Epigambrialongipennis Handlirsch, 1939

Makarkin, Vladimir N., Ansorge, Jörg & Khramov, Alexander V., 2021, Revision of Epigambriinae Handlirsch, stat. nov., a subfamily of Early Jurassic Ithonidae s. l. (Neuroptera), Palaeoentomology 4 (6), pp. 516-531 : 517-518

Epigambrialongipennis Handlirsch, 1939

( Fig. 1 )

Epigambrialongipennis Handlirsch, 1939: 76 , Pl. 7, Fig. 119; Martynova, 1949: 167; Lambkin, 1988: 449, 453; Herrig & Nestler,1989:33; Carpenter,1992:355; Ross& Jarzembowski, 1993: 418; Makarkin & Archibald, 2003: 176.

Material. Holotype FGWG 123/51, deposited in the collections of the Institute of Geography and Geology, University of Greifswald, Germany. Apoorly preserved crumpled complete hind wing. Germany: Dobbertin. Early Jurassic (early Toarcian, “Grüne Serie”, Harpoceras falciferum ammonite zone).

Re-description. Hindwing 6.9mmlong,1.8 mmwide aspreserved (estimated completewidth ≥ 2 mm). Costal space relatively broad, slightly broadened proximally, moderately broadened distally. Subcostal veinlets widely spaced; basally simple, perpendicular to Sc; more proximally bent toward wing apex, simple or shallowly forked; distally once forked. Subcostal space moderately broad; crossveins not detected. Sc stout, approaching RA distally, with or without fusion unknown. RA stout, relatively long, entering margin somewhat proximad wing apex; with four long veinlets, once to twice forked. RA space narrow, with four preserved crossveins (nearly perpendicular to RA). RP originating relatively near wing base, with seven pectinate branches. RP 1, RP 3, RP 4, RP 6 deeply dichotomously forked; RP 2 similarly (but not so deep) branched; RP 5 once forked. Proximal part of Mnot preserved. MA, MP deeply dichotomously forked. CuA pectinately branched; its anterior trace, three branches once forked. CuP not clearly discernible (possibly few branched). Anal veins very poorly preserved. Crossveins in radial space very poorly preserved, probably forming one gradate series (three crossveins discernible).

Remarks. The drawing of Handlirsch ( Handlirsch, 1939: Pl. 7, Fig. 119) is largely incorrect, especially in the area of the cubital space; we revise it here ( Fig. 1 ).

Figures

FIGURE 1. Epigambria longipennis Handlirsch, 1939, holotype FGWG 123/51. A, Part. B, counterpart. C, Hind wing venation. Scale bars = 1 mm (A, B to same scale).