Empis (Enoplempis) montywoodi Brooks

Sinclair, Bradley J., Brooks, Scott E. & Cumming, Jeffrey M., 2013, Revision of the Empis subgenus Enoplempis Bigot, east of the Rocky Mountains (Diptera: Empididae), Zootaxa 3736 (5), pp. 401-456 : 425-428

Empis (Enoplempis) montywoodi Brooks sp. nov.

( Figs 12 A, B, 13A, 18B)

Type material. HOLOTYPE, ♂ labelled: “CAN: QC: Gatineau/ Masham, Duncan Lk/ N 45°40'59.43'' W 76°3'21.38''/ 21.vi.1993 / D.M. Wood”; “ HOLOTYPE / Empis (Enoplempis) / montywoodi / Brooks [red label]” (CNC).

Diagnosis. This species is distinguished by the male hind tibia bearing a pair of blunt basiventral processes, each with narrow brush of long, black apical setae.

Description. Wing length 7.0 mm. Male. Head mostly dark in ground-colour, with dense greyish pruinescence on face, frons and occiput, postgena pale yellow. Dichoptic, eye with ommatidia of similar size. Frons parallelsided, subequal to width of ocellar triangle, bearing short dark setulae along inner margin of eye. Face slightly divergent towards mouthparts; bare with oral margin pale. Ocellar triangle dark, subshining, with pair of short, parallel, proclinate ocellar setae and several shorter posterior setulae. Occiput bearing 2 strong, black upper postocular setae with remainder of postocular row comprised of short black setae; occipital setae black, similar in size to strong upper postoculars. Postpedicel and stylus dark; scape, pedicel and extreme base of postpedicel palebrownish. Scape 2× longer than pedicel; postpedicel about 3.7× longer than basal width, about 2× longer than stylus. Palpus pale yellow, with dark setulae. Proboscis largely pale; apex and base of labrum darkened; labellum with dark setae.

Thorax mostly yellowish in ground-colour but with darkened scutum; postpronotum, notopleuron, postalar ridge, posteromedial region of scutum and scutellum yellowish brown. Scutum with pair of distinct blackish vittae between acr and dc rows; distinct pair of vittae lateral to dc and above notopleural depression; with greyish pruinose vittae along acr and dc rows. Pleura entirely yellowish. Proepisternum with several short, dark setae on upper part; prosternum bare. Antepronotum with row of short dark setae. Postpronotum with several short dark setae and 1 long seta. Scutum with inconspicuous, uniserial row of fine acr setulae ending at transverse suture; dc row slightly stronger than acr; 1 strong posterior npl, with several weak anterior npl setae; 1–3 weak presut spal; 1 weak psut spal; 1 strong pal. Scutellum with pair of short sctl, similar to pal, 1–2 marginal setulae. Laterotergite with 2 long dark setae and 1 lower setula. Anterior and posterior spiracles pale.

Legs long, entirely yellow, except for dark ring at trochanter-femur junction and apices of femora; tarsi becoming darker apically. Coxae with numerous dark lateral and apical setae, longer and stouter along apical anterior margins. Hind trochanter lacking spine-like or modified setae. Femora clothed in short dark setae, setae longer at apex, mid and hind femora with distinct anterior preapical seta. Hind femur slender in proximal 2/3, apical third slightly swollen bearing anteroventral brush-like tuft of setae adjacent to posteroventral thumb-like process ( Fig. 12 A), with more distally positioned conical process ventrally. Fore tibia clothed in dark setae, shorter than width of tibia, setae dense and shorter along anterior surface; 3 posteroventral setae and 4 preapical setae. Mid tibia clothed in dark setae, shorter than width of tibia; 2–3 anterodorsal, 2 posterodorsal, 2 posteroventral and 6 preapical setae. Hind tibia weakly bowed medially, clothed in dark setae, shorter than width of tibia; basally with pair of blunt ventral processes, each with narrow brush of long, black apical setae ( Figs 12 A, B), distal process with anteroventral fringe of long setae proximal to base; 3 anterodorsal, 4 posterodorsal setae and 5 preapical setae. Tarsomeres 1–5 of all legs with rows of antero- and posteroventral spine-like setae; fore tarsomere 1 slender, not swollen.

Wing clear with brownish veins; all veins complete (except Sc), well sclerotized. Basal costal seta indistinct from surrounding costal setae. R5 and M1 parallel near wing margin; R5 ending beyond wing tip; radial fork acute. Halter yellow.

Abdominal tergites 1–7 mainly dark, concolorous with scutum with marginal setae distinct and dark; sternites 1– 7 pale with pale setae. Segment 8 with marginal setae dark, longer than proceeding segment. Pregenital segments unmodified; sclerites of segment 8 weakly fused to form complete ring, not weakly sclerotized dorsally. Terminalia ( Fig. 13 A) largely yellow. Cercus narrow, appressed to dorsal margin of epandrium, linear, broader at anterior end; dorsal margin lacking tooth-like projection at inner apex; posterior end simple, truncate; clothed in setae. Hypoproct with several long setae. Epandrial lamella largely clothed with short setae, posterior margin rounded. Hypandrium prolonged posteriorly with rounded margin, about 0.5× as long as epandrium; with numerous long setae, about equal to length of hypandrium. Phallus with broad base, apical portion straight; apex just emerging beyond cercus; ejaculatory apodeme slightly shorter than length of epandrium; inverted Y-shaped, with short median keel.

Female. Unknown.

Geographical distribution and seasonal occurrence ( Fig. 18 B). Empis (En.) montywoodi is known only from the holotype collected from Gatineau, Quebec in mid-June.

Etymology. This species is named after colleague, mentor and friend Dr. Monty Wood who collected the unique holotype.

Nuptial gift presentation. Form unknown.

Figures

FIGURE 12. Male legs of E. (Enoplempis). A, E. montywoodi, apex of hind femur and base of hind tibia, anterior view; B, E. montywoodi, apex of hind femur and base of hind tibia, posterior view; C, E. nodipoplitea, apex of hind femur and base of hind tibia, anterior view; D, E. nuda, hind trochanters; E, E. nuda, hindleg, anterior view.

FIGURE 13. Male terminalia, lateral view of E. (Enoplempis). A, E. montywoodi; B, E. nodipoplitea.

FIGURE 18. Distribution of E. (Enoplempis). A, E. nuda and E. prodigiosa; B, E. enodis, E. montywoodi, E. pectinata, and E. penicillata.