Dichotomius (Cephagonus) fonsecae ( Luederwaldt 1926 )

Nunes, Rafael V. & Vaz-de-Mello, Fernando Z., 2019, Taxonomic revision of Dichotomius (Cephagonus) Luederwaldt 1929 and the taxonomic status of remaining Dichotomius Hope 1838 subgenera (Coleoptera: Scarabaeidae: Scarabaeinae: Dichotomiini), Journal of Natural History 53 (37), pp. 2231-2351 : 2297-2298

2.2.: Dichotomius (Cephagonus) fonsecae ( Luederwaldt 1926) ( Figure 21 )

Pinotus fonsecae Luederwaldt 1926 ı p. 135 (description)

Pinotus (Cephagonus) fonsecae ı Luederwaldt 1929 ı p. 110

Dichotomius (selenocopris) sp.ı Sarmiento-Garcés & Amat-Garcia 2014 ı p. 104.

Diagnosis. distinguishable from the other species in the group by the following combination of characteristics: males ’ cephalic carina longer than wideı emarginated at apex producing two tubercles (as in D. fissus of the same group); anterior and posterior portion of the pronotum lacking distinct punctuation (at most some fineı sparse punctures as in D. seag of the same group); males ’ pronotal disc with a pair of acute tubercles at anterior portion; pronotal lateral borders almost completely smooth – except for a slightly irregular portion near medio-lateral pronotal pit and elytra lacking distinct microsculpture.

● HOLOTYPE of Pinotus fonsecae : O. ECUADOR. Macas. Feyer {whiteı printed}/TIPO {red with black marginsı printed}/ Pinotus fonsecae Lüd. Lüd Det 23. {whiteı handwritten}/ 25904 {whiteı printed} [MZSP].

Non type material. COLOMBIA: Caquetá. Solano. Parque Nacional Chiribiquete. 0³10 ʹ 54 ” Nı 72 ³36 ʹ 57 ” W. 800 mosl. E Gonzalez [1 Ƌ at CEMT]; PERU [1 Ƌ at UNMSM]; Júnin: Satipo. 1100 mosl. Fezes humanas. A Santibañez [3 ƋƋı 2 ♀♀ at CEMT].

Redescription. Colour: blackı shinyı with a brown sheen near the elytral base. Head: fronto-clypeal and genal surface smooth. Inter-ocular surface with fine punctures. Gena borders slightly curved inwards. Cephalic carina longer than wideı emarginate and producing two tubercles at the apex. Pronotum: lateral borders slightly irregular near the median lateral pronotal pit – rest of it smooth. Pronotal disc shinyı having a few spaced fine punctures and delimited anteriorly by a pair of acute tubercles. Anterior portionı including excavation and anterior angles with very few and inconspicuous sparse punctures. Posterior margin lacking punctuation. Metasternum: sides and anterior portion of anterior lobe covered by dense red setaeı setigerous punctures equally spacedı where present. Elytra: striae weakly impressed with weak punctures. Punctures separated by twice their diameter. Interstriae flatı lacking microsculpture. Abdomen: glabrousı punctuation of the anterior margin of the ventrites interrupted medially. Aedeagus ( Figure 21 (f-h)): parameraı apexes dorsally roundedı convergent; longitudinal excavation present taking more than 2/3 of the paramera length; basal excavation absent. Laterallyı abrupt declivity towards the apex. Ventrallyı subgenital plate rounded at the base and strongly emarginated at apex.

Morphological variation. males BL: 21 – 22 mmı PW: 12 mm. Females BL 18 – 20 mmı PW: 10 – 12 mm. Females distinguishable from males as follows: Head: fronto-clypeal surface transversely wrinkled. Cephalic carina very lowı producing two small tubercles. Pronotum: punctuation coarser than on males but following same distribution pattern. Elytra: striae deeply impressedı punctures separated by twice their diameter. Abdomen: sixth ventrite bearing a pair of triangular projections reaching the apical margin of the pygidium ( Figure 7 (b)).

Distribution. Macası West Equador (type-locality)ı Peru (Junín) and Colombia (Caquetá) ( Figure 17 ).

Figures

Figure 7. (a) Mandible of D. fissus (Harold), inc – incisory lobe (membranous), mo – molar lobe. (b) Maxilla and maxillary palp of same species, mxp – maxillary palp, ga – galea, las – lacinia. (c) Epipharynx of same species. (d) Labrum and labial palpomere of same species. (e) Ventral clypeal process of D. fissceps indicated by white arrow.(f) Antena of same species.esc – escapo, lam – lamella. (g) Macropterous hind wig of D. fissiceps. (h) Brachypterous hindwing of D. ingens.

Figure 17. D. mundus: (a) male dorsal habitus; (b) female dorsal habitus; (c) female ventral habitus; (d) paramera dorsal view; (e) paramera ventral view; (f) paramera lateral view; (g) type labels.

Figure 21. D. fonsecae: (a) male dorsal habitus; (b) Holotype dorsal habitus; (c) holotype labels; (d) female dorsal view; (e) female ventral view; (f) paramera dorsal view; (g) paramera ventral view; (h) paramera lateral view.