Falcidius scipionis, Gnezdilov & Wilson, 2008
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publication ID |
https://doi.org/ 10.1080/00222930802106229 |
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persistent identifier |
https://treatment.plazi.org/id/03FD87FF-FFF1-FFA2-FE3B-FA93FCAEFD44 |
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treatment provided by |
Carolina |
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scientific name |
Falcidius scipionis |
| status |
sp. nov. |
Falcidius scipionis View in CoL sp. nov.
( Figures 15, 16 View Figures 9–16 , and 33)
Falcidius doriae Dlabola 1984: 66 View in CoL , figs 236–244.
Type material
Algeria: „, holotype, Souq Ahras, Coll. E. de Bergevin ( MNHN) .
Paratypes. 2 „, as holotype ( MNHN, ZIN); 1 „, 1♀, Oued Taga, 6–11 km SW of Timgad, ‘stat. 45, wet cultivated area’, 1300–1400 m, 3–4.V.1980, E v. Nieukerken, G. Bryan & P. Oosterbroek leg. ( ZMAN); 1 „, near Aïn Amara, 18 km SW of Guelma, 650 m, 5.V.1980, olive yard, E v. Nieukerken, G. Bryan & P. Oosterbroek leg. ( MNHN); 3 „, 3♀, Constantine, Coll. E. de Bergevin ( MNHN, ZIN); 1 „, 2♀, Djebel Ouach, Coll. E. de Bergevin ( MNHN); 1 „, 1♀, St. Charles , A. The «ry ( MNHN, ZIN) .
Description
Metope with weak median and sublateral keels jointed at its upper margin. Coryphe three times as wide as long, with widely convex anterior margin. Pronotum without keels. Scutellum without keels or sometimes with weak median keel. Forewings nearly oval, with narrow hypocostal plate. Radius bifurcate, median trifurcate, cubitus anterior simple ( R 2 M 3 CuA 1). Metatarsomere I with single intermediate spine.
Coloration ( Figures 15 and 16 View Figures 9–16 ). General coloration light brown yellowish. Metope with dark brown dots and traces of sensory pits. Femora and tibiae with longitudinal dark brown stripes. Apices of spines, metatarsomeres III, and claws dark brown. Abdominal tergites black.
Male genitalia ( Figure 33 View Figure 33 ). Pygofer with weakly convex hind margin. Anal tube elongate, narrowing apically, lateral margins turned down. Anal column about onethird as long as anal tube. Each phallobase lobe with narrow supporting process and wide covering process. Ventral phallobase lobe long and wide. Aedeagal hooks turned to dorsal surface of the phallobase. Style with weakly concave hind margin. Capitulum weakly narrowing to rounded apex (in dorsal view); lateral tooth with straight margin.
Female genitalia. Hind margin of sternum VII with short semicircular median process. Anal tube widely rounded apically.
Body length. Males 3.7–3.8 mm; females 4.1–4.2 mm.
Distribution
Algeria.
Etymology
The species name is derived from the name of Roman general Scipio in the third Punic war.
Falcidius duffelsicus Dlabola, 1982 View in CoL
( Figures 17–19 View Figures 17–22 , and 34)
Falcidius duffelsicus Dlabola 1982: 135 View in CoL .
Material examined
Tunisia: „, holotype, 1 „, 1♀, paratypes, 5 km N of Hammamet , 0–150 m, 9– 17.VI.1980, J.P. Duffels leg. ( ZMAN) ; 1♀, about 100 km N of Hammamet , sea cliff ,
31.III.1980, J.M. Cox leg. ( BMNH); 1 „, 1♀, 3–7 km S of Korbous , 0–100 m , 26.V.1984, J.P. Duffels leg. ( ZIN) .
Supplementary description
Metope convex (visible from above), with sublateral keels jointed at its upper margin. Coryphe 2.5 times as wide as long, sometimes with weak median keel, its anterior margin strongly convex. Pronotum sometimes with weak median keel. Scutellum with median groove. Forewings with narrow hypocostal plate. Radius and median bifurcate, cubitus anterior simple ( R 2 M 2 CuA 1). Metatarsomere I with single intermediate spine.
Coloration ( Figures 17–19 View Figures 17–22 ). General coloration light brown yellowish with dark brown or black spots and stripes. Metope with large approximately rectangular black spot above clypeus and a pair of dark brown or black large rounded spots between sublateral keels apically. Upper angles of metope and traces of sensory pits dark brown. Clypeus with longitudinal brown stripes, it is edged by black stripes on each side. Rostrum with terminal segment dark brown. Coryphe with two pairs of dark brown elongate spots on each side. Forewings with dark brown longitudinal veins and light costal margin. Femora and tibiae with longitudinal dark brown strips. Spines of legs with dark brown apices. Abdominal sternites including pygofer (in males with dark brown apical angles) light yellow with brown dots at seta bases. Abdominal tergites dark brown or black excluding yellow brownish median line. Anal tube light yellow. Style light yellow excluding dark brown capitulum.
Male genitalia ( Figure 34 View Figure 34 ). Pygofer with weakly convex hind margin. Anal tune elongate, narrowing apically (in dorsal view), apical angles turned down (in lateral view). Anal column one-third as long as anal tube. Each dorsolateral phallobase lobe with narrow supporting process and wide covering process. Ventral phallobase lobe long and wide, not narrowing apically. Ventral aedeagal hooks strongly curved, directed to dorsal suface of the phallobase. Style with weakly concave hind margin, caudodorsal angle narrowly rounded. Capitulum on long neck, narrow, narrowing apically; lateral tooth with concave margin.
Female. Hind margin of sternum VII with semicircular median process.
Body length. Males 3.4 mm (after Dlabola 3.4–3.6 mm); females 3.8 mm (after Dlabola 4.1–4.2 mm).
Distribution
Tunisia ( Dlabola 1982; present data).
Falcidius ebejeri sp. nov.
( Figures 20–22 View Figures 17–22 , and 35)
Type material
Malta: „, holotype, Mtahleb , 4.III.2001, M.J. Ebejer leg. ( NMWC) . Paratypes: 6 ♀, as holotype ( NMCW and ZIN) ; 1♀, Isole Maltesi , Ghar Lapsi (Siggizwi), 19.IV.1965, C. Baroni leg. ( MNHN) .
Description
Metope convex (visible from above), with very weak sublateral keels. Coryphe transverse (three times as wide as long), anterior margin weakly convex. Pronotum and scutellum without keels. Forewings with narrow hypocostal plate. Radius bifurcate, median trifurcate, cubitus anterior simple ( R 2 M 3 CuA 1); small number of transverse veins. Metatarsomere I with single intermediate spine.
Coloration ( Figures 20–22 View Figures 17–22 ). General coloration light brown. Metope with large rectangular black spot above clypeus. Apical angles of metope terminated by narrow dark brown stripes. Traces of sensory pits dark brown. Anteclypeus with four longitudinal black stripes. Terminal segment of rostrum black. Forewings with dark brown longitudinal veins and yellowish transversal veins. Abdominal tergites excluding median line black. Abdominal sternites and genital segments light yellow brownish. Femora and tibiae with longitudinal dark brown stripes. Spines and metatarsomeres III with dark brown apices. Gonoplacs with dark brown margins. Female anal tube with dark apex.
Male genitalia ( Figure 35 View Figure 35 ). Pygofer with convex hind margin. Anal tube not narrowing apically, its apical angles turned down. Anal column one-quarter to onethird as long as anal tube. Each dorsolateral phallobase lobe with rudimentary supporting process and wide covering process. Ventral phallobase lobe long and wide. Aedeagal hooks directed to dorsal surface of the phallobase. Style with weakly concave hind margin; caudodorsal angle widely rounded. Capitulum of style narrowing to apex (in dorsal view); lateral tooth with concave margin.
Female genitalia. Hind margin of sternum VII with semicircular median process. Gonoplacs with a pair of weak transverse keels. Anal tube weakly narrowing apically.
Body length. Male 3.6 mm; females 4.0– 4.5 mm.
Distribution
Endemic species in Malta.
Etymology
The species is named for the collector and Maltese dipterist Dr Martin Ebejer.
Falcidius hannibal View in CoL sp. nov. ( Figures 23–26 View Figures 23–26 , 36 View Figure 36 , and 37) Falcidius chlorizans Dlabola 1980: 215 View in CoL
Type material
Tunisia: „, holotype, 5 km W of Sbeitla , 300 m, 17.V.1984, J.P. Duffels leg. ( ZMAN) . Paratypes: 4 „, 8♀, as holotype ( ZMAN, ZIN); Algeria: 5 „ , 2♀, massif des Aurès, Bouhamana, 1300 m, 11.V.1984, J.P. Duffels leg. ( ZMAN, ZIN); 4 „, 2♀, Batna, ‘Exp. Obenb.’ ( MNHN; NMPC); 9 „ , 13♀, Djurdjura , 15–19.VI.1971, 1000– 2000 m, A. Hoffer et J. Hora « k leg. ( NMPC, ZIN, NMWC); 1♀, Lambe«ze, VI.1885, L. Bleuse leg. ( MNHN) .
Description
Metope convex, with distinct or very weak (traces) sublateral keels jointed close to its upper margin, sometimes with weak median keel. Coryphe 2.5 times as wide as long, sometimes with median keel, anterior margin convex. Pronotum and scutellum without keels. Forewings with narrow hypocostal plate. Radius bifurcate, median tri- or tetrafurcate, cubitus anterior simple ( R 2 M 3–4 CuA 1). Metatarsomere I with one or two intermediate spines.
Coloration ( Figures 23–26 View Figures 23–26 ). General coloration light yellow brownish. Metope with dark brown or black wide transverse band above clypeus. Metope and pronotum with dark brown traces of sensory pits. Postclypeus with transverse lateral dark brown stripes. Anteclypeus with dark brown spots. Rostrum with dark brown terminal segment. Forewings with light costal margin. Longitudinal veins light brown. Femora and tibiae with dark brown longitudinal stripes and spots. Abdominal tergites excluding median line black. Abdominal sternites including genital segments light yellow. Apices of spines and metatarsomeres III dark brown.
Male genitalia ( Figures 36 View Figure 36 and 37 View Figure 37 ). Pygofer with nearly straight or convex hind margin. Anal tube narrowing apically, its apical angles in shape of semicircular processes turned down. Anal column one-quarter as long as anal tube. Each phallobase lobe with narrow supporting process and wide covering process. Aedeagal hooks directed to dorsal surface of the phallobase. Style with weakly concave hind margin. Capitulum narrowing to truncated or rounded apex; lateral tooth with concave margin.
Female genitalia. Hind margin of sternum VII with semicircular median process. Gonoplacs with a pair of weak transverse keels. Anal tube weakly narrowing apically.
Body length. Males 2.7–3.2 mm; females 3.0– 3.5 mm.
Distribution
Tunisia, Algeria.
Etymology
The species name is derived from the name of Carthaginian general Hannibăl.
Note
The material cited by Dlabola ( Dlabola 1980: 215) under the name F. chrorizans belongs to F. hannibal sp. nov. This material is included in the type series.
Key to species
(1) Cells of forewings with dark brown edging inside ( Figure 2 View Figures 1–8 )......... 2
Cells of forewings without dark brown edging inside ( Figure 13 View Figures 9–16 ).... 4
(2) Phallobase strongly enlarged proximally, with very wide covering processes ( Figure 29A,B View Figure 29 )......................... F. marocanus Bergevin Phallobase View in CoL not enlarged proximally, with moderately wide covering processes ( Figures 28A,B View Figure 28 )................................ 3
(3) Each dorsolateral phallobase lobe with vertical supporting process situated
near to its ventral margin ( Figure 27A View Figure 27 ). Aedeagal hooks directed to the
ventral margin of the phallobase. Style with strongly concave hind margin
( Figure 27D View Figure 27 )........................... F. apterus (Fabricius) View in CoL
Each dorsolateral phallobase lobe with horizontal supporting process
situated near to its dorsal margin ( Figure 28A View Figure 28 ). Aedeagal hooks directed
to the dorsal margin of the phallobase. Style with weakly concave hind
margin ( Figure 28E View Figure 28 )................... F. limbatus (A. Costa) View in CoL
(4) Metope with large black spot or band above clypeus ( Figures 19, 22 View Figures 17–22 , 25 View Figures 23–26 , and 26).............................................. 5 Metope without large black spot or band above clypeus ( Figure 9 View Figures 9–16 ).. 7
(5) Metope with black rectangular spot above clypeus ( Figures 19 and 22 View Figures 17–22 ). Forewings with dark brown longitudinal veins ( Figures 18 and 21 View Figures 17–22 ).... 6 Metope with dark brown or black band above clypeus ( Figures 25 and 26 View Figures 23–26 ). Forewings with light brown longitudinal veins ( Figure 24 View Figures 23–26 ).......................................... F. hannibal View in CoL sp. nov.
(6) Metope with a pair of distinct large dark brown or black spots apically ( Figure 19 View Figures 17–22 ). Coryphe with anterior margin strongly convex ( Figure 17 View Figures 17–22 ). Supporting processes of the phallobase narrow ( Figure 34A View Figure 34 )............................................. F. duffelsicus Dlabola Metope View in CoL without distinct large dark brown or black spots apically ( Figure 22 View Figures 17–22 ). Coryphe with anterior margin weakly convex ( Figure 20 View Figures 17–22 ). Supporting processes of the phallobase rudimentary ( Figure 35A View Figure 35 )......................................... F. ebejeri View in CoL sp. nov.
(7) Male anal tube strongly enlarged medially in dorsal view ( Figure 32D View Figure 32 )......................................... F. diphtheriopsis Bergevin View in CoL Male anal tube weakly enlarges medially in dorsal view ( Figure 30C View Figure 30 ). 8
(8) Phallobase with wide supporting processes ( Figure 30A View Figure 30 ). Lateral margins of male anal tube straight in lateral view ( Figure 30B View Figure 30 )..... F. doriae (Ferrari) Phallobase View in CoL with narrow supporting processes ( Figure 31A View Figure 31 ). Lateral margins of male anal tube convex in lateral view ( Figure 33F View Figure 33 )...... 9
(9) Phallobase with waist proximally ( Figures 31A,B View Figure 31 ). Ventral phallobase lobe with a pair of longitudinal bulge-like projections basally which cover supporting processes of the phallobase (in ventral view) ( Figure 31B View Figure 31 )..
........................................ F. chlorizans (Rey) View in CoL
Phallobase without waist proximally ( Figures 33A,B View Figure 33 ). Ventral phallobase lobe glib, not covering supporting processes of the phallobase (in ventral view) ( Figure 33B View Figure 33 )................. F. scipionis View in CoL sp. nov.
Acknowlegements
We are sincerely grateful to Prof. Thierry Bourgoin (Paris, France), Prof. Vera D’Urso (Catania, Italy), Prof. Adalgisa Guglielmino ( Viterbo , Italy), Dr Roberto Poggi ( Genova , Italy), Dr Hans Duffels ( Amsterdam , the Netherlands), Dr Petr Kment (Prague, Czech Republic), Dr Pavel Lauterer and Dr Igor Malenovsky ( Brno , Czech Republic), Dr Anna Maryan «ska-Nadachowska (Krako« w, Poland), Dr Stuart McKamey (Washington, USA), Mr Mick Webb ( London , UK), Mr Jerome Constant (Bruxelles, Belgium), Dr Larry Hulde «n ( Helsinki , Finland) for an opportunity to examine the material, Prof. Niels P. Kristensen ( Copenhagen , Denmark) for the photos of the type specimen of Cercopis aptera and Mme Claude Pierre (Paris, France) for her help with preparation of the pictures. The study was financially supported by grants from the Royal Society (London, UK), the Russian Fund for Basic Researches (06–04–48427), and a Museum National d`Histoire Naturelle (Paris, France) grant 2007 for invited scientists. The collection of the Zoological Institute of the Russian Academy of Sciences is supported by Rosnauka for UFC no. 2–2.20 .
No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.
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Genus |
Falcidius scipionis
| Gnezdilov, Vladimir M. & Wilson, Michael R. 2008 |
Falcidius doriae
| Dlabola J 1984: 66 |
Falcidius duffelsicus Dlabola 1982: 135
| Dlabola J 1982: 135 |
Falcidius hannibal
| Dlabola J 1980: 215 |