Drasteria rada altivaga Alphéraky, 1894

Yu, Alexey & Korb, Stanislav K., 2019, A revision of the genus Drasteria of Central Asia and Kazakhstan with special attention to the adjacent areas (Lepidoptera: Erebidae), Zootaxa 4673 (1), pp. 1-104 : 32-33

Drasteria rada altivaga Alphéraky, 1894

(= D. rada schlumbergeri Fuchs, 1903 , = D. rada sibirica Kozhnatchikov, 1924 ) ( Figs. 79–90 (imago), 228, 229 (male genitalia), 277 (female genitalia))

Alphéraky, 1894: 347 ( Leucanitis Altivaga ). Type locality: “Kaschgaria. Altit.: 9000’” (by the original description); “Kaschgaria, Russian Mts.” (by the holotype data).

Type material. Holotype (by monotypy) ( Figs. 82–84 ), labelled: “18.” (white paper, handwritten); “Кол. Вел. Кн. | НиколаЯ | Михаиловича” <Collection of the Great Duke Nikolai Mikhailovich> (white paper, printed); “slide 0001 | Matov” (white paper, printed); “Kaschgaria | Русскiй хр.” [Kashgaria, Russian Mts.] (green paper, handwritten); “9000’ a.l. | 11 VII 1890.” (green paper, handwritten) ( ZISP).

= schlumbergeri Fuchs, 1903: 50 ( Leucanitis rada B. var schlumbergeri ). Type locality: “aus Aksu ” (by the original description).

Type material. Not found.

= sibirica Kozhantshikov, 1924 ( Leucanitis roda sibirica ). Type locality: “Minussinsk” (by the original description).

Type material. Holotype male ( Figs. 88–90 ), labelled: golden circle; “Leuc.[anitis] roda | sibirica m. det. | W Koshantschikov” (white paper, handwritten 1 st and 2 nd line, printed 3 rd line); “Окр. Минусинска | 7.VI | 1924 КоЖанчиков» <Minusinsk vicinity, 7.06.1924, Kozhanchikov> (white paper, letters printed, digits handwritten) ( ZISP).

Staudinger, 1882: 53 [Saisan]; Staudinger, Rebel, 1901: 243 [Saisan, Ili]; Hampson, 1913: 284 [Saisan, Ili]; Warren W., 1913: 342 [Saisan, Ili]; Shekh, 1975: 411 [Central Kazakhstan, Alma-Ata Prov]; Remm, Viidalepp, 1979: 76 [Kyzyl]; Ronkay, 1983: 234 [Bajanchongor aimak: Oase Echin gol, cca. 90 km NO von Grenzposten Caganbulag, 950 m; zwischen Somon Bajangobi und Somon Bajanleg, 26 km SO von Bajanleg, 1450 m; Bulgan aimak: SO con Somon Dashincilen; Central aimak: 25 km O von Somon Lun, 1200 m; Chovd aimak: Chovd (Kobdo), cca 5 km SW von der Stadt, 1500 m; Südgobi aimak: Nojon nuruu Gebirge, unweit von Dzun adu chudag, 34 km NO von Grenzposten Ovot Chuural, 1800 m; Nojon nuruu Gebirge, Grenzposten Ovot chuural, 1500 m; 100 km W von der Grenzposten Ovot Chuural, 22 km W von Sajryn chudag, 1250 m; Tachilga ul Gebirge, zwischen Somon Cogt-Ovoo und Dalanzadgad, 68 km S von Cogot-Ovoo, cca 1550 m; Uvs aimak: Senke des Sees Uvs nuur am SW-Rand des Sees, 84 km W von Somon Zuungobi und 63 km O von der Stadt Ulaangom, 790 m; Mogoin arschaan, N-Rand des Sees Chajargas nuur, 48 km OSO von Somon Naranbu-lag, 1100 m]; Nekrasov et al., 1988: 51 [Akulen, Pokrovka, Chon-Dzhargylchak]; Poole, 1989: 330; Milko, 1996: 258 [Issyk-Kul depression (including northern slopes of Terskey Ala-Too Mts., southern slopes of Kungey Ala-Too Mts. and Tyup river valley)]; Lehmann et al., 1998: 526 [Toraygyr Mts., 1400 m, 43.19’22”N/78.55‘96”O; Ili area, Charyn river, 400m; 42.10’N. 77.90’E, S Ufer des Issyk Kul-Sees. ca 10 km W Tasor; Sarikamysch Mts., River Kjokjomeren, 5 km n Amok, 41.45’N/74.10”O]; Lehmann, 2000: 95 [Jurievka bei Syn-Tasch, 42°42’ N, 75°03’ E]; Lehmann, Bergmann, 2005: 20 [Issyk Kul depression, Susamyrtoo, Dzhumgaltoo, Sarykamish Mts., Kirgizsky Mts.]; Bálint et al., 2006: 107 [ Mongolia, Övörkhangay aimak, Khan Khögshin Uul, 81 km SW of Arvaykheer, N 45°51’, E 101°55’, 1890 m; Mongolia, Gov’-Altay aimak, Mongolian Altay Mts., Sutay Uul, 16 km SE of Dzuyl, N 46°11’, E 94°01’, 2070 m; Mongolia, Khovd aimak, Dzhungarian Gobi, 81 km km SW of Dzuyl, N 45°45’, E 93°15’, 1810 m; Mongolia, Gov’-Altay aimak, Mongolian Altay Mts., Sutay Uul, 16 km SE of Dzuyl, N 46°11’, E 94°01’, 2070 m]; Bálint, Benedek, 2009: 7 [ Mongolia, Töv aimag, 58 km north of Delgertsogt, N46°38.154’, E106°33.128’, 1,374 m]; Benedek, Bálint, 2009: 163, 165 [Prov. Almaty, Toraygir Mts, near Alasay pass, 1340 m, N43°18’ E078°56’; Prov. Almaty, Syugeti Mt., 8 km NW of Kok-Pek, 980 m, N43°29’; E078°37’];:165 [Prov. Almaty, Boguty Mt., 830 m, N43°34’ E078°39’]; Benedek, Bálint, 2013: 138 [Province Almaty, 20 km NW of Kapchagai, 650 m, N 43° 58’, E 077° 01’]; Korb, 2013: 28 [Chulak gorge in Dzhungarian Alatau Mts., Ili River Valley, Altyn Emel Nature Reserve, N 43 54.255 E 77 47.980, 681 m; Singing Dune in Altyn Emel Nature Reserve, eastern border of Kapchagai Reservoir, Ili River Valley, N 43 50.273 E 78 35.759, 467 m; Saksaul forest 56.5 km S of Kugaly, Altyn Emel Nature Reserve, N 43 57.688 E 78 46.164, 645 m; Charyn National Park, Charyn River Valley, N 43 38.124 E 79 21.641, 638 m]; Shovkoon, Trofimova, 2016: 607 [Aktobe Prov., Mugodzhary Mts., N 48°32’; E 58°43’; Aktobe Prov., Mugodzhary Mts., N 48°11’; E 58°17’; Aktobe Prov., N 47°59’ E 59°24’; Aktobe Prov., Bolshie Barsuki sands, N 46°58’; E 59°13’; Aktobe Prov., Shagyrai Plateau, N 46°52’; E 58°02’; Aktobe Prov., Shagyrai Plateau, N 46°50’; E 57°47’; Aktobe Prov., Shagyrai Plateau, N 46°45’; E 57°31’; Kyzyl-Orda Prov., Karatup penin., Aral lake, N 46°20’; E 59°43’; Aktobe Prov., near Bozoe vill., sands, N 46°17’; E 58°56’; Mangystau Prov., N 46°05’; E 54°35’; Mangistau Prov., Ustyurt Plateau, N 43°57’; E 53°45’; Mangistau Prov., Ustyurt Plateau, N 43°55’; E 53°34’]; Makhov et al., 2018: 245 [Buryatia: Gusinoe].

Habitat ( Figs. 298, 299–304 ). Lowlands and mountain foothills, deserts and semideserts, sometimes (in mountains) steppes, but mostly on sands. Elevation: 200–1800 m.

Distribution. South Siberian Mountains, South Kazakhstan (Saisan and its environs, Saur, Tarbagatai, Dzhungar and Boro-Khoro, Ili and Charyn river valleys), West and Central Mongolia, northern part of Sinjan-Uigur autonomous region of China, foothills of North and West Tian-Shan, Ferghana valley, deserts of Uzbekistan, Turkmenistan and North Iran.

Material examined (in total: 117 specimens). China. 1 ♂, Tura , leg. Grum-Grzhimailo, ex coll. Grand Duke Nikolai Mikhailovich ( ZISP) ; 1 ♀, 23.04.1874, Kuldja, Talki , leg. S. Alphéraky ( ZISP) ; 1 ♂, 11.07.1890, Kasch- garia, Русский хр. [ Russky Mts. ], h=9000’, leg. Pevtsov, slide Matov 0001 ( ZISP) (holotype) ; 1 ♂, 3.07.[?], ур. Арчалын-удЗур [ Archalyn-Udzur valley ], ex coll. Grand Duke Nikolai Mikhailovich ( ZISP) ; 1 ♂, 1902, Saichin- Kette nördl. von Korla, Rückbeil (holotype of pia Strand, 1917) ( ZMHU) . Kazakhstan. 3 ♂, 1897, Asia centr., Ili ( ZMHU) ; 1 ♀, 3.08.1904, West Tarbagatai, Kara-Kitat river , southern foothills, <no collector data> ( ZISP) ; 1 ♀, 22.05.1910, Semiretchye Prov., Kopalsky uezd, Kutun’ -tay station, Nikolaevskoe settlement, leg. N. Tchetverikov ( ZISP) ; 8 ♂, 9 ♀, 1915, Semiretchye Prov., Dzharkent , leg. A. Dyakonov ( ZISP) ; 1 ♂, 1 ♀, 17.05.1934, Transil- ian Alatau Mts. , Issyk, leg. Slivinsky ( ZMMU) ; 1 ♂, 9.10.1936, Alma-Ata Prov., Ili river , Iliysk, leg. Slivinsky ( ZMMU) ; 1 ♀, 21– 22.10.1953, Kapchagai on Ili river, leg. Rustambekova ( ZISP) ; 1 ♂, 2 ♀, 11– 16.06.1988, 150 km NE Alma-Ata, right bank of Ili river, Mynbulak valley , leg. M.I. Falkovich ( ZISP) ; 1 ♀, 2– 7.06.1989, Aidarly , leg. Amirguzhin ( ZISP) ; 1 ♂, 21.05.1990, Dzhambulskaya obl. , 70 km NNE Frunze, 43 º 24’ N 75 º 02’E, [leg. A.V. Nekrasov?], slide N 2263 [glycerol], ex coll. A.V. Nekrasov ( ZISP) ; 1 ♀, 12.05.1991, Khantau, Balkhash See , leg. Danilevsky ( HNHM) ; 1 ♂, 7– 15.05.1994, Prov. Almaty, Bayseit, Chilik valley , 700 m, 78º21´E, 43º14´N, leg. Gy. Fabian and I. Retezar ( HNHM) ; 1 ♂, 1 ♀, 27– 31.05.1994, Prov. Almaty, Zailisky Alatau, Bokaydin-tau 4 km S Malibay, 850–1450 m, 78º24´E, 43º26´N, leg. Gy. Fabiani and I. Retezar ( HNHM) ; 2 ♂, 2– 3.05.2010, Charyn river valley, Charyn relict forest, 638 m, 43°38’07.5” N, 79° 21’38.5” E, leg. S.K. Korb ( SKK) ; 1 ♂, 5.05.2010, Karatau Mt. Range, Besaryk River , 35 km N of Talap, h= 440 m, 43 º 49’ N 67 º 41’ E, leg. P. Gorbunov ( ZISP) ; 1 ♀, 6– 8.05.2010, Karatau Mt. Range, Kurkal dry valley, 40 km N of Igilik, h= 540 m, 43 º 47’ N 68 º 03’ E, leg. P. Gorbunov ( ZISP) ; 1 ♂, 29.06.2010, Karatau Mt. Range, Zhilagan-Ata , 1100–1200 m, 43 º 50’ N 68 º 21’ E, leg. P. Gorbunov ( ZISP) ; 1 ♀, 25.04.2011, Almaty Prov., 32 km NW of Bakanas, Ili river , leg. S.A. Knyazev ( CKO) ; 2 ♂, 30.04.2011, Dzhungarian Alatau Mts., Chulak mountain gorge, 823 m, 43° 56’27.1”N, 77°48’16.3”E, leg. S.K. Korb ( SKK) ; 3 ♂, 23– 24.04.2011, Almaty Prov., 5 km N of Kapchagai , leg. S.A. Knyazev ( CKO) ; 1 ♂, 30.04.2011, Alma-Ata Prov., Altyn-Emel Nature Reserve, Singing Dunes , near Ili river , 577 m, leg. S.K. Korb ( SKK) ; 1 ♂, 1.05.2011, Altyn-Emel Nature Reserve , saksaul 43 36147N, 78 76941E, 645 m, leg. S.K. Korb ( SKK) . Kyrgyzstan. 1 ♂, Kyrgyzstan , leg. N. Danilov ( ZISP) ; 2 ♂, 2 ♀, 15.08.1955, southern shore of Issyk-Kul lake, Chongdzhargylchak , leg. A.V. Zvetaev ( ZMMU) ; 1 ♂, 17– 24.07.1986, Frunze, Chon-Aryk , leg. A. Kondratyev, ex coll. A.V. Nekrasov ( ZISP) ; 1 ♀, 8– 9.07.2000, Issyk-Kul, prope Kadzhi-Saj , 1650 m, leg. E. Rutjan ( OPB) ; 1 ♂, 29.07.2009, Terskey Ala-Too Mts., Kok-Moynok environs, 42°26’18”N, 75°50’39”E, 1700 m, leg. S.K. Korb ( SKK) ; 2 ♂, 09.07.2014, Suusamyrtoo Mts., Kekemeren river valley, 41°59.211’N, 74°09.396’E, 1700 m, leg. S.K. Korb ( SKK) . Mongolia. 1 ♂, 26– 28.07.1967, South Gobi aimak, 30 km NNE Bulgan, Baiin-Dzag Mts. , leg. I.M. Kerzhner ( ZISP) ; 1 ♂, 31.07.1967, South Gobi aimak, 60 km WNW Bayan-Dalay, Khongoryn-els , 30-leg. I.M. Kerzhner ( ZISP) ; 1 ♀, 31.07.1967, South Gobi aimak, 20 km WNW Bayan-Dalay, leg. V.F. Zaycev ( ZISP) ; 1 ♀, 2.08.1967, South Gobi ai- mak, 10 km E dwell Sudzhiyn-khuduk , leg. V. Zaitsev ( ZISP) ; 1 ♂, 17– 18.08.1967, Bayan-Khongorsky aimak, Ikh- Bogdo , 15 km SW Orog-Nur lake, h= 1700 m, leg. I.M. Kerzhner ( ZISP) ; 1 ♂, 4.08.1969, Uver-Khangaisky aimak, near eastern shore of the lake Tazyn-Cagan-Nur, leg. I.M. Kerzhner ( ZISP) ; 1 ♀, 20.08.1969, South Gobi aimak, near Dund-Gol settlement, leg. I.M. Kerzhner and M.A. Kozlov ( ZISP) ; 2 ♂, 22.07.1970, Kobdo aimak, Bodon- chin-Gol river, 12 km SW Altai (low), leg. I.M. Kerzhner ( ZISP) ; 1 ♂, 28.07.1970, Kobdo aimak, 10 km N Uench, leg. I.M. Kerzhner ( ZISP) ; 3 ♀, 5.08.1970, Ubsunur aimak, Ubsu-Nur , 50 km E Ulangom, leg. I.M. Kerzhner and Chogsomzhav ( ZISP) ; 1 ♀, 29.06.1971, East Gobi aimak, Dulan-Khara Mt. , 18 km SE Tal-Khongoryn-Khuduka, leg. I.M. Kerzhner ( ZISP) ; 1 ♂, 2.07.1971, East Gobi aimak, 50 km ENE Sayn-Shanda, leg. I.M. Kerzhner ( ZISP) ; 4 ♂, 5 ♀, 4.08.1971, East Gobi aimak, 30 km SSE lake Shokhoy-Nur, Nomt-Ula Mt. , leg. M.A. Kozlov ( ZISP) ; 10 ♂, 4 ♀, 27.06– 3.08.1981, Bayan-Khongorsky aimak, 140 km S Shine-Dzhinsta, Ekhiin-Gol , leg. A.L. Lvovsky ( ZISP) ; 1 ♀, 17.07.1981, Bayan-Khongorsky aimak, 20 km S Shine-Dzhinsta, steppe-desert, leg. A.L. Lvovsky ( ZISP) ; 1 ♀, 13.07.1987, Khoduut Khar , Dayryn-Am, leg. S. Dorzh ( ZISP) ; 1 ♂, 24– 26.07.1987, Bayanhongor aimak, Mts. Ih Bogd Uul, 1850 m, valley of Pitut river , 100º13´E, 45º00´N, leg. L. Peregovits, M. Hreblay, T. Stéger ( HNHM) ; 1 ♂, 3.06.1997, Ömnögovi aimag, Mts. Tost , 42 km WSW Gurt, 2450 m, 43º11´11´´N, 100º36´60´´E, leg. L. Lȍkös and L. Peregovits ( HNHM) ; 1 ♂, 25.06.2005, Ömnögovi aimag, Khan Khögshin Uul , 81 km SW of Arvaykheer, 45º51´N, 101º55´E, leg. B. Benedek and T. Csövár ( HNHM) . Russia. 1 ♂, 2 ♀, Minusinsk , leg. N. Danilov ( ZISP) ; 1 ♂, 7.06.1924, Minusinsk vic., Tagarsky island , leg. V. Kozhantchikov ( ZISP) (holotype of D. sibirica Kozhantschikov, 1925 ) ; 1 ♂, 15.05.1990, Tuva, 5 km W Kyzyl, Ulug-Khem river valley, leg. V.V. Dubatolov ( HNHM) ; 1 ♂, 26– 27.07.2015, Selenginsky distr., 18 km SW of Gusinoozyorsk, lake Gusinoe , 51011’37,14” N 106015’53,35” E, leg. S.A. Knyazev ( ZISP) ; 3 ♂, 1 ♀, 26– 27.07.2015, Buryatia, Selenga distr., 18 km SW of Gusinoozersk, Gusinoe lake , leg. S.A. Knyazev ( CKO) . Uzbekistan. 1 ♀, 4– 16.05.1892, Samarkand , leg. O. Herz ( ZISP) ; 1 ♀, 8.07.1905, Namangansky uyezd, Uzun-Akhmat , leg. Abramov ( ZISP) .

Figures

FIGURES 64–84. Drasteria specimens. 64–69. D. herzi (Alphéraky, 1895), lectotype (64: upperside; 65: underside; 66: labels), paralectotype (67), Tuz Golu, Turkey (68), Kugitang, Tajikistan (69). 70–72. D. kusnezovi (John, 1910), lectotype (70: upperside; 71: underside; 72: labels), paralectotype (73), Aral Sea, Kazakhstan (74), paralectotype (75), Altyn-Emel Nature Reserve, Kazakhstan (76), Koro-Goo Pass, Moldo-Too Mts., Kyrgyzstan (77, 78). 79–81. D. rada pia (Strand, 1917), holotype (79: upperside, 80: underside; 81: labels). 82–84. D. rada altivaga (Alphéraky, 1893), lectotype (82: upperside; 83: underside; 84: labels).

FIGURES 85–105. Drasteria specimens. 85–87. D. rada (Boisduval, 1848), Altyn-Emel Nature Reserve, Kazakhstan (85), Charyn Canyon, Kazakhstan (86), Ili river valley, Kazakhstan (87). 88–90. D. rada sibirica (Kozhantschikov, 1925), holotype (88: upperside; 89: underside; 90: labels). 91–96. D. christophi (Alphéraky, 1895), lectotype (91: upperside; 92: underside; 93: labels), holotype of dubia Bang-Haas in litt. (94: upperside; 95: underside; 96: labels). 97–99. D. kabylaria (Bang-Haas, 1906), lectotype (97: upperside; 98: underside; 99: labels). 100–102. D. kabylaria columbina (Brandt, 1941), paratype (100: upperside; 101: underside; 102: labels). 103–105. D. caucasica (Kolenati, 1848), Arzni, Armenia (103, 104: male; 105: female).

FIGURES 297–304. Drasteria habitats. 297. 25 km N of Panfilov, Dzhungarian Alatau, Kazakhstan (one of the authors in the collecting trip 2010). 298. Ili river valley near Koktal bridge, Almaty Prov., Kazakhstan (Russian Entomological Society and Institute of Zoology of Kazakhstan collecting trip, 2010). 299. Sands near Dosang environs, Astrakhan Prov., Russia. 300. Saksaul forest near Bakanas, Almaty Prov., Kazakhstan. 301. Chulak gorge, foothills of Dzhungarian Alatau Mts. in the Altyn-Emel Nature Reserve, Kazakhstan. 302. Ili river shore near Singing Dunes, Altyn-Emel Nature Reserve, Kazakhstan. 303. Katutau mountains, Altyn-Emel Nature Reserve, Kazakhstan. 304. Koram environs, northern slopes of Transilian Alatau Mts., Kazakhstan. Photos: S.K. Korb.