903 (MCZC:2, UMRM:11), vi:21:1904 (UMRM:2), vi:30:1904 (UMRM:3), vii:1 (MCZC:2), vii:1:1903 (UMRM:2); Union Co., Pine Hills, vii:3:1972 (SIUC:1); Vermillion Co., Oakwood, vi:18:1939 (INHS:1), vii:31:1947 (INHS:1), Oakwood, Kickapoo St. Pk., vii:5:1948 (INHS:1); No County, Eichorn, Hick's Branch, vi:13:1934 (INHS:1), N. of E. Peoria, viii:25:1960 (INHS;1), White Heath, vi:25:1939 (INHS:3), vii:20:1939 (INHS:1); State only, no date (INHS:3, NDSU:1). INDIANA: Clark Co., State Forest, vi:14:1933 (PURC:2), vi:12:1934 (USNM:1); Crawford Co., county only, vi:24:03 (PURC:1); Knox Co., county only, vii:2:03 (PURC:1), vi:11:06 (TAMU:1), ix:4:21 (PURC:1), vii:11:1929 (PURC:1), vi:26:1937 (UCDC:1); Lawrence Co., county only, iii:24:07 (PURC:1); Marion Co., Ind'ap'lis, vii:14:10 (MCZC:1), Indianapolis, vii:14:1910 (CUIC:1), vii:23:1951 (FSCA:1), vii:27:1958 (FSCA:1); county only, vi:4:22 (PURC:1), vi:16:23 (PURC:1), no date (BMNH:5); Owen Co., county only, vi:18:1982 (PURC:1); Parke Co., county only, vii:5:1970 (SEMC:9); Putnam Co., county only, no date (BMNH:2); Tippecanoe Co., 7m. NW Lafayette, vii:4:1983 (JEWC:2), Lafayette, vii:23:1961 (AMNH:2), vii:6:1987 (FSCA:1), McCormick Woods, viii:10:1981 (JEWC:1), W, Lafayette McCormick Woods, vii:4:1967 (FSCA:2), county only, vii:11:1932 (USNM:1), vii:3:1953 (AMNH:4), vi:17:1955 (USNM:2), vi:26:1955 (USNM:4), vi:27:1956 (AMNH:2), vi:21:1957 (AMNH:2), vi:22:1977 (AMNH:3), vi:9:1982 (FSCA:1); Vermillion Co., county only, vi:14:04 (PURC:1), viii:11:04 (PURC:1), viii:17:21 (CUIC:2); Vigo Co., county only, iv:2:92 (CUIC:4); Warren Co., Pine Village, 7:10:50 (INHS:1); White Co., Springsboro, vii:2:1967 (FSCA:1); County not determined: Bear Wallow, vi:22:1963 (FSCA:1), vi:20:1964 (FSCA:1), vi:7:1965 (FSCA:2), vii:3:1965 (FSCA:1); State only, vii:13:10 (MCZC:1). IOWA: Boone Co., Ledges State Pk., vi:9:1956 (AMNH:3), county only, vii:4:32 (USNM:1); Polk Co., Brown Frst Prsv, vi:12–13:1982 (JEWC:1), Jester Park, vi:1,4:85 (JEWC:3), vi:22:85 (JEWC:2), W. Saylorville Lk., vi:9:85 (JEWC:1), vi:15:1985 (JEWC:2), vi:22:85 (JEWC:3), Walnut Woods, vii:3:1981 (JEWC:6); State only, no date, (AMNH:1, UAIC:1, USNM:1). KENTUCKY: Christian Co., county only, vi:15:1960 (INHS:1, CNCI:1); Franklin Co., Frankfort, 5:7:93 (USNM:1), viii:18:89 (USNM:1); Henderson Co., Henderson, vii:15:23 (LACM:2); Jefferson Co., Louisville, vii:17:94 (UMRM:2); State only, no date (USNM:1). LOUI- SIANA: East Baton Rouge Par., Baton Rouge, vi:3:1986 (LSUC:1), v:14:1992 (MSS:4), LSU Campus, vii:11:1985 (LSUC:2); Natchitoches Par., Kis. N.F., Cunningham Brake at end of FS 348, vi:18:1988 (LSUC:1); St. Landry Par., Opelousas, v:7:08 (USNM:1), Thistlethwaite W.M.A., vii:16:1988 (LSUC:1); St. Tammany Par., county only, vi:15:1980 (LSUC:2); West Feliciana Par., Hardwood on Hwy 61, v:16:1992 (MSS:22), near Weyanoke, v:16:1981 (EGRC:1); State only, v:21:89 (MCZC:1), v:27:97 (MCZC:1), v:31:97 (MCZC:1). MINNESOTA: Olmsted Co., county only, no date (DEFW:1). MISSISSIPPI: Adams Co., Natchez, vi:6:95 (MUIC:1); Benton Co., 6.1 mi. S. of Ashland on Hwy 4/5, vi:2:1992 (MSS:5); Greene Co., county only, vi:19:1970 (MUIC:1); Itawamba Co., 10.4 mi. N. of Hwy 78 on Hwy 25, vi:2:1992 (MSS:4); Jackson Co., Ocean Springs, v:10:1981 (MUIC:2); Lamar Co., county only, v:28:74 (MUIC:1); Lauderdale Co., Meridian, vi:19 (USNM:1); Lincoln Co., 6 mi. NE Bude, viii:11:1982 (MUIC:1); Lowndes Co., 5.1 mi. W. of AL on Hwy 12, v:23:1992 (MSS:2); Oktibbeha Co., 8 mi. SSW Starkville, vi:20:1975 (MUIC:1), vi:30:1975 (MUIC:2), Ag. Coll., 5:9:1921 (MUIC:1), vii:94 (MUIC:1), Ag. College, v:18:1921 (MUIC:1); Pontotoc Co., 1 mi. SE Ecru, vii:2:1980 (MUIC:1), vii:16:1980 (MUIC:1); Scott Co., Forest, vi:9:1989 (RHT:6); Smith Co., 2.3 mi. E Polkville, vi:10:1989 (RHT:3); Tippah Co., 3.4 mi. E. of Hwy 2 on Hwy 4, vi:2:1992 (MSS:14); Tishomingo Co., 5.6 mi. N. of county line on Hwy 25, vi:2:1992 (MSS:21); Warren Co., 1 mi. S Redwood, vi:10:1984 (RHT:1); Wilkinson Co., Pond; Clark Creek Nat. Area, vi:26:1987 (LSUC:1); Winston Co., 13 mi. S. Starkville, vi:14–25:1982 (MUIC:1), vi:28:1982 (MUIC:1), vii:12–19:1982 (MUIC:1); Nanih Waiya, v:29:1977 (MUIC:1). MISSOURI: Barry Co., vic overlook on Sugar Camp Rd., 3 mi, SE R.R. St., vii:3:1986 (LSUC:1), SW Roar. Riv. St. Pk., Sugar Camp rd. scenic over, vii:2:1984 (EGRC:1), Roaring River St. Park, vi:15:54 (CASC:3), Roaring River State Park, vi:10:1974 (UMRM:1); Boone Co., Ashland, viii:3:1968 (UMRM:1), Ashland Wildlife Area, vii:4:74 (EGRC:3), viii:3:76 (EGRC:1), Columbia, vii:1:1967 (UMRM:1), Hallsville; The Pinnacles, vi:27:1971 (UMRM:1); Camden Co., 1/ 2 mi. N.E. jct J on U.S. 54, vi:25:1975 (UMRM:1); Carter Co., Sky Line Drive nr. Van Buren, vii:22:1978 (EGRC:1); Franklin Co., Fern Glen, vi:14:1901 (USNM:1); Greene Co., Springfield, vii:15:1915 (USNM:1), vii:18:1915 (USNM:4); Jackson Co., Atherton, vi:4:22 (SEMC:1); Montgomery Co., Graham Cave State Park, vi:22:1974 (UMRM:1); Morgan Co., 1 mi. N jct. J on Rt.5, vi:25:1975 (UMRM:1); Polk Co., La Petit Gemme Pr. 3 mi. W. Bolivar, vi:13:1978 (EGRC:1); Pulaski Co., Ft. L. Wood, vii:11:1971 (TAMU:1); Randolph Co., 1 mi. E. Moberly, vi:28:72 (EGRC:4), vii:13:75 (EGRC:1), vii:3–4:1982 (EGRC:1), vii:3–4:1992 (EGRC:1), Sugar Creek, vi:17:72 (EGRC:1), Sugar Creek L., vi:15:72 (EGRC:1), vi:23:72 (EGRC:1), Sugar Creek Lake, Moberly, vi:16:74 (AJG:1, JEWC:1), vi:22:74 (AJG:1, EGRC:2, JEWC:1), vi:14:80 (EGRC:1); St. Charles Co., St. Charles, viii:20:99 (UMRM:1); St. Francois Co., St. Francois St. Pk., vi:21:1978 (EGRC:1); St. Louis Co., 2 m W St. Louis, vi:26:1904 (USNM:1), Creve Coeur Lake, vi:10:1900 (USNM:1), St. Louis, v:23:99 (UMRM:1), county only, v:25:02 (UMRM:1); Saline Co., Van Meter St. Pk., vi:27:1972 (UMRM:1); Stoddard Co., 2.8 mi. NE of Dexter; Holly Preserve, vi:10:1975 (UMRM:1), vi:22:1978 (EGRC:1); Mingo Wildlf. Refuge, vi:12:1975 (UMRM:1); Stone Co., 1 mi. E jct. 160 & 248, vi:7:1975 (UMRM:1); Wayne Co., 3 mi. W Waynesville, vi:23:1978 (EGRC:5); State only, no date (DEFW:1). OHIO: Champaign Co., Westville, vii:19:62 (FSCA:1); Greene Co., county only, v:20:53 (OSUC:1), vii:8 (OSUC:2, CDAE:1); Hamilton Co., Cin., vi:15:01 (ANSP:2), no date (ANSP:6); State only, vi:24 (MCZC:1), no date (MCZC:3, UAIC:1, USNM:1). OKLAHOMA: Adair Co., Watts, vi:16:1939 (OSEC:1); Delaware Co., Flint, vi:6:1934 (OSEC:1). TENNESSEE: Davidson Co., Nashville, vi:23:93 (UMRM:2, USNM:4); Knox Co., Knoxville, vi:1:1957 (CMNC:1); Lake Co., Reelfoot Lk., vi:2:54 (OSUC:1); Shelby Co., Shelby For. St. Pk., vi:28:1954 (CMNC:1); Smith Co., Elmwood, no date (CASC:4). TEXAS: Brazos Co., College Sta., vii:3:1963 (CMNC:1). No locality, no date (INHS:1, TAMU:2).

Temporal Data. Collecting dates ranged from 2 April to 4 September.

Natural History. Plant associations included various members of the Vitaceae , including Ampelopsis arborea (L.), Parthenocissus quinquefolia (L.), and Vitis spp. as well as clover, cotton, locust, and nettle. Specimens were also taken at lights and in Leggett, Malaise, pitfall, and yellow sticky traps.

Taxonomic History. This species was not previously recognized as a distinct taxon, and the name F. longipes has been invariably applied to it. Schultz (1970) mentioned examining two males, which lacked the typical spatulate protibiae of F. longipes , and suggested that they might represent a new species. However, he did not note the differences between the apex of the penis of the two species and applied the name F. longipes to both taxa.