Enodinympha Ren & Engel, 2007

Winterton, Shaun L., Martins, Caleb Califre, Makarkin, Vladimir, Ardila-Camacho, Adrian & Wang, Yongjie, 2019, Lance lacewings of the world (Neuroptera: Archeosmylidae, Osmylidae, Saucrosmylidae): review of living and fossil genera, Zootaxa 4581 (1), pp. 1-99 : 18-19

Enodinympha Ren & Engel, 2007

( Fig. 7C )

Type species. Enodinympha translucida Ren & Engel, 2007: 214 (by original designation).

Diagnosis. Antenna distinctly longer than forewing; scape elongate; wings narrow, medium sized; forewing with numerous crossveins throughout wing, regularly arranged but not forming gradates; RP with 12 branches; RP1 originating distant from origin of RP, approximately 1/3 wing length; M fork proximal to origin of RP1, closer to origin of RP, near 1/6 wing length; MA unbranched except for end-twigging near wing margin; MP with few pectinate branches near wing margin; CuA pectinately branched near wing margin, 4–5 branches, some with endtwigging; CuP strongly pectinate (nine simple branches); A1 pectinate with six branches; HW with numerous crossveins throughout wing; MP pectinate; CuA strongly pectinate (13 branches); CuP simple.

Comments. Placement of this genus in Gumillinae is clear based on the long antennae, narrow costal margin and distal origin of RP1. The forewing medial vein forking very close to the wing base is uncommon among genera of Gumillinae , as it typically forks closer to the middle of the wing. The relatively large number of RP branches (12) and numerous regular crossveins is also uncommon in Gumillinae , as most members of this subfamily have 5– 7 (or fewer) branches and far fewer, irregular crossveins. This genus is likely closely related to Tenuosmylus and Epiosmylus . Li et al. (2018) recently transferred Nilionympha imperfecta to Enodinympha .

Figures

FIGURE 7. Gumillinae fossils. A. Allotriosmylus uniramosus Yang et al. (from Yang et al., 2010); B. Epiosmylus panfilovi Ren & Yin (from Ren & Yin, 2002); C. Enodinympha translucida Ren & Engel (forewing); D. Nuddsia longiantennata Menon & Makarkin (from Menon & Makarkin, 2008); E. Nilionympha pulchella Ren & Engel (from Ren & Engel, 2007); F. Tenuosmylus brevineurus Wang et al. (from Wang et al., 2009b) (figures not to scale).