Eocypraea judithsmithae, Groves & Squires, 2023

Groves, Lindsey & Squires, Richard L., 2023, Revision of northeast Pacific Paleogene cypraeoidean gastropods, including recognition of three new species: implications for paleobiogeographic distribution and faunal turnover, PaleoBios 40 (10), pp. 1-52 : 24

EOCYPRAEA JUDITHSMITHAE GROVES AND SQUIRES N. SP.

FIGS. 7M–O

Zoobank ID — LSID: urn: lsid: z ooba nk. org:act: 515D4573-172F-457E-AAB9-BD37AC242CD9

Diagnosis— Inflated-pyriform shell of small to medium size; spire involute; narrow elongate aperture; fossula broad, smooth, concave.

Holotype and Type Locality— LACMIP 40372.1 , LACMIP Type 14940 ( Figs. 7M–O ), length 26.7 mm, width 18.3 mm, height 13.8 mm. LACMIP Locality 40372 [ex CSUN Locality 372], Llajas Formation , approximately 25 m stratigraphically below “Stewart bed,” Devil Canyon (formerly known as Aliso Canyon), southwestern Santa Susana Mountains , Los Angeles County, California.

Paratype — LACMIP 7171.1 , LACMIP type 14926, length 24.8 mm, width 18.4 mm. height 13.3 mm, LAC- MIP Locality 7171 [ex CIT Locality 1074], Domengine Formation, Domengine Ranch , Fresno County, California. The paratype is worn but has a well preserved labial lip .

Occurrence— Llajas Formation, upper lower Eocene, Devil Canyon (formerly Aliso Canyon), southwestern Santa Susana Mountains , Los Angeles County, southern California and Eocene Domengine Formation, Fresno County, California .

Etymology— Named in honor of friend and colleague Judith T. Smith ( USNM, Paleobiology) for her numerous contributions to molluscan paleontology.

Description— Shell of medium size, noticeably inflated and smooth. Constricted anteriorly, spire completely covered, dorsum highly arched, maximum height slightly posterior of center, maximum width nearly center, aperture slightly s-shapped, curves towards columella posteriorly, denticulation semi-coarse, outer lip with 18 teeth, inner lip with 13 teeth (tooth count incomplete due to missing posterior portion of inner lip), fossula smooth and wide, anterior and posterior canals deep, anterior terminal ridges short, base rounded, very slight basal callus on outer lip.

Remarks— The holotype has good preservation and the paratype has moderately good preservation. This new species resembles closely the middle Eocene Eocypraea inflata ( Lamarck, 1802) , known from France, but the new species differs by having being less globular in shell shape and having weaker denticulations on both lips of the aperture.Two additional specimens ( LACMIP 7171.2) from the paratype locality ( LACMIP Locality 7171 [ex CIT Locality 1074] are poorly preserved specimens with little original shell material.

Figures

Figure 7A–C. Cypraeidae, genus and species indeterminate, Lodo Formation (basal) (Paleocene, Seldanian Stage), Tumey Hills, Fresno County, central California. Basal (A), dorsal (B), and right (labral) (C) views, 1.3x, length 39 mm, width 25 mm, height 22 mm, of hypotype CASG 61666.06, CASG Locality 61666. D–F. Eocypraea batequensis (Groves, 2011), Bateque Formation (Eocene, “Capay Stage”), Mesa La Salina, Baja California Sur, México. Basal (D), dorsal (E), and right (labral) (F) views, 4.8x, length 9.4 mm, width 6.5 mm, height 5.1 mm, of holotype IGM 5174, LACMIP Locality 16951 [ex CSUN Locality1220b]. G–I. Eocypraea crescentensis (Groves, 2011), Crescent Formation (Eocene, “Capay Stage”), Larch Mountain, Black Hills area, Thurston County, Washington. Basal (G), dorsal (H), and left (columellar) (I) views, 3.9x, length 10.9 mm, width 7.6 mm, height 6.1 mm, of holotype LACMIP 16655.40, LACMIP Type 13646, LACMIP Locality 16655 [= LACMIP Locality 41563; ex CSUN Locality 1563]. J–L. Eocypraea jimgoederti (Groves, 2011), lower McIntosh Formation (middle Eocene, “Transition Stage” to “Tejon Stage”), Doty Hills, Lewis County, Washington. Basal (J), dorsal (K), and left (columellar) side (L) views, 3.5x, length 12.7 mm, width 8.2 mm, height 6.9 mm, of holotype LACMIP 41573.1, LACMIP Type 13647, LACMIP Locality 41573 [ex CSUN Locality 1573]. M–O. Eocypraea judithsmithae n. sp., Llajas Formation (Eocene, “Domengine Stage”), north side Simi Valley, Ventura County, southern California. Basal (M), dorsal (N), and left (labral) (O) views, 1.5x, length 26.6 mm, width 18.4 mm, height 12.6 mm, of holotype LACMIP 40372.1, LACMIP Type 14940 (LACMIP Locality 40732).