Eumenes truncatus Nugroho, 2010

Nugroho, Hari, Ubaidillah, Rosichon & Kojima, Jun-ichi, 2010, Potter Wasps Of The Genus Eumenes Latreille (Hymenoptera: Vespidae: Eumeninae) In The Western Part Of The Papuan Region, With Description Of Two New Species And Taxonomic Notes On E. Inconspicuus Smith, Raffles Bulletin of Zoology 58 (2), pp. 179-187 : 182-184

Eumenes truncatus Nugroho , new species

( Figs. 5 – 8 ; 11 – 12, 14, 17 – 18 )

Material examined. – Holotype – female ( MZB), labeled “ INDONESIA, Papua, Sentani , forest edge, 100–150 m, I. J. [= Irian Jaya expedition], 26 Mar.1973 ” (handwritten), “ HOLOTYPE, MZB. HYMN. 19213” and “ Eumenes truncatus Nugroho ” .

Paratype – 1 male ( MZB), INDONESIA, Papua, Manokwari, Ambau , 110 m, Irian Jaya expedition, 13 May 1973 .

Diagnosis. – This species can be separated from all other known species in the Papuan Region by the following combination of characters: clypeus with dorsal margin distinctly emarginate medially, produced dorsally beyond level of tentorial pits and very close to antennal socket, with ventral margin distinctly emarginate medially; labrum truncate apically; propodeum strongly convex, with deep median furrow; first metasomal sternum and tergum fused, with suture between them distinct only posteriorly; body black; female clypeus yellow dorsally, black ventrally.

Female. – Body length about 17 mm; forewing length about 12 mm.

Similar to E. batantanensis except for the following characteristics: clypeus with dorsal margin close to antennal sockets and deeply emarginate medially ( Fig. 7 ), ventral margin distinctly emarginate medially ( Fig. 11 ); labrum truncate apically ( Figs. 7 , 11 ); mandibular dorsalmost tooth more obtuse ( Fig. 7 ); mesosoma not so stubby as in E. batantanensis , about 1.2 times as long as high in lateral view, about 1.2 times as long as wide; mesoscutum nearly as wide as long; propodeum strongly convex, bulging in anterior part of posterior face, in lateral view outline of posterior face strongly curved ( Fig. 5 ), median longitudinal groove deeper; first metasomal tergum in dorsal view weakly produced laterally at spiracles; punctures on first and second metasomal segments denser than in E. batantanensis ( Figs. 17, 18 ). Antennal scape slender, weakly curved, 4 times as long as its maximum width; first flagellomeres about 3 times as long as wide; second to fourth flagellomere slightly longer than its maximum width; fifth to ninth flagellomeres square in outline, about as long as wide.

Body densely covered with long, ferruginous hairs. Head strongly punctured in frons and vertex. Pronotum and mesoscutum with punctures slightly larger than those on head. Scutellum, metanotum and propodeum with punctures stronger and larger than those on mesoscutum. First metasomal tergum and sternum with punctures sparser and smaller than those on head and mesosoma, but stronger than those on E. batantanensis ; second sternum with punctures superficial, sparser than those on second tergum.

Body black, with following yellow markings: Band along inner eye margin occupying eye emargination and extending ventrally to clypeus; supraclypeal area, dorsal half of clypeus, narrow band along apical margin of labrum, narrow band in dorsal half of gena along posterior margin of eye, anterior part of dorsal face of pronotum, narrow band in each posterolateral corner of pronotum, short lateral band along each anterolateral margin of mesoscutum, parategula, outer margin of tegula, anterior half of metanotum, large scrobal spot connected with large yellow spot below it to occupy posterior half of mesepisternum, paired large posterolateral spots on propodeum, propodeal valvula, apical band of first metasomal tergum, paired large lateral spots and apical band of second tergum, and large irregular-shaped posterior spot on second sternum. Apical margin of clypeus, labrum except for apical yellow band, mandible except for black base, ferruginous. Antenna blackish brown, but ferruginous beneath. Wings fusco-hyaline, darker than in E. batantanensis . All coxae black; femora dark brown, but fore femur largely and midfemur apically yellow; fore and mid-tibiae yellow; hind-tibia ferruginous; all tarsi ferruginous, darker apically.

Male. – Body length about 15 mm; forewing length about 10 mm. Similar to female, but clypeus more slender, about 1.3 times as high as wide, more strongly narrowed dorsally ( Fig. 12 ), entirely yellow except for apical, semitransparent, ferruginous band; space between clypeus and antennal socket wider than in female; mandible black but ferruginous at apex, with dorsalmost tooth longer and more slender than in female; labrum yellow; punctures on first and second terga sparser and weaker than in female. Antennal scape slightly curved, about 4 times as long as its maximum width (near apex); first flagellomere about 3.5 times as long as its maximum width; second to fifth flagellomeres slightly longer than their maximum width; sixth to tenth flagellomeres square, about as long as wide; terminal flagellomeres slightly curved, sharply pointed apically, about 2.2 times as long as its basal width ( Fig. 14 ).

Etymology. – The specific name, truncatus , is a Latin adjective and refers to the apically truncate labrum.

Remarks. – This new species is similar to E. batantanensis Nugroho , new species, but can be easily distinguished from the latter in the following characters [states for E. batantanensis are given in brackets]: Both dorsal and ventral margins of clypeus rather deeply emarginate medially ( Figs. 11 – 12 ) [emargination shallow or very shallow ( Figs. 9 – 10 )]; female clypeus close to antennal sockets ( Fig. 7 ) [distinctly separated ( Fig. 3 )]; propodeum more strongly convex ( Fig. 5 ) than in E. batantanensis ( Fig. 1 ); base of second tergum slightly flattened ( Fig. 5 ) [barely flattened ( Fig. 1 )]. This species is also similar to E. dorycus , according to the redescription of the lectotype by Giordani Soika (1941), but can be easily distinguished from E. dorycus by having the clypeus black in dorsal half (nearly entirely yellow in E. dorycus ). Furthermore, E. dorycus is distinguished from either E. truncatus and E. batantanensis by having the first metasomal segment much more elongated than the latter two species (see Giordani Soika, 1941: Fig. 19 )

Distribution. – Known from the two localities listed herein, Sentani near Jayapura and Manokwari. This species may occur in northwestern part of New Guinea Island.

Figures

Figs. 5–8. Eumenes truncatus Nugroho, new species, holotype female: 5, body, lateral view; 6, first metasomal tergum, dorsal view; 7, head, frontal view; 8, left antenna. Scale bar = 1 mm.

Figs. 9–18. Eumenes batantanensis Nugroho, new species (9–10, 13, 15–16) and E. truncatus Nugroho, new species (11–12, 14, 17–18). Female, holotype (9, 11, 15–16) and male (10, 12, 13–14). 9–12, Clypeus and labrum, 13–14, apical part of antenna, 15, 17, punctures on first metasomal tergum, 16, 18, punctures on second metasomal tergum. Scale bar (for only Figs. 15–18) = 1 mm.

Figs. 1–4. Eumenes batantanensis Nugroho, new species, holotype female: 1, body, lateral view; 2, first metasomal tergum, dorsal view; 3, head, frontal view; 4, fore femur and trochanter. Scale bar = 1 mm.

Figs. 19–23. Eumenes inconspicuus Smith. Male (19–22) and female (23). 19, metasoma, lateral view, 20, first metasomal tergum, dorsal view, 21, 23, clypeus, 22, apical part of antenna. Scale bar (for only Fig. 19) = 1 mm.