Dianthus ciliatus subsp. medunensis ( Beck & Szyszyłowicz 1888: 66 ) Trinajstić (1979b: 8)

Terlević, Ana, Ljubičić, Ivica, Tokić, Petar, Rešetnik, Ivana, Shuka, Donald, Siracusa, Giuseppe & Bogdanović, Sandro, 2024, Morphological and nomenclatural notes on Dianthus ciliatus subspecies (Caryophyllaceae), Phytotaxa 661 (3), pp. 213-234 : 224-225

4. Dianthus ciliatus subsp. medunensis ( Beck & Szyszyłowicz 1888: 66) Trinajstić (1979b: 8)

≡ Dianthus medunensis Beck & Szyszyłowicz (1888: 66) ≡ Dianthus ciliatus subsp. dalmaticus var. medunensis ( Beck & Szyszyłowicz 1888: 66) Hayek (1924: 246) .

Lectotype (designated here):— MONTENEGRO. Medun , July 1886, Szyszyłowicz s.n. ( PRC 455887!, Fig. 8).

— Dianthus ciliatus subsp. medunensis ( Beck & Szyszyłowicz 1888: 66) Trinajstić (1979a: 751) , nom. inval. (Art. 6, 29–31, 31–45 of ICN).

Notes: — Dianthus medunensis was described by Beck and Szyszyłowicz (1888) and illustrated in Beck & Szyszyłowicz (1888: table 3, fig. a, b, c) from Medun in Montenegro. Subsequently, Szyszyłowicz (1889: 118) republished again the same description in a separate paper. We were able to find original material in PRC herbarium that without any doubt belongs to Szyszyłowicz’s original collection ( PRC 455887!), and here we choose this specimen as lectotype ( Fig. 8).

Ecology and phenology: — Dianthus ciliatus subsp. medunensis ( Fig. 2E, F, G ) grows within sub-Mediterranean zone on calcareous rocky places, rocky grasslands, vertical inland cliffs, along roads and deciduous forest margins, where the specimens are rare in the field. It flowers from the end of June to the end of July, and fruits in August to September.

General distribution: — Albania and Montenegro ( Fig. 1 ). It occurs in the central part of Montenegro and central and north-western part of Albania.

Figures

FIGURE 1. Localities of sampled populations of taxa belonging to the endemic amphi-Adriatic Dianthus ciliatus used for morphometric analyses (n = 27 populations). Type localities are indicated with thick black symbol outlines.

FIGURE 2. Morphological details of corolla and calyx in Dianthus ciliatus subsp. ciliatus (A, B), D. ciliatus subsp. dalmaticus (C, D), and D. ciliatus subsp. medunensis (E, F, G). Habitus of D. ciliatus from Popoli (locus classicus) in Abruzzo, Italy (H), and floral quantitative characters analysed (I). Photographs by S. Bogdanović.