Exocelina gorokaensis Shaverdo & Balke

Shaverdo, Helena, Sagata, Katayo, Panjaitan, Rawati, Menufandu, Herlina & Balke, Michael, 2014, Description of 23 new species of the Exocelinaekari-group from New Guinea, with a key to all representatives of the species group (Coleoptera, Dytiscidae, Copelatinae), ZooKeys 468, pp. 1-83 : 8

Taxon classification Animalia Coleoptera Dytiscidae

4. Exocelina gorokaensis Shaverdo & Balke sp. n. Figs 14, 42

Exocelina undescribed sp. MB1307: Toussaint et al. 2014: Supplementary figs 1-4, Tab. 2.

Type locality.

Papua New Guinea: Eastern Highlands Province, 37 km S Goroka, Hogave village, Mt. Michael, 06°22.48'S; 145°15.26'E.

Type material.

Holotype: male "Papua New Guinea: Eastern Highlands, 37 km S Goroka, Hogave vill., Mt. Michael, 2179-2800m, 9.-15vii.2009, 06.22.479S 145.15.256E, Sagata (PNG 230)" (ZSM). Paratypes: Eastern Highlands: 23 males, 39 females with the same label as the holotype, 1 male additionally with a green label "DNA M.Balke 4038" (NHMW, ZSM). 5 males, 4 females "Papua New Guinea: Eastern Highlands, Goroka, Mt. Gahavisuka, 2200m, 8.iv.2006, 06.00.896S 145.24.753E, Balke & Sagata (PNG 35)" (NHMW, ZSM). 1 male, 1 female "Papua New Guinea: Eastern Highlands, Hogu, Mt. Barola, 1900m, 9.v.2006, 06.17.556S 145.45.036E, Balke & Sagata (PNG 56)" (ZSM). 10 males, 5 females "Papua New Guinea: Eastern Highlands, Goroka, below Mt. Otto, 2000m, 11.v.2006, 06.01.687S 145.26.493E, Balke (PNG 57)", one male additionally with a green label "DNA M.Balke 1305" (NHMW, ZSM). 21 males, 14 females "Papua New Guinea: Eastern Highlands, Goroka, Daulo Pass, 2500m, 19.v.2006, 06.02.432S 145.13.333E, John & Balke (PNG 67)", one male additionally with a green label "DNA M.Balke 1307" (NHMW, ZSM). 2 males, 1 female "Papua New Guinea: Eastern Highlands, Akameku - Brahmin, Bismarck Range, 2200m, 23.xi.2006, 05.56.801S 145.22.238E, Balke & Kinibel (PNG 106)" (ZSM). 1 female "Papua New Guinea: Eastern Highlands, Akameku - Brahmin, Bismarck Range, 2400m, 23.xi.2006, 05.55.615S 145.22.699E, Balke & Kinibel (PNG 107)", "DNA M.Balke 1518" [green] (ZSM). Simbu: 8 males, 5 females "Papua New Guinea Simbu prov L. Cizek lgt.", "Kundiawa, Mu vill. 145°02'E 4°42'S [6°05'S; 145°02'E] III.2001, 1900m" (ZSM). Western Highlands: 9 males, 4 females "Papua New Guinea: Western Highlands, Mondmill,5 Km SE Minj, small pools near creek, 1741m, 12.vi.2006, 05.56.801S 144.39.898E, John (PNG 77)", one male additionally with a green label "DNA M.Balke 1343" (NHMW, ZSM).

Diagnosis.

Beetle medium-sized, dark brown to piceous, with paler clypeus, vertex, and pronotal sides, submatt; pronotum with distinct lateral bead; male antennomeres 3-5 evidently enlarged, slightly rounded, almost equal in size, antennomeres 6 and 7 somewhat enlarged; male protarsomere 4 with medium-sized, slender, evidently curved anterolateral hook-like seta; median lobe with very weak submedian constriction and apex very slightly concave in ventral view and with apex slightly pointed and broadened in lateral view; paramere with distinct notch on dorsal side and subdistal part elongate, with numerous, dense, more or less long, thin setae. The species is similar to Exocelina jimiensis sp. n., from which differs with evident punctation of the dorsal surface and shape of the median lobe. Also it is similar to Exocelina vovai sp. n. and Exocelina bismarckensis sp. n., from which differs with weaker punctation of the dorsal surface, paramere with distinct notch on the dorsal side, and shape of the median lobe.

Description.

Size and shape: Beetle medium-sized (TL-H 3.7-4.3 mm, TL 4.1-4.8 mm, MW 1.95-2.4 mm), with oblong-oval habitus, broadest at elytral middle. Coloration: Dorsal surface more or less uniform dark brown to piceous, paler on clypeus, vertex, pronotal sides, and along elytral suture; head appendages and legs yellowish red to dark reddish, legs darker distally (Fig. 42). Teneral specimens paler.

Surface sculpture: Head with very dense, coarse punctation (spaces between punctures 1-2 times size of punctures). Pronotum with punctation finer than on head. Elytra with punctation sparser than on pronotum. Pronotum and elytra with relatively strongly impressed microreticulation, dorsal surface submatt. Head with microreticulation stronger. Metaventrite and metacoxa distinctly microreticulate, metacoxal figs with longitudinal strioles and transverse wrinkles. Abdominal ventrites with distinct microreticulation, strioles, and fine sparse punctation, coarser and denser on two last abdominal ventrites.

Structures: Pronotum with distinct lateral bead. Base of prosternum and neck of prosternal process and neck of prosternal process with distinct ridge, smooth and not rounded anteriorly, with small anterolateral extensions. Blade of prosternal process lanceolate, relatively narrow, convex, with distinct lateral bead and few setae; neck and blade of prosternal process evenly jointed. Abdominal ventrite 6 broadly rounded or slightly truncate apically.

Male: Antennomeres 3-5 evidently enlarged, slightly rounded, almost equal in size, antennomeres 6 and 7 somewhat enlarged (Fig. 14A), antennomeres 3-7 rugose ventrally. Protarsomere 4 with medium-sized, slender, evidently curved anterolateral hook. Protarsomere 5 ventrally with anterior row of 17 and posterior row of 6 short setae (Fig. 14B). Abdominal ventrite 6 with 11-14 lateral striae on each side, broadly rounded or slightly truncate apically. Median lobe with very weak submedian constriction and apex very slightly concave in ventral view and with apex slightly pointed and broadened in lateral view (Fig. 14C, D). Paramere with distinct notch on dorsal side and subdistal part elongate, with numerous, dense, more or less long, thin setae (Fig. 14E).

Holotype: TL-H 4.0 mm, TL 4.5 mm, MW 2.15 mm.

Female: Antennae simple, abdominal ventrite 6 broadly rounded apically, without striae.

Distribution.

Papua New Guinea: Eastern Highlands, Simbu, and Western Highlands Provinces. The species is known mainly from the area around Goroka and Kundiawa (Fig. 53).

Etymology.

The species is named after Goroka, where it was mostly collected. The name is an adjective in the nominative singular.