Exocelina bagus (Balke & Hendrich, 2001)

Shaverdo, Helena, Wild, Michael, Sumoked, Bob & Balke, Michael, 2017, Six new species of the genus Exocelina Broun, 1886 from Wano Land, New Guinea (Coleoptera, Dytiscidae, Copelatinae), ZooKeys 665, pp. 93-120 : 95

3. Exocelina bagus (Balke & Hendrich, 2001) Figs 6, 14

Copelatus (Papuadytes) speciosus Balke & Hendrich, 1998: 336; Nilsson 2001: 77 (catalogue).

Copelatus (Papuadytes) bagus Balke & Hendrich, in Balke 2001: 361.

Exocelina bagus (Balke & Hendrich, 2001): Nilsson 2007: 33 (as E. baga , comb.n.).

Exocelina bagus MB4915: Toussaint et al. 2014: Supplementary figs 1-4, Tab. 2.

Type locality.

Papua: Nabire Regency, 54-55 km of road Nabire to Enarotali, ca. 03°29.80'S; 135°43.89'E. Note: the road only goes up to Enarotali, Ilaga is much further in the mountains, therefore, people now refer to the road as Nabire-Enarotali.

Type material studied.

Holotype: male "IR90-11: W. New Guinea, Trek Nabire-Ilaga, km55, 19.-25.ix.1990, Balke", “HOLOTYPUS” [red], " Copelatus speciosus sp.n. Balke des. 1997" [red] (NHMW). Paratypes: 9 males, 12 females with the same label as the holotype (NHMW). 6 females "W.-Neuguinea/Paniai Prov. Strasse Nabire-Ilaga km 54 700m, 22.-25.9.1990/IR 11 leg: Balke & Hendrich" (NHMW). 7 males, 8 females "IR 20-W. New Guinea, track Nabire-Ilaga KM 59, ca.750m, 18.vii.1991, Balke & Hendrich leg." (NHMW). 1 female "IR 21-W. New Guinea track Nabire-Ilaga KM 65, Kali Utowa, 250 M, 18-19.vii.1991 Balke & Hendrich leg." (NHMW), note: the original description says "one male", it is probably a mistake in the sex identification or type error. 6 females "IR 23-W. New Guinea, track Nabire-Ilaga, KM 62, 250m, 24.vii.1991 Balke & Hendrich leg." (NHMW). 1 female "IR 24-W. New Guinea, track Nabire-Ilaga Km 54, basecamp, 750m, 25.vii.1991 Balke & Hendrich leg." (NHMW). All paratypes are additionally with red labels "Paratypus Copelatus speciosus Balke des. 1997".

Additional material.

10 males, 15 females "IR #91-7 (IR 24). West New Guinea, Nabire-Ilaga km 54, 750m, 25.&27.1991 Balke" (NHMW). 8 males, 8 females "IRIAN JAYA: Paniai Prov. road Nabire-Ilaga, km 54 26./27.8.1996, 750-800m leg. M. Balke (96 # 2)" (NHMW). 6 males, 5 females "IRIAN JAYA: Paniai Prov. road Nabire-Ilaga, km 54 30.8.1996, 750m leg. M. Balke (96 # 9)" (NHMW). 2 males "IRIAN JAYA: Paniai Prov. road Nabire-Ilaga, km 54 10.9.1996, 900m leg. M. Balke (96 # 19)" (NHMW). 7 males, 3 females "IRIAN JAYA: Paniai Prov. road Nabire-Ilaga, km 54 10.9.1996, 800m leg. M. Balke (96 # 20)" (NHMW). Note: although most of these mentioned above specimens are with the paratype labels, they are not included in the type material of the original description in Balke (1998, p. 336). 1 female "Indonesia: Papua, Road Nabire-Enarotali KM 55, 774m, 22.x.2011, 03 29.796S 135 43.885E, Uncen (PAP09)" (NHMW). 2 males, 1 female "Indonesia: Papua, Road Nabire-Enarotali KM 60, 640m, 22.x.2011, 03 30.474S 135 42.611E, Uncen (PAP10)" (NHMW).

Diagnosis.

Beetle medium-sized (TL-H 3.8-4.8 mm), elongate; dark brown, with reddish brown pronotal sites and head anteriorly; submatt, with fine but evident punctation and rather strongly impressed microreticulation; pronotum without lateral bead (Fig. 6; fig. 3 in Balke (1998)); male antennae strongly and modified: antennomeres 2-3 very strongly reduced, 4-6 excessively enlarged and 3 and 7 strongly enlarged (Fig. 6; fig. 16 in Balke (1998)); male protarsomere 4 with large, strongly curved anterolateral hook-like setae; male protarsomere 5 slightly concave ventrally, with anterior band of more than 30 and posterior row of five relatively long, not pointed setae (Fig. 14D); median lobe strongly curved, evenly tapering to apex, apex straight, pointed, with short, thick lateral setae in lateral view, subdistal part of median lobe strongly broadened in ventral view; paramere with distinct dorsal notch and large, elongate subdistal part; subdistal setae very dense, strong, long, proximal setae very sparse, thin, small, weakly visible (Fig. 14 A–C; figs 39, 86, 92 in Balke (1998)). For complete description, see Balke (1998).

Distribution.

Papua: Nabire Regency. The species is known only from the Mount Gamey area (Fig. 19).