Eustrongylosoma tumbuna, Golovatch, Sergei I. & Stoev, Pavel, 2011

Golovatch, Sergei I. & Stoev, Pavel, 2011, Review of the millipede genus Eustrongylosoma Silvestri, 1896 in the island of New Guinea, with descriptions of nine new species (Diplopoda, Polydesmida, Paradoxosomatidae), International Journal of Myriapodology 5, pp. 1-26 : 5-6

Eustrongylosoma tumbuna sp. n. Figs 7-14

Type material.

Holotype♂(NMNHS),Papua New Guinea, Western Province, Mt Fugilil, near camp, 2980 m, 9 September 1975, B.S.E. (leg. Ph. Chapman & P. Beron). - Paratype: 1 ♂(NMNHS), West Sepik Province, Telefomin area, 1700 m, September 1975, B.S.E. (leg. P. Beron).

Name.

“tumbuna” means “ancestor” or “ancestors” in Pidgin English. A noun in apposition.

Diagnosis.

Differs from congeners by the rather wide paratergal calluses strongly delimited not only dorsally, but also ventrally, by the distal part of the solenophore being dentate/lobulate, not split (see also Key below).

Description.

Length of both holo- and paratype ca 20 mm, width of pro- and metazona 1.4 and 2.1 (holotype) or 1.2 and 1.8 mm (paratype), respectively. Coloration rather uniformly light brown to brown, only front 1/3 of body slightly more intensely brown; dorsal sides of prozona, calluses on paraterga, epiproct and venter light yellow-brown to nearly pallid. Sometimes (paratype) region near stricture between pro- and metazona slightly infuscate, brown, thus creating a faint cingulate pattern.

In width, head >> collum = segment 3 = 4 <2 <5-16; thereafter body gently tapering towards telson. Entire head densely setose. Antennae rather long, slightly clavate, reaching beyond midlength of segment 3 or 4 dorsally; antennomere 6 longer than 2nd. Collum transversely oval, lateral flaps small, regularly and broadly rounded. Tegument generally smooth and shining, only below paraterga faintly microgranulate; rear halves of metaterga sometimes faintly rugulose. Paraterga of postcollum segments moderately well developed (Figs 7, 8), set low (at about 1/3 midbody height), subhorizontal, evidently thicker in pore-bearing segments than in poreless ones; paratergum 2 with a small anterolateral denticle, its front margin straight; following paraterga without denticulations at lateral margin, sometimes with only a very faint undulation near midway. Caudal corner of all postcollum paraterga pointed to narrowly rounded, starting from 5th extending beyond rear tergal margin. Lateral calluses of paraterga rather wide, especially so on pore-bearing segments, each delimited by a sulcus both dorsally and ventrally. Ozopores lateral to dorsolateral, placed inside an ovoid groove. Tergal setae mostly abraded, retained ones about 1/5 the length of metaterga, pattern traceable at least as 1+1 paramedian insertion points in front row (Figs 7, 8). Axial line nearly wanting, vaguely traceable only here and there on metaterga. Transverse sulcus on metaterga rather superficial, nearly reaching bases of paraterga in metaterga 5-18. Stricture between pro- and metazona rather faintly striolate. Pleurosternal carinae small, complete or nearly so in segments 2-8, a sharp or rounded caudal denticle retained also in segments 9 and 10, thereafter entirely missing. Epiproct (Fig. 9) short, flattened dorsoventrally, digitiform; apical papillae small, tip faintly concave; subapical papillae evident, removed from tip. Hypoproct subtrapeziform, setigerous paramedian knobs at caudal margin well-separated and small.

Sterna rather densely setose; a rather high, linguiform, caudally rounded, ventral, setose lamina only between ♂ coxae 4 (Fig. 10). Legs relatively long, midbody ones ca 1.6-1.7 times longer than body height, apparently slightly incrassate and longer compared to ♀; ♂evident tarsal and distotibial brushes increasingly well reduced towards legs of segment 10.

Gonopods as in Figs 11-14. Tip of solenophore simple, faintly trifid, with three apical denticles or lobules. Femorite with a large rounded lobe (l) somewhat hollow on mesal side.