Desognaphosa yabbra, PLATNICK, 2002

PLATNICK, NORMAN I., 2002, A Revision Of The Australasian Ground Spiders Of The Families Ammoxenidae, Cithaeronidae, Gallieniellidae, And Trochanteriidae (Araneae: Gnaphosoidea), Bulletin of the American Museum of Natural History 2002 (271), pp. 1-1 : 1-

Desognaphosa yabbra , new species Figures 152 , 181–190 ; Map 14

TYPES: Female holotype and male allotype taken in pitfall trap in Yabbra State Forest , 28 ° 30 ̍ S, 152 ° 40 ̍ E, New South Wales (Feb. 7–Mar. 15, 1992; M. Gray, P. Croft), deposited in AMS (KS38455) .

ETYMOLOGY: The specific name is a noun in apposition taken from the type locality.

DIAGNOSIS: Males resemble those of D. dryander in having a moderately long embolus (fig. 187), but have a larger retrolateral tibial apophysis (fig. 188); females also resemble those of D. dryander in having a relatively narrow anterior epigynal margin (fig. 189), but have a much more elongated epigynum, with the anterior margin far removed from the ducts (fig. 190).

MALE: Total length 5.7. Carapace dark reddish brown; abdominal dorsum gray with two paramedian longitudinal rows of white spots, most anterior pair of spots much longer than others, posterior pairs grading into chevrons, sides gray, venter white with three longitudinal gray stripes; legs light reddish brown, femora darkest. Leg spination: tibiae: III v1p­1p­2; IV r0­1­1; metatarsi: III p0­0­ 1, v2­0­1p; IV p0­0­1, v2­1p­1p, r1­0­0. Retrolateral tibial apophysis triangular, thickened, cymbial excavation deep, extending over half length of segment (fig. 188); embolus extending posterior of tibial margin, with prominent, prolaterally directed protuberance at base (fig. 187).

FEMALE: Total length 5.3. Coloration as in male. Leg spination: tibia IV d1­0­0, r0­1­1; metatarsi: III v2­0­2; IV v2­1p­2, r1­0­0. Epigynum with elevated anterior margin, openings medially situated (fig. 189); ducts bridged by w­shaped, transverse bar (fig. 190).

OTHER MATERIAL EXAMINED: New South Wales: Blue Gum Lookout, via Mallanganee , 28 ° 47 ̍ S, 152 ° 43 ̍ E, Oct. 12, 1977 – Feb. 7, 1978, rainforest pitfall, elev. 580 m (G. Monteith, QMB S26658 ), 1♀ ; Cambridge Plateau, Richmond Range State Forest , 28 ° 47 ̍ S, 152 ° 45 ̍ E, Dec. 17, 1988, subtropical rainforest pitfall, elev. 300–600 m (Smith, Hines, Pugh, Webber, AMS KS44244, 44816, 57716), 33 ; Elkhorn Road, Ewingar State Forest , 29 ° 06 ̍ S, 152 ° 26 ̍ E, Feb. 4–Apr. 9, 1993, pitfall, elev. 670–710 m (M. Gray, G. Cassis, AMS KS40471, 40472), 73, 2♀ ; Ewingar Creek, Elkhorn Road, Ewingar State Forest , 29 ° 05 ̍ S, 152 ° 26 ̍ E, Feb. 4–Apr. 9, 1993, pitfall, elev. 640 m (M. Gray, G. Cassis, AMS KS40470 ), 53, 1♀ ; Fosters Road, 300 m from Running Road, Whian Whian State Forest , 28 ° 36 ̍ S, 153 ° 20 ̍ E, Feb. 4–Apr. 9, 1993, pitfall, elev. 310 (M. Gray, G. Cassis, AMS KS35795 ), 2♀ ; Gibburgunyah Range Road, 150 m W Rocky Creek crossing, Big Scrub Flora Reserve , 28 ° 38 ̍ S, 153 ° 19 ̍ E, Feb. 4–Apr. 9, 1993, pitfall, elev. 180 (M. Gray, G. Cassis, AMS KS35796 ), 43, 1♀ ; Lionsville Road, Ewingar State Forest , 29 ° 11 ̍ S, 152 ° 25 ̍ E, Feb. 4–Apr. 9, 1993, pitfall, elev. 740 m (M. Gray, G. Cassis, AMS KS42184 ), 23, 1♀ ; NW Rocky Creek crossing, Big Scrub Flora Reserve , 28 ° 38 ̍ S, 153 ° 19 ̍ E, Feb. 4–Apr. 9, 1993 (M. Gray, G. Cassis, AMS KS35794 ), 13 ; Running Road, 100 m E Rocky Creek, Whian Whian State Forest , 28 ° 36 ̍ S, 153 ° 21 ̍ E, Feb. 4–Apr. 9, 1993, pitfall, elev. 250 (M. Gray, G. Cassis, AMS KS35797 ), 2♀ ; Sheepstation Creek, Wiangaree State Forest , 28 ° 24 ̍ S, 153 ° 04 ̍ E, Feb. 29–Mar. 3, 1980, Berlese, leaf and log litter, subtropical rainforest, elev. 600 m (A. Newton, M. Thayer, AMNH), 2♀ ; Solferino Creek, track off Lionsville Road, Ewingar State Forest , 29 ° 10 ̍ S, 152 ° 26 ̍ E, Feb. 4–Apr. 9, 1993, pitfall, elev. 520 m (M. Gray, G. Cassis, ex AMS KS40463 ), 1♀ ; Terania Creek, N Lismore, 28 ° 34 ̍ S, 153 ° 19 ̍ E, Apr.– May , 1976, rainforest litter, elev. 340 m (M. Gray, C. Horseman, AMS KS10070 ), 1♀ ; Tributary Grasstree Creek, Nogrigar Road, Ewingar State Forest , 29 ° 08 ̍ S, 152 ° 25 ̍ E, Feb. 4–Apr. 9, 1993, pitfall, elev. 720 m (M. Gray, G. Cassis, AMS KS40473 ), 33 ; Tuck­ er Box Road, Beaury State Forest , 28 ° 28 ̍ S, 152 ° 24 ̍ E, Feb. 4–Apr. 9, 1993, pitfall, elev. 740 m (M. Gray, G. Cassis, AMS KS36076 ), 33 ; Tweed Range Road, 4.6 km SW Bridle Creek Road, Border Ranges National Park , 28 ° 24 ̍ S, 153 ° 02 ̍ E, Feb. 4–Apr. 9, 1993, pit­ fall, elev. 580 m (M. Gray, G. Cassis, AMS KS40460 ), 1♀ ; Victoria Park, via Alstonville , 28 ° 53 ̍ S, 153 ° 23 ̍ E, Dec. 26, 1974 – Mar. 23, 1975, rainforest pitfall, elev. 213 m (G., S. Monteith, QMB S26624 ), 13, Aug. 3– Nov. 16, 1975, rainforest pitfall (G., S. Monteith, QMB S26629 ), 1♀ ; N along Wallaby Creek, Beaury State Forest , 28 ° 26 ̍ S, 152 ° 27 ̍ E, Feb. 4–Apr. 9, 1993, pitfall, elev. 830 m (M. Gray, G. Cassis, AMS KS36087 ), 43, 1♀ ; Whian Whian State Forest , 28 ° 41 ̍ S, 153 ° 19 ̍ E, Dec. 26, 1974 – Mar. 23, 1975, rainforest pitfall (G., S. Monteith, QMB S26638 ), 1♀, Sept. 12, 1976, rainforest litter (R. Raven, QMB S34522 ), 2♀ ; Woobah Fire Trail, Bundjalung National Park , 29 ° 15 ̍ S, 153 ° 17 ̍ E, Feb. 4–Apr. 9, 1993, pitfall (M. Gray, G. Cassis, AMS KS40459 ), 13 ; Yabbra State Forest, 28 ° 30 ̍ S, 152 ° 40 ̍ E, Feb. 7– Mar. 15, 1992, pitfall (M. Gray, P. Croft, AMS KS38447, 38463, 38798), 13, 2♀. Queensland: Booloumba Creek, Conondale Range , 26 ° 39 ̍ S, 152 ° 39 ̍ E, Nov. 29, 1974 – Feb. 22, 1975, rainforest pitfall, elev. 518 m (G., S. Monteith, QMB S26625 ), 1♀, Feb. 22–Apr. 19, 1975, rainforest pitfall, elev. 550 m (G., S. Monteith, QMB S26630 ), 13, May 13–18, 1976, rainforest litter (R. Raven, QMB S28770 ), 23, 9♀ ; Booloumba Creek Road, Kenilworth State Forest , 4 km W Cambroon, 26 ° 38 ̍ S, 152 ° 39 ̍ E, May 5, 1998, wet sclerophyll forest (G. Milledge, AMS KS52278 ), 3♀ ; Cold Creek, via Imbil , 26 ° 28 ̍ S, 152 ° 41 ̍ E, Nov. 9–Dec. 31, 1974, rainforest pitfall, elev. 122 m (G., S. Monteith, QMB S26633 ), 13 ; Conondale Range , 26 ° 50 ̍ S, 152 ° 45 ̍ E, May 1–3, 1976 (R. Raven, AMS S29330 ), 1♀ ; Cooran Plateau, via Traveston , 26 ° 21 ̍ S, 152 ° 47 ̍ E, Mar. 28–Aug. 13, 1975, rainforest pitfall, elev. 366 m (G., S. Monteith, QMB S26627 ), 13 ; Cunninghams Gap , 28 ° 03 ̍ S, 152 ° 24 ̍ E, Dec. 28, 1974 – Mar. 30, 1975, rainforest pitfall, elev. 762 m (G., S. Monteith, QMB S26626 , 26662 ), 13, 1♀, June 28, 1991, dry forest litter (D. Black, WAM 99 /647, 648), 13, 1♀, Jan. 6– Mar. 1, 1992, rainforest pitfall, elev. 790 m (D. Cook, QMB S25045 ), 13 ; Hiller property, Mount Glorious , 27 ° 20 ̍ S, 152 ° 46 ̍ E, Nov. 18, 1977 ­ Jan. 20, 1978, rainforest pitfall (G., S. Monteith, QMB S26701 ), 1♀, June 26–Nov. 18, 1978, same ( QMB S26704 ), 1♀ ; Kenilworth State Forest, 5.2 km SW Kenilworth , 26 ° 37 ̍ S, 152 ° 41 ̍ E, May 5, 1998, rainforest (G. Milledge, AMS KS52289 ), 2♀ ; Levers Plateau , 28 ° 19 ̍ S, 152 ° 15 ̍ E, Feb. 29–Apr. 11, 1993, rainforest pitfall, elev. 720 m (D. Cook, QMB S43395 ), 13 ; Little Yabba Creek , 26 ° 37 ̍ S, 152 ° 41 ̍ E, Aug. 10–Dec. 29, 1974, rainforest pitfall, elev. 152 m (G., S. Monteith, QMB S26622 , 26639 ), 23, 2♀, Jan. 27, 1980 (V. Davies, QMB S26668 ), 1♀ ; Marys Creek State Forest, via Gympie , 26 ° 15 ̍ S, 152 ° 35 ̍ E, Aug. 11–Nov. 10, 1974, rainforest pitfall, elev. 183 m (G., S. Monteith, QMB S40714 ), 13, 1♀, Dec. 31, 1974 – Mar. 28, 1975, same ( QMB S26637 ), 1♀, June 16–Aug. 23, 1975, same ( QMB S26623 ), 1♀ ; Mount Coolum, Fern Gully , 26 ° 34 ̍ S, 153 ° 05 ̍ E, Jan. 1984, pitfall (B. Jahnke, QMB S26670 ), 13 ; Mount Cotton , upper gully, 27 ° 36 ̍ S, 153 ° 13 ̍ E, May 7, 1988, Berlese, rainforest litter, elev. 150 m (G. Monteith, G. Thompson, QMB S42445 ), 1♀ ; Mount Cotton, Scott’s Dam , 27 ° 36 ̍ S, 153 ° 13 ̍ E, May 7, 1988, Berlese, rainforest litter, elev. 120 m (G. Monteith, G. Thompson, QMB S42444 ), 1♀ ; Mount Cotton , 27 ° 37 ̍ S, 153 ° 13 ̍ E, Sept. 13, 1997, Berlese, rainforest litter, elev. 200 m (G. Monteith, QMB S43873 ), 1♀ ; Mount Glorious , 27 ° 20 ̍ S, 152 ° 46 ̍ E, Sept. 20, 1979, rainforest, Berlese, stick brushings (G. Monteith, QMB S26686 ), 13 ; Mount Mee, Neurum Creek , 27 ° 05 ̍ S, 152 ° 42 ̍ E, Jan. 20–Feb. 26, 1978, rainforest pitfall, elev. 210 m (G., S. Monteith, QMB S26697 ), 1♀ ; Mount Mee, via Samford , 27 ° 05 ̍ S, 152 ° 47 ̍ E, Mar. 20–June 1, 1975, rainforest pitfall (G., S. Monteith, QMB S26635 , 26636 ), 23 ; Mount Mee , top, 27 ° 05 ̍ S, 152 ° 41 ̍ E, Oct. 28, 1977 – Jan. 20, 1978, rainforest pitfall, elev. 550 m (G., S. Monteith, QMB S26659 ), 23, Feb. 26–June 26, 1978, same ( QMB S26664 ), 13 ; Mount Tamborine , 27 ° 56 ̍ S, 153 ° 12 ̍ E, July 10, 1974 (C. Wilton, V. Davies, R. Raven, QMB S26707 ), 1♀ ; N side, Mount Tamborine, near Curtis Falls , 27 ° 58 ̍ S, 153 ° 12 ̍ E, May 8, 1953, rainforest litter (T. Woodward, OMD), 2♀ ; Mount Tenison Woods , 27 ° 18 ̍ S, 152 ° 45 ̍ E, Mar. 22, 1979, rainforest, Berlese, stick brushings (G. Monteith, QMB S26666 ), 13, 1♀ ; Mount Tenison Woods , 27 ° 19 ̍ S, 152 ° 44 ̍ E, May 15, 1997, Berlese, rainforest litter (G. Monteith, QMB S43876 ), 1♀ ; Palm Grove, Mount Tamborine , 27 ° 56 ̍ S, 153 ° 12 ̍ E, Oct. 26–Dec. 14, 1974, rainforest pitfall, elev. 670 m (G., S. Monteith, QMB S26628 ), 1♀ ; Sunday Creek , 26 ° 40 ̍ S, 152 ° 34 ̍ E, Jan. 20–May 15, 1997, rainforest pitfalls (G. Monteith, QMB S38078, 38091 ), 23 ; Sunday Creek Road, junction with Gigh­ er Creek Road, Kenilworth State Forest , 26 ° 42 ̍ S, 152 ° 33 ̍ E, May 7, 1998, wet sclerophyll forest (G. Milledge, AMS KS52282 ), 2♀ ; Upper Tallebudgera Valley, below Springbrook , 28 ° 15 ̍ S, 153 ° 16 ̍ E, Jan. 8–Mar. 17, 1985, rainforest pitfall, elev. 550 m (G. Monteith, G. Thompson, D. Cook, QMB S26653 ), 1♀ ; Yabba Creek forest , 7 km SW Kenilworth, 26 ° 35 ̍ S, 152 ° 44 ̍ E, Aug. 15, 1982, rainforest litter, elev. 150 m (S., J. Peck, AMNH), 1♀ .

DISTRIBUTION: Southeastern Queensland and adjacent New South Wales (map 14).

Desognaphosa bulburin , new species Figures 193, 194 ; Map 15 TYPE : Female holotype taken in a pitfall trap in rainforest at an elevation of 609 m on

the Bulburin Plateau, via Miriamvale, Bulburin State Forest , 24 ° 30 ̍ S, 151 ° 35 ̍ E, Queensland (June 1–Oct. 5, 1974; G., S. Monteith), deposited in QMB ( S26634 ) .

ETYMOLOGY: The specific name is a noun in apposition taken from the type locality.

DIAGNOSIS: Females of this distinctive species can easily be recognized by the oval, medially situated epigynal atrium (fig. 193).

MALE: Unknown.

FEMALE: Total length 4.5. Coloration as in D. yabbra , except pars thoracica with dark markings, legs with dark markings distally on femora and patellae, proximally and distally on tibiae. Leg spination: tibiae: III v1p­ 1p­2; IV r0­1­1; metatarsi: III v1p­0­2; IV v2­1p­2, r1­0­0. Epigynum with transverse, oval atrium (fig. 193); anterolateral portion of ducts making complete coil (fig. 194).

OTHER MATERIAL EXAMINED: Queensland: Bulburin State Forest, 24 ° 30 ̍ S, 151 ° 35 ̍ E, Mar. 17–24, 1975, rainforest litter (V. Davies, R. Kohout, QMB S28302 ), 1♀, Mar. 25–28, 1977 (R. Raven, V. Davies, ex QMB S26953 ), 1♀.

DISTRIBUTION: Known only from Bulburin State Forest in mideastern Queensland (map 15).

Figures

Map 14. Records of Desognaphosa yabbra, new species (circles), D. malbon, new species (square), and D. boolbun, new species (diamonds).

Map 15. Records of Desognaphosa bulburin, new species (circle), D. tyson, new species (square), and D. carbine, new species (diamonds).

Figs. 149–154. Palpal endites of females, anterior views. 149. Meedo mullaroo, new species. 150. Neato kioloa, new species. 151. Tinytrema bondi, new species. 152. Desognaphosa yabbra, new species. 153. Trachycosmus sculptilis Simon. 154. Trachytrema castaneum Simon.

Figs. 181–186. Desognaphosa yabbra, new species, spinnerets of male (left) and female (right), distal views. 181, 182. Anterior lateral spinnerets. 183, 184. Posterior median spinnerets. 185, 186. Posterior lateral spinnerets.

Figs. 187–190. Desognaphosa yabbra, new species. 187. Left male palp, ventral view. 188. Same, retrolateral view. 189. Epigynum, ventral view. 190. Same, dorsal view.

Figs. 191–196. Epigyna, ventral (top row) and dorsal (bottom row) views. 191, 192. Olin platnicki Deeleman­Reinhold. 193, 194. Desognaphosa bulburin, new species. 195, 196. D. kroombit, new species.