Episcomitra bouei ( Hoernes & Auinger 1880 ) Harzhauser & Landau, 2021

Harzhauser, Mathias & Landau, Bernard, 2021, The Mitridae (Gastropoda: Neogastropoda) of the Miocene Paratethys Sea, Zootaxa 4983 (3), pp. 1-72 : 10-11

Episcomitra bouei ( Hoernes & Auinger 1880) nov. comb.

Figs 3C 1 –C 2

[ Mitra ] Bouéi nov. form.—Hoernes 1880: 125 [nomen nudum].

* Mitra Bouéi nov. form.— Hoernes & Auinger 1880 (pars): 79, pl. 9, figs 6a–b [non figs 7a–b = Episcomitra pseudoincognita nov. sp.].

Mitra (Mitra) bouei Hö. Au. — Boettger 1906: 7.

Mitra (Tiara) bouei ( Hoernes & Auinger, 1880) — Atanacković 1985: 161, pl. 36, figs 7–8.

non Mitra Bouei R. Hoern. i Auinger— Friedberg 1911: 14, pl. 1, fig. 7 [= Episcomitra pseudoincognita nov. sp.].

non Mitraria (Mitraria) bouei ( Hoernes & Auinger, 1880) — Bałuk 1997: 29, pl. 8, figs 7–8.

Type material. Lectotype (designated herein): NHMW 1854 /0035/0100d, 16.4 mm, MD: 6.1 mm, Lăpugiu de Sus ( Romania), illustrated in Hoernes & Auinger (1880, pl. 9, figs 6a–b), figs 3C 1 –C 2.

Revised description. Shell small, moderately broad ovoid with cyrtoconoid spire and weakly impressed suture. Protoconch unknown. Teleoconch of six whorls. First two teleoconch whorls weakly convex with five convex, wide-spaced spiral cords. Spiral cords weakening on subsequent whorls, number of cords increasing to 13 on penultimate whorl by intercalation of secondary spiral threads. Last whorl high, ovoid, moderately constricted with shallow basal concavity. Periphery of last whorl coinciding with adapical tip of aperture. Mode of spiral sculpture changing from delicate densely spaced flattish spiral cords to widely spaced, crest-like spiral cords below periphery. Aperture moderately narrow. Columellar callus narrow, sharply delimited from adapical columellar fold to tip of siphonal canal. Columella with four oblique columellar folds, weakening abapically. Outer lip thin. Siphonal canal moderately short, moderately wide, straight with shallow siphonal notch.

Shell measurements and ratios. SL = 16.4 mm, MD: 6.1 mm; AA = 47°, SL/ MD: 2.7, AL/AW: 5.2, AH/S: 2.6.

Discussion. Hoernes & Auinger (1880) illustrated two syntypes, which in our opinion are not conspecific. To clarify the status of the species, we select the specimen from Lăpugiu de Sus ( Romania), illustrated by Hoernes & Auinger (1880, pl. 9, fig. 6), as lectotype of Episcomitra bouei , and exclude the specimen from Jerutek at Lysice ( Czech Republic) (1880, pl. 9, fig. 7), described herein as Episcomitra pseudoincognita nov. sp.

Episcomitra facilis (Mayer, 1890) , which co-occurs with E. bouei at Lăpugiu de Sus ( Romania), differs in its more ovoid profile, the coarser spiral sculpture and weakly canaliculate suture.

Palaeoenvironment. Unknown.

Distribution in Central Paratethys. Badenian (middle Miocene): Făget Basin: Lăpugiu de Sus ( Romania) ( Hoernes & Auinger 1880); Southern Pannonian Basin: Hrvaćani ( Bosnia and Herzegovina) ( Atanacković 1985).

Figures

FIGURE 1. A. Map of central and south–eastern Europe, representing the area that was partly covered by the Central Paratethys Sea (for detailed maps with the localities mentioned in the text see Kroh (2005) and Nosowska (2020). The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Palaeogeography of the Middle Miocene Paratethys Sea (modified from Harzhauser & Landau 2017) showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, E–Sopr B: Eisenstadt–Sopron Basin).

FIGURE 2. Measurements for shell morphology and whorl profiles. SL: shell length, MD: maximum diameter, AA: apical angle, LWH: last whorl height, AH: aperture height. AL: aperture length. AW: aperture width. S: length of siphonal canal.

FIGURE 3. A1–A2. Episcomitra antibellardii nov. sp., holotype, Lăpugiu de Sus (Romania), NHMW 1854/0035/0100b. B1–B2. Episcomitra antibellardii nov. sp., paratype, NHMW 1854/0035/0100c. C1–C2. Episcomitra bouei (Hoernes & Auinger 1880), lectotype, Lăpugiu de Sus (Romania), NHMW 1854/0035/0100d. D1–D2. Episcomitra cochlearella (Mayer-Eymar, 1890), lectotype, Lăpugiu de Sus (Romania), NMB Inv. Nr. t3238). E1–E2. Episcomitra cochlearella (Mayer-Eymar, 1890), paralectotype, Lăpugiu de Sus (Romania), NMB Inv. Nr. t3238. F1–F2. Episcomitra facilis (Mayer-Eymar, 1890), holotype, Lăpugiu de Sus (Romania), Inv. Nr. t3308. G1–G2. Episcomitra hilberi (Hoernes & Auinger 1880), paralectotype, Lăpugiu de Sus (Romania), NHMW 1854/0035/0100. H1–H2. Episcomitra hilberi (Hoernes & Auinger 1880), lectotype, Jerutek at Lysice (Czech Republic), NHMW 1865/0015/0014. I1–I2. Episcomitra pilsbryi (Boettger, 1906), holotype, Coşteiu de Sus (Romania), SFM 360347 (= SFM XII 12.2207a).